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<document id="21FBE09C4E854D2CDCA8BC4492DCB265" ID-CLB-Dataset="4378" ID-DOI="10.1046/j.1096-3642.2003.00044.x" ID-GBIF-Dataset="8ef22762-4fd3-4d65-8d0b-20b07d88d5d1" ID-ISSN="0024-4082" ID-Zenodo-Dep="5437155" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630436240609" checkinUser="carolina" docAuthor="Crochet, Pierre-André, Geniez, Philippe &amp; Ineich, Ivan" docDate="2003" docId="03C1AF1C8314FFF6FF543BD815D0F992" docLanguage="en" docName="j.1096-3642.2003.00044.x.pdf" docOrigin="Zoological Journal of the Linnean Society 137 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2003.00044.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Acanthodactylus longipes Boulenger 1918" docType="treatment" docVersion="7" lastPageNumber="145" masterDocId="FFF8D7648308FFEAFFFC3B1A1610FFDF" masterDocTitle="A multivariate analysis of the fringe-toed lizards of the Acanthodactylus scutellatus group (Squamata: Lacertidae): systematic and biogeographical implications" masterLastPageNumber="155" masterPageNumber="117" pageNumber="145" updateTime="1699279316374" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="78E29229CC4F2BB4E3A4820608CE14DA">A multivariate analysis of the fringe-toed lizards of the Acanthodactylus scutellatus group (Squamata: Lacertidae): systematic and biogeographical implications</mods:title>
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<mods:namePart id="AEA0407A25A183FE8E970BF19B6FEDCD">Ineich, Ivan</mods:namePart>
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<treatment id="03C1AF1C8314FFF6FF543BD815D0F992" ID-DOI="http://doi.org/10.5281/zenodo.5490910" ID-GBIF-Taxon="185447196" ID-Zenodo-Dep="5490910" LSID="urn:lsid:plazi:treatment:03C1AF1C8314FFF6FF543BD815D0F992" httpUri="http://treatment.plazi.org/id/03C1AF1C8314FFF6FF543BD815D0F992" lastPageNumber="145" pageId="28" pageNumber="145">
<subSubSection id="C3724D818314FFF6FF543BD814CCFF05" box="[168,732,194,218]" pageId="28" pageNumber="145" type="nomenclature">
<paragraph id="8BD71E0A8314FFF6FF543BD814CCFF05" blockId="28.[168,732,194,218]" box="[168,732,194,218]" pageId="28" pageNumber="145">
<heading id="D09FA9668314FFF6FF543BD814CCFF05" box="[168,732,194,218]" centered="true" fontSize="9" level="2" pageId="28" pageNumber="145" reason="2">
<taxonomicName id="4C6865898314FFF6FF543BD814CCFF05" authority="BOULENGER, 1918" authorityName="Boulenger" authorityYear="1918" box="[168,732,194,218]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FF543BD81411FF06" box="[168,513,194,218]" italics="true" pageId="28" pageNumber="145">ACANTHODACTYLUS LONGIPES</emphasis>
<bibRefCitation id="EFF963FB8314FFF6FDF53BD814CCFF05" author="Boulenger GA" box="[521,732,194,218]" pageId="28" pageNumber="145" pagination="143 - 155" refId="ref21726" refString="Boulenger GA. 1918. Sur les lezards du genre Acanthodactylus Wiegm. Bulletin de la Societe Zoologique de France 43: 143 - 155." type="journal article" year="1918">BOULENGER, 1918</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3724D818314FFF6FF723BF11473FCAD" pageId="28" pageNumber="145" type="reference_group">
<paragraph id="8BD71E0A8314FFF6FF723BF117C5FD12" blockId="28.[142,759,235,717]" pageId="28" pageNumber="145">
<treatmentCitationGroup id="AB7839248314FFF6FF723BF117C5FD12" pageId="28" pageNumber="145">
<treatmentCitation id="0AC9381B8314FFF6FF723BF11715FEFF" author="Boulenger GA" page="154" pageId="28" pageNumber="145" year="1918">
<taxonomicName id="4C6865898314FFF6FF723BF11715FEFF" authority="Boulenger, 1918: 154" authorityName="Boulenger" authorityPageNumber="154" authorityYear="1918" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="variety" species="scutellatus" variety="longipes">
<emphasis id="B91CC2188314FFF6FF723BF117C1FEDF" box="[142,465,235,256]" italics="true" pageId="28" pageNumber="145">Acanthodactylus scutellatus</emphasis>
var.
