300 lines
36 KiB
XML
300 lines
36 KiB
XML
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<mods:title id="D6517C9A5424FAA0498E72CA67F06F68">A multivariate analysis of the fringe-toed lizards of the Acanthodactylus scutellatus group (Squamata: Lacertidae): systematic and biogeographical implications</mods:title>
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<mods:namePart id="24B35A3E1AF6B0E8DC9B28A9D12E5944">Ineich, Ivan</mods:namePart>
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<treatment id="03C1AF1C8314FFF5FCA43D8314B1F9B1" ID-DOI="http://doi.org/10.5281/zenodo.5490912" ID-GBIF-Taxon="185447197" ID-Zenodo-Dep="5490912" LSID="urn:lsid:plazi:treatment:03C1AF1C8314FFF5FCA43D8314B1F9B1" httpUri="http://treatment.plazi.org/id/03C1AF1C8314FFF5FCA43D8314B1F9B1" lastPageId="31" lastPageNumber="148" pageId="28" pageNumber="145">
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<subSubSection id="C3724D818314FFF6FCA43D83134CF96E" box="[856,1372,1689,1713]" pageId="28" pageNumber="145" type="nomenclature">
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<paragraph id="8BD71E0A8314FFF6FCA43D83134CF96E" blockId="28.[856,1372,1689,1713]" box="[856,1372,1689,1713]" pageId="28" pageNumber="145">
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<heading id="D09FA9668314FFF6FCA43D83134CF96E" box="[856,1372,1689,1713]" centered="true" fontSize="9" level="2" pageId="28" pageNumber="145" reason="2">
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<taxonomicName id="4C6865898314FFF6FCA43D83134CF96E" authority="GUNTHER, 1903" authorityName="Gunther" authorityYear="1903" box="[856,1372,1689,1713]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="species" species="aureus">
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<emphasis id="B91CC2188314FFF6FCA43D83128CF96F" box="[856,1180,1689,1713]" italics="true" pageId="28" pageNumber="145">ACANTHODACTYLUS AUREUS</emphasis>
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<bibRefCitation id="EFF963FB8314FFF6FB583D83134CF96E" author="Gunther A" box="[1188,1372,1689,1713]" pageId="28" pageNumber="145" pagination="298 - 299" refId="ref22138" refString="Gunther A. 1903. Reptiles from Rio de Oro, Western Sahara. Novitates Zoologicae 10: 298 - 299." type="journal article" year="1903">GÜNTHER, 1903</bibRefCitation>
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3724D818314FFF7FCDA3DD817A6F9B3" lastPageId="29" lastPageNumber="146" pageId="28" pageNumber="145" type="reference_group">
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<paragraph id="8BD71E0A8314FFF7FCDA3DD81491FA3D" blockId="28.[806,1423,1730,1905]" lastBlockId="29.[155,771,1177,1506]" lastPageId="29" lastPageNumber="146" pageId="28" pageNumber="145">
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<treatmentCitationGroup id="AB7839248314FFF7FCDA3DD81491FA3D" lastPageId="29" lastPageNumber="146" pageId="28" pageNumber="145">
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<treatmentCitation id="0AC9381B8314FFF6FCDA3DD81542F929" author="Gunther A" page="298" pageId="28" pageNumber="145" year="1903">
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<taxonomicName id="4C6865898314FFF6FCDA3DD81542F929" authority="Gunther, 1903: 298" authorityName="Gunther" authorityPageNumber="298" authorityYear="1903" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="145" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="aureus">
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<emphasis id="B91CC2188314FFF6FCDA3DD812DAF907" box="[806,1226,1730,1752]" italics="true" pageId="28" pageNumber="145">Acanthodactylus scutellatus aureus</emphasis>
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<bibRefCitation id="EFF963FB8314FFF6FB273DD81542F929" author="Gunther A" pageId="28" pageNumber="145" pagination="298 - 299" refId="ref22138" refString="Gunther A. 1903. Reptiles from Rio de Oro, Western Sahara. Novitates Zoologicae 10: 298 - 299." type="journal article" year="1903">Günther, 1903: 298</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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. Name-bearing type: Günther’s original description was based on ‘a considerable number of this species’. More than
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<specimenCount id="9D6ED5838314FFF6FC223C041264F8EB" box="[990,1140,1822,1844]" pageId="28" pageNumber="145" type="generic">30 specimens</specimenCount>
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were examined from
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<collectingCountry id="F37F5E9A8314FFF6FA9B3C041569F88D" name="Western Sahara" pageId="28" pageNumber="145">Rio de Oro</collectingCountry>
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(former
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<collectingCountry id="F37F5E9A8314FFF6FC163C2712B9F88D" box="[1002,1193,1852,1874]" name="Western Sahara" pageId="28" pageNumber="145">Western Sahara</collectingCountry>
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) and several from Southern
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<collectingCountry id="F37F5E9A8314FFF6FC623C4115E5F8AE" box="[926,1013,1883,1905]" name="Algeria" pageId="28" pageNumber="145">Algeria</collectingCountry>
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. The
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<typeStatus id="54D3A0A88314FFF6FBC33C4612B4F8AE" box="[1087,1188,1884,1905]" pageId="28" pageNumber="145" type="syntype">syntypes</typeStatus>
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listed by
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<collectingCountry id="F37F5E9A8314FFF6FAD53C41139EF8AE" box="[1321,1422,1883,1905]" name="El Salvador" pageId="28" pageNumber="145">Salvador</collectingCountry>
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(1982) are thus clearly a part of the types only: BMNH 1946.8.5.2-26, USNM 33109-33111, MNHN 1918.11- 12. The
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<typeStatus id="54D3A0A88315FFF7FEF93FCD177AFB33" box="[261,362,1239,1260]" pageId="29" pageNumber="146" type="syntype">syntypes</typeStatus>
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from Southern
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<collectingCountry id="F37F5E9A8315FFF7FDC93FCC1499FB33" box="[565,649,1238,1260]" name="Algeria" pageId="29" pageNumber="146">Algeria</collectingCountry>
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are most likely specimens of
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<emphasis id="B91CC2188315FFF7FE743FEF143CFAD5" box="[392,556,1269,1290]" italics="true" pageId="29" pageNumber="146">
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<taxonomicName id="4C6865898315FFF7FE743FEF1439FAD5" authorityName="Geniez & Foucart" authorityYear="1995" box="[392,553,1269,1290]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="146" phylum="Chordata" rank="species" species="taghitensis">A. taghitensis</taxonomicName>
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,
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</emphasis>
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but we could not find them. It is thus important to select a
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<typeStatus id="54D3A0A88315FFF7FD673E0E1511FAF5" box="[667,769,1300,1322]" pageId="29" pageNumber="146" type="lectotype">lectotype</typeStatus>
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which belongs to
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<taxonomicName id="4C6865898315FFF7FE8D3E2917F3FA97" authorityName="Gunther" authorityYear="1903" box="[369,483,1331,1352]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="146" phylum="Chordata" rank="species" species="aureus">
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<emphasis id="B91CC2188315FFF7FE8D3E2917F3FA97" box="[369,483,1331,1352]" italics="true" pageId="29" pageNumber="146">A. aureus</emphasis>
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</taxonomicName>
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as understood now. We select as
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<typeStatus id="54D3A0A88315FFF7FEF43E4B177EFAB8" box="[264,366,1361,1383]" pageId="29" pageNumber="146" type="lectotype">lectotype</typeStatus>
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the specimen MNHN 1918-11, an adult male from ‘