<emphasis id="B91CC2188314FFF6FDEF3BF11460FEDF" box="[531,624,235,256]" italics="true" pageId="28" pageNumber="145">longipes</emphasis>
<bibRefCitation id="EFF963FB8314FFF6FD813BF11715FEFF" author="Boulenger GA" pageId="28" pageNumber="145" pagination="143 - 155" refId="ref21726" refString="Boulenger GA. 1918. Sur les lezards du genre Acanthodactylus Wiegm. Bulletin de la Societe Zoologique de France 43: 143 - 155." type="journal article" year="1918">Boulenger, 1918: 154</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Name-bearing type: Boulenger does not refer in his original description to any specimen but he clearly used several individuals housed in the British Museum and coming from the Algerian Sahara. He latter talks about
<specimenCount id="9D6ED5838314FFF6FE903A9E1406FE45" box="[364,534,388,410]" pageId="28" pageNumber="145" type="generic">five specimens</specimenCount>
(
<bibRefCitation id="EFF963FB8314FFF6FDD63A9E14F9FE44" author="Boulenger GA" box="[554,745,388,411]" pageId="28" pageNumber="145" refId="ref21752" refString="Boulenger GA. 1921. Monograph of the Lacertidae, II. London: British Museum (Natural History)." type="book" year="1921">Boulenger, 1921</bibRefCitation>
), which are apparently all
<typeStatus id="54D3A0A88314FFF6FE413ABE1434FE66" box="[445,548,420,441]" pageId="28" pageNumber="145" type="syntype">syntypes</typeStatus>
. These specimens are (see also
<collectingCountry id="F37F5E9A8314FFF6FEC53AD8178EFE07" box="[313,414,450,472]" name="El Salvador" pageId="28" pageNumber="145">Salvador</collectingCountry>
, 1982): BMNH
<date id="FFD638CA8314FFF6FD983AD814C3FE07" box="[612,723,450,472]" pageId="28" pageNumber="145" value="1946-08-30">1946.8.30</date>
-32 (Wargla), BMNH 1946.9.3.75 (between the Wed Nça and El Alia), BMNH 1946.9.3.74 (El Wed, East of Tuggurt). As the original type series might have included more specimens, including members of other species, we select as
<typeStatus id="54D3A0A88314FFF6FEA7394117D1FDAE" box="[347,449,603,625]" pageId="28" pageNumber="145" type="lectotype">lectotype</typeStatus>
the specimen BMNH 1946.8.4.31 (an adult male from Wargla,
<collectingCountry id="F37F5E9A8314FFF6FD6F396014FBFD4F" box="[659,747,634,656]" name="Algeria" pageId="28" pageNumber="145">Algeria</collectingCountry>
). Type locality: restricted by
<typeStatus id="54D3A0A88314FFF6FE2739821451FD71" box="[475,577,664,686]" pageId="28" pageNumber="145" type="lectotype">lectotype</typeStatus>
designation to Wargla [=
<collectingRegion id="49ACD0E88314FFF6FEFB39AC1778FD13" box="[263,360,694,716]" country="Algeria" name="Ouargla" pageId="28" pageNumber="145">Ouargla</collectingRegion>
],
<collectingCountry id="F37F5E9A8314FFF6FE8639AD17C1FD12" box="[378,465,695,717]" name="Algeria" pageId="28" pageNumber="145">Algeria</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD71E0A8314FFF6FF5B39F91473FCAD" blockId="28.[142,758,738,883]" pageId="28" pageNumber="145">
<emphasis id="B91CC2188314FFF6FF5B39F91770FD27" box="[167,352,739,760]" italics="true" pageId="28" pageNumber="145">Junior synonym</emphasis>
.
<taxonomicName id="4C6865898314FFF6FE5D39F8142FFCC9" authority="Bons &amp; Girot, 1964: 327" authorityName="Bons &amp; Girot" authorityPageNumber="327" authorityYear="1964" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="subSpecies" species="longipes" subSpecies="panousei">
<emphasis id="B91CC2188314FFF6FE5D39F816E3FCC8" italics="true" pageId="28" pageNumber="145">Acanthodactylus longipes panousei</emphasis>
<bibRefCitation id="EFF963FB8314FFF6FEF83818142FFCC9" author="Bons J &amp; Girot B" box="[260,575,769,791]" pageId="28" pageNumber="145" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons &amp; Girot, 1964: 327</bibRefCitation>
</taxonomicName>
. Name-bearing types:
<specimenCount id="9D6ED5838314FFF6FF23383A176AFCEA" box="[223,378,800,821]" pageId="28" pageNumber="145" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
MNHN 1963.1013-1014 (called ‘holotypes’ in
<bibRefCitation id="EFF963FB8314FFF6FED638251419FC8B" author="Bons J &amp; Girot B" box="[298,521,830,852]" pageId="28" pageNumber="145" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons &amp; Girot, 1964</bibRefCitation>
). Type locality: ‘Bord de l’Erg Chebbi au niveau de Merzouga’.
</paragraph>
</subSubSection>
<subSubSection id="C3724D818314FFF6FF5B389215D0F992" pageId="28" pageNumber="145" type="description">
<paragraph id="8BD71E0A8314FFF6FF5B38921771FC04" blockId="28.[142,758,904,988]" pageId="28" pageNumber="145">
<emphasis id="B91CC2188314FFF6FF5B38921721FC42" box="[167,305,904,925]" italics="true" pageId="28" pageNumber="145">Chresonyms</emphasis>
.
<taxonomicName id="4C6865898314FFF6FEBA389216D8FC63" authority="Boulenger, 1918" authorityName="Boulenger" authorityYear="1918" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FEBA3892147EFC42" box="[326,622,904,925]" italics="true" pageId="28" pageNumber="145">Acanthodactylus longipes</emphasis>
<bibRefCitation id="EFF963FB8314FFF6FD81389216D8FC63" author="Boulenger GA" pageId="28" pageNumber="145" pagination="143 - 155" refId="ref21726" refString="Boulenger GA. 1918. Sur les lezards du genre Acanthodactylus Wiegm. Bulletin de la Societe Zoologique de France 43: 143 - 155." type="journal article" year="1918">Boulenger, 1918</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="EFF963FB8314FFF6FF2A38B217EAFC63" author="Bons J &amp; Girot B" box="[214,506,935,957]" pageId="28" pageNumber="145" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons &amp; Girot, 1964: 324</bibRefCitation>
;
<collectingCountry id="F37F5E9A8314FFF6FDF638BD147FFC62" box="[522,623,935,957]" name="El Salvador" pageId="28" pageNumber="145">Salvador</collectingCountry>
, 1982: 132;
<bibRefCitation id="EFF963FB8314FFF6FF7238DC174DFC04" author="Arnold EN" box="[142,349,966,988]" pageId="28" pageNumber="145" pagination="291 - 339" refId="ref21308" refString="Arnold EN. 1983. Osteology, genitalia and the relationships of Acanthodactylus (Reptilia: Lacertidae). Bulletin of the British Museum (Natural History) Zoology 44: 291 - 339." type="journal article" year="1983">Arnold, 1983: 324</bibRefCitation>
.
</paragraph>
<paragraph id="8BD71E0A8314FFF6FF5B38EB142DFB7E" blockId="28.[142,758,1009,1185]" pageId="28" pageNumber="145">
<emphasis id="B91CC2188314FFF6FF5B38EB1722FBD9" box="[167,306,1009,1030]" italics="true" pageId="28" pageNumber="145">Distribution</emphasis>
(
<figureCitation id="1353028F8314FFF6FEBB38EB1786FBD8" box="[327,406,1009,1031]" captionStart="Figure 19" captionStartId="29.[155,233,1033,1052]" captionTargetBox="[157,1431,198,996]" captionTargetId="figure-430@29.[155,1432,194,1001]" captionTargetPageId="29" captionText="Figure 19. Geographical distribution of Acanthodactylus longipes. Data from Salvador (1982), Baha El Din (1994), Bons &amp; Geniez (1996), Nouïra (1996), this study." figureDoi="http://doi.org/10.5281/zenodo.5437193" httpUri="https://zenodo.org/record/5437193/files/figure.png" pageId="28" pageNumber="145">Fig. 19</figureCitation>
). Most of the Sahara from coastal
<collectingCountry id="F37F5E9A8314FFF6FF183F0A1777FBF9" box="[228,359,1040,1062]" name="Mauritania" pageId="28" pageNumber="145">Mauritania</collectingCountry>
(this study; A. Foucart, pers. com.), southern
<collectingCountry id="F37F5E9A8314FFF6FEFF3F351770FB9B" box="[259,352,1071,1092]" name="Morocco" pageId="28" pageNumber="145">Morocco</collectingCountry>
(Tafilalet, Iriki) (
<bibRefCitation id="EFF963FB8314FFF6FDC13F3416D7FBBC" author="Geniez P &amp; Soto P" pageId="28" pageNumber="145" pagination="33 - 39" refId="ref22077" refString="Geniez P, Soto P. 1994. Nouvelles observations sur l'herpetofaune marocaine, 5. Bulletin de la Societe Herpetologique de France 69 - 70: 33 - 39." type="journal article" year="1994">Geniez &amp; Soto, 1994</bibRefCitation>
), northern
<collectingCountry id="F37F5E9A8314FFF6FEB53F571790FBBC" box="[329,384,1101,1123]" name="Serbia" pageId="28" pageNumber="145">Mali</collectingCountry>
, northern Niger, northern
<collectingCountry id="F37F5E9A8314FFF6FD4E3F5714E1FBBC" box="[690,753,1101,1123]" name="Chad" pageId="28" pageNumber="145">Chad</collectingCountry>
, Algerian Sahara,
<collectingCountry id="F37F5E9A8314FFF6FEA73F7617A4FB5D" box="[347,436,1132,1154]" name="Tunisia" pageId="28" pageNumber="145">Tunisia</collectingCountry>
,
<collectingCountry id="F37F5E9A8314FFF6FE3D3F761412FB5D" box="[449,514,1132,1154]" name="Libya" pageId="28" pageNumber="145">Libya</collectingCountry>
(
<collectingCountry id="F37F5E9A8314FFF6FDEE3F761468FB5D" box="[530,632,1132,1154]" name="El Salvador" pageId="28" pageNumber="145">Salvador</collectingCountry>
, 1982; this study), to
<collectingCountry id="F37F5E9A8314FFF6FEFF3F911757FB7F" box="[259,327,1163,1184]" name="Egypt" pageId="28" pageNumber="145">Egypt</collectingCountry>
(
<bibRefCitation id="EFF963FB8314FFF6FEAB3F901421FB7E" author="Baha El Din SM" box="[343,561,1162,1185]" pageId="28" pageNumber="145" pagination="18 - 27" refId="ref21524" refString="Baha El Din SM. 1994. A contribution to the herpetology of Sinai. British Herpetological Society Bulletin 48: 18 - 27." type="journal article" year="1994">Baha El Din, 1994</bibRefCitation>
).
</paragraph>
<paragraph id="8BD71E0A8314FFF6FF5B3FAC1297FA59" blockId="28.[142,759,1206,1902]" lastBlockId="28.[806,1423,196,1414]" pageId="28" pageNumber="145">
<emphasis id="B91CC2188314FFF6FF5B3FAC1707FB14" box="[167,279,1206,1227]" italics="true" pageId="28" pageNumber="145">Diagnosis</emphasis>
. Medium-sized species (reaching
<quantity id="4C90B3EF8314FFF6FD543FAC14E6FB14" box="[680,758,1206,1228]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="28" pageNumber="145" unit="mm" value="61.0">61 mm</quantity>
snout-vent length, mean: 52.4). The subocular in contact with three or (rarely) four supralabials in 100% of the individuals (
<tableCitation id="C6EA2BB18314FFF6FEB03E0817B0FAF7" box="[332,416,1298,1320]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="28" pageNumber="145">Table 3</tableCitation>
) distinguishes
<taxonomicName id="4C6865898314FFF6FDAF3E081705FA9A" authorityName="Boulenger" authorityYear="1918" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FDAF3E081705FA9A" italics="true" pageId="28" pageNumber="145">Acanthodactylus longipes</emphasis>
</taxonomicName>
from
<taxonomicName id="4C6865898314FFF6FEA73E2B17D7FA99" authorityName="Gunther" authorityYear="1903" box="[347,455,1329,1350]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="aureus">
<emphasis id="B91CC2188314FFF6FEA73E2B17D7FA99" box="[347,455,1329,1350]" italics="true" pageId="28" pageNumber="145">A. aureus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6865898314FFF6FDFD3E2B148CFA9A" authorityName="Geniez &amp; Foucart" authorityYear="1995" box="[513,668,1328,1350]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="taghitensis">
<emphasis id="B91CC2188314FFF6FDFD3E2B148CFA9A" box="[513,668,1328,1350]" italics="true" pageId="28" pageNumber="145">A. taghitensis</emphasis>
</taxonomicName>
. Dorsal scales small, very numerous (range: 55–77, mean: 66.4;
<tableCitation id="C6EA2BB18314FFF6FF2C3E741734FA5B" box="[208,292,1390,1412]" captionStart="Table 2" captionStartId="5.[155,219,196,215]" captionText="Table 2. Means, standard deviations (in parentheses) and ranges of the quantitative variables used in this study. See Table 1 for variable definitions" pageId="28" pageNumber="145">Table 2</tableCitation>
), elongate, smooth except in the vertebral area where they can be weakly to moderately keeled (code 3 or less in 97% of the individuals, no individual reaching code 5;
<tableCitation id="C6EA2BB18314FFF6FE363ED0140DFA00" box="[458,541,1482,1504]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="28" pageNumber="145">Table 3</tableCitation>
, see
<figureCitation id="1353028F8314FFF6FDAA3ED014A3FA3F" box="[598,691,1482,1504]" captionStart="Figure 16" captionStartId="23.[155,233,1091,1110]" captionTargetBox="[154,1434,199,1058]" captionTargetId="figure-466@23.[153,1435,199,1058]" captionTargetPageId="23" captionText="Figure 16. (a) Acanthodactylus scutellatus audouini, male, paralectotype of Acanthodactylus scutellatus var. audouini, BMNH 97.10.28.316, Wadi Halfa (Egypt), close-up of dorsal scales. (b) A. senegalensis, male, EPHE Mch1, coast north of Dakar (Senegal), close-up of dorsal scales. (c) A. dumerili, male, MNHN 1997.3764, Blaouakh (Mauritania), close-up of dorsal scales. (d) A. longipes, male, EPHE AF4, Akreïdil, 50 km NE of Nouakchott (Mauritania), close-up of dorsal scales." figureDoi="http://doi.org/10.5281/zenodo.5437187" httpUri="https://zenodo.org/record/5437187/files/figure.png" pageId="28" pageNumber="145">Fig. 16d</figureCitation>
). Two rows or more of supraciliary granules in 86% of the individuals (
<tableCitation id="C6EA2BB18314FFF6FEDE3D1D1767F9C3" box="[290,375,1543,1565]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="28" pageNumber="145">Table 3</tableCitation>
, see
<figureCitation id="1353028F8314FFF6FE443D1D1406F9C2" box="[440,534,1543,1565]" captionStart="Figure 20" captionStartId="30.[142,220,1753,1772]" captionTargetBox="[166,1398,199,1719]" captionTargetId="figure-93@30.[364,1196,612,1169]" captionTargetPageId="30" captionText="Figure 20. (a) Acanthodactylus longipes, male, MNHN 1997.4771, Azzefâl (Mauritania). (b) Acanthodactylus longipes, male, lectotype, BMNH 1946.8.4.31, Wargla (Algeria), close-up of head. (c) Acanthodactylus aureus, male, PHG 35, Sidi Ouassaï, near Massa (Morocco), picture by P. Geniez. (d) Acanthodactylus taghitensis, female, PHG 28, 5 km SSW of Taghit (Algeria), picture by M. Geniez." figureDoi="http://doi.org/10.5281/zenodo.5437195" httpUri="https://zenodo.org/record/5437195/files/figure.png" pageId="28" pageNumber="145">Fig. 20b</figureCitation>
). Large number of longitudinal rows of ventral plates (15 or more in 95% of the individuals, range: 13–19, mean: 16.1;
<tableCitation id="C6EA2BB18314FFF6FD673D5E14FEF984" box="[667,750,1604,1627]" captionStart="Table 2" captionStartId="5.[155,219,196,215]" captionText="Table 2. Means, standard deviations (in parentheses) and ranges of the quantitative variables used in this study. See Table 1 for variable definitions" pageId="28" pageNumber="145">Table 2</tableCitation>
) arranged in oblique rows. Large number of femoral pores (range 17–28, mean = 21.5;
<tableCitation id="C6EA2BB18314FFF6FDE83D981479F947" box="[532,617,1666,1688]" captionStart="Table 2" captionStartId="5.[155,219,196,215]" captionText="Table 2. Means, standard deviations (in parentheses) and ranges of the quantitative variables used in this study. See Table 1 for variable definitions" pageId="28" pageNumber="145">Table 2</tableCitation>
). Coloration distinctive: flanks have a mottled pattern which usually tends to disappear towards the vertebral area (
<figureCitation id="1353028F8314FFF6FF6A3DC416E4F92B" box="[150,244,1758,1780]" captionStart="Figure 20" captionStartId="30.[142,220,1753,1772]" captionTargetBox="[166,1398,199,1719]" captionTargetId="figure-93@30.[364,1196,612,1169]" captionTargetPageId="30" captionText="Figure 20. (a) Acanthodactylus longipes, male, MNHN 1997.4771, Azzefâl (Mauritania). (b) Acanthodactylus longipes, male, lectotype, BMNH 1946.8.4.31, Wargla (Algeria), close-up of head. (c) Acanthodactylus aureus, male, PHG 35, Sidi Ouassaï, near Massa (Morocco), picture by P. Geniez. (d) Acanthodactylus taghitensis, female, PHG 28, 5 km SSW of Taghit (Algeria), picture by M. Geniez." figureDoi="http://doi.org/10.5281/zenodo.5437195" httpUri="https://zenodo.org/record/5437195/files/figure.png" pageId="28" pageNumber="145">Fig. 20a</figureCitation>
). Females can have small, regularly disposed spots. Red spots can occur on the dorsum. Pileus weakly vermiculated with red. The combination of dorsal pattern and scale structure gives to the skin of
<taxonomicName id="4C6865898314FFF6FF723C43171AF8B2" authorityName="Boulenger" authorityYear="1918" box="[142,266,1880,1902]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FF723C43171AF8B2" box="[142,266,1880,1902]" italics="true" pageId="28" pageNumber="145">A. longipes</emphasis>
</taxonomicName>
a fragile and translucent aspect. This species is further characterized by an elongate and pointed snout compared to
<taxonomicName id="4C6865898314FFF6FB9C3BF91377FF27" authorityName="Boulenger" authorityYear="1918" box="[1120,1383,227,248]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="audouini">
<emphasis id="B91CC2188314FFF6FB9C3BF91377FF27" box="[1120,1383,227,248]" italics="true" pageId="28" pageNumber="145">A. scutellatus audouini</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6865898314FFF6FA8A3BF91599FEC9" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="dumerili">
<emphasis id="B91CC2188314FFF6FA8A3BF91599FEC9" italics="true" pageId="28" pageNumber="145">A. dumerili</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6865898314FFF6FC2A3A18129AFEC9" authorityName="Chabanaud" authorityYear="1918" box="[982,1162,257,279]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="senegalensis">
<emphasis id="B91CC2188314FFF6FC2A3A18129AFEC9" box="[982,1162,257,279]" italics="true" pageId="28" pageNumber="145">A. senegalensis</emphasis>
</taxonomicName>
. See
<taxonomicName id="4C6865898314FFF6FB263A18139EFEC9" authorityName="Chabanaud" authorityYear="1918" box="[1242,1422,257,279]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="senegalensis">
<emphasis id="B91CC2188314FFF6FB263A18139EFEC9" box="[1242,1422,257,279]" italics="true" pageId="28" pageNumber="145">A. senegalensis</emphasis>
</taxonomicName>
for additional differences from that species. Several scalation characters separate
<taxonomicName id="4C6865898314FFF6FB6F3A251306FE8B" authorityName="Boulenger" authorityYear="1918" box="[1171,1302,319,340]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FB6F3A251306FE8B" box="[1171,1302,319,340]" italics="true" pageId="28" pageNumber="145">A. longipes</emphasis>
</taxonomicName>
from the broadly sympatric
<taxonomicName id="4C6865898314FFF6FBF83A441299FEAD" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[1028,1161,349,371]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="dumerili">
<emphasis id="B91CC2188314FFF6FBF83A441299FEAD" box="[1028,1161,349,371]" italics="true" pageId="28" pageNumber="145">A. dumerili</emphasis>
</taxonomicName>
and
<emphasis id="B91CC2188314FFF6FB353A44137CFEAD" box="[1225,1388,349,371]" italics="true" pageId="28" pageNumber="145">A. s. audouini</emphasis>
. A strongly fragmented fourth supraocular is found in 73% of the
<taxonomicName id="4C6865898314FFF6FC583A81120FFE6F" authorityName="Boulenger" authorityYear="1918" box="[932,1055,411,432]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FC583A81120FFE6F" box="[932,1055,411,432]" italics="true" pageId="28" pageNumber="145">A. longipes</emphasis>
</taxonomicName>
individuals but in 14% of the
<taxonomicName id="4C6865898314FFF6FA8A3A811599FE11" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="dumerili">
<emphasis id="B91CC2188314FFF6FA8A3A811599FE11" italics="true" pageId="28" pageNumber="145">A. dumerili</emphasis>
</taxonomicName>
and 37% of the
<emphasis id="B91CC2188314FFF6FBB53AA012F8FE11" box="[1097,1256,441,463]" italics="true" pageId="28" pageNumber="145">A. s. audouini</emphasis>
. Granules are often inserted between the parietal plates in
<taxonomicName id="4C6865898314FFF6FABD3AC21570FDD3" authorityName="Boulenger" authorityYear="1918" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FABD3AC21570FDD3" italics="true" pageId="28" pageNumber="145">A. longipes</emphasis>
</taxonomicName>
, which is exceptional in
<emphasis id="B91CC2188314FFF6FB6E3AED1323FDD3" box="[1170,1331,503,524]" italics="true" pageId="28" pageNumber="145">A. s. audouini</emphasis>
and
<taxonomicName id="4C6865898314FFF6FA8A3AED1599FDF5" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="dumerili">
<emphasis id="B91CC2188314FFF6FA8A3AED1599FDF5" italics="true" pageId="28" pageNumber="145">A. dumerili</emphasis>
</taxonomicName>
. The number of longitudinal rows of ventrals is often nearly diagnostic: most
<taxonomicName id="4C6865898314FFF6FB72392E1319FD96" authorityName="Boulenger" authorityYear="1918" box="[1166,1289,564,585]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FB72392E1319FD96" box="[1166,1289,564,585]" italics="true" pageId="28" pageNumber="145">A. longipes</emphasis>
</taxonomicName>
(81% of the specimens) have 16 ventrals rows or more, which is extremely rare in
<emphasis id="B91CC2188314FFF6FC0839681280FD59" box="[1012,1168,625,647]" italics="true" pageId="28" pageNumber="145">A. s. audouini</emphasis>
(1% of the specimens) or
<taxonomicName id="4C6865898314FFF6FCBA398A15D9FD7A" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[838,969,656,677]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="dumerili">
<emphasis id="B91CC2188314FFF6FCBA398A15D9FD7A" box="[838,969,656,677]" italics="true" pageId="28" pageNumber="145">A. dumerili</emphasis>
</taxonomicName>
(less than 1% of the specimens).
<taxonomicName id="4C6865898314FFF6FAB0398A121EFD1B" authorityName="Boulenger" authorityYear="1918" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FAB0398A121EFD1B" italics="true" pageId="28" pageNumber="145">Acanthodactylus longipes</emphasis>
</taxonomicName>
is the only species to possess dorsal scales small, elongate and smooth except in the vertebral area. The dorsal scales in the vertebral area are only weekly keeled, a further distinction from most
<emphasis id="B91CC2188314FFF6FC9838301210FCE1" box="[868,1024,809,831]" italics="true" pageId="28" pageNumber="145">A. s. audouini</emphasis>
and
<taxonomicName id="4C6865898314FFF6FBC5383012AAFCE1" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[1081,1210,809,831]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="dumerili">
<emphasis id="B91CC2188314FFF6FBC5383012AAFCE1" box="[1081,1210,809,831]" italics="true" pageId="28" pageNumber="145">A. dumerili</emphasis>
</taxonomicName>
: 84% of the
<taxonomicName id="4C6865898314FFF6FAB838301570FC81" authorityName="Boulenger" authorityYear="1918" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FAB838301570FC81" italics="true" pageId="28" pageNumber="145">A. longipes</emphasis>
</taxonomicName>
individuals have CARE code 2 or (rarely) 1, whereas less than 2% of the
<taxonomicName id="4C6865898314FFF6FB7B387D131DFCA3" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[1159,1293,871,892]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="dumerili">
<emphasis id="B91CC2188314FFF6FB7B387D131DFCA3" box="[1159,1293,871,892]" italics="true" pageId="28" pageNumber="145">A. dumerili</emphasis>
</taxonomicName>
specimens have CARE code 2 or 1. The typical dorsal pattern is also characteristic. It should be noted, however, that some
<taxonomicName id="4C6865898314FFF6FC9438D915F4FC07" authorityName="Boulenger" authorityYear="1918" box="[872,996,963,984]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FC9438D915F4FC07" box="[872,996,963,984]" italics="true" pageId="28" pageNumber="145">A. longipes</emphasis>
</taxonomicName>
females present a uniformly pinkish coloration with small dark spots, making them similar to
<taxonomicName id="4C6865898314FFF6FCBE3F1A15D3FBCA" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[834,963,1024,1045]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="dumerili">
<emphasis id="B91CC2188314FFF6FCBE3F1A15D3FBCA" box="[834,963,1024,1045]" italics="true" pageId="28" pageNumber="145">A. dumerili</emphasis>
</taxonomicName>
or
<emphasis id="B91CC2188314FFF6FC1B3F1A1292FBCA" box="[999,1154,1024,1046]" italics="true" pageId="28" pageNumber="145">A. s. audouini</emphasis>
, although their pattern is less contrasting than in these species. In conclusion, although no single character is fully diagnostic between
<taxonomicName id="4C6865898314FFF6FC703F461218FBAE" authorityName="Boulenger" authorityYear="1918" box="[908,1032,1116,1137]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B91CC2188314FFF6FC703F461218FBAE" box="[908,1032,1116,1137]" italics="true" pageId="28" pageNumber="145">A. longipes</emphasis>
</taxonomicName>
and the sympatric species, a combination of several scalation and coloration features will enable to identify the vast majority of specimens. In ambiguous cases, the elongate and pointed shape of the snout and the structure of the dorsal scales are often useful, although occasional individuals remain impossible to identify safely. It should be stressed than other characters proposed by earlier authors (number of supralabials in contact with the subocular, length of hindlegs) proved to be useless.
</paragraph>
<paragraph id="8BD71E0A8314FFF6FCC33E8415D0F992" blockId="28.[806,1422,1438,1613]" pageId="28" pageNumber="145">
<emphasis id="B91CC2188314FFF6FCC33E841258FA6B" box="[831,1096,1438,1460]" italics="true" pageId="28" pageNumber="145">Geographical variation</emphasis>
. The subspecies
<emphasis id="B91CC2188314FFF6FADE3E851397FA6B" box="[1314,1415,1439,1460]" italics="true" pageId="28" pageNumber="145">panousei</emphasis>
, described from south-eastern
<collectingCountry id="F37F5E9A8314FFF6FB833EA712CFFA0D" box="[1151,1247,1469,1490]" name="Morocco" pageId="28" pageNumber="145">Morocco</collectingCountry>
, does not seem to be valid (
<collectingCountry id="F37F5E9A8314FFF6FC503EC11202FA2E" box="[940,1042,1499,1521]" name="El Salvador" pageId="28" pageNumber="145">Salvador</collectingCountry>
, 1982; own results). In some populations from the eastern part of the species distribution, specimens tend to have more strongly carinate dorsal scales.
</paragraph>
</subSubSection>
</treatment>
</document>
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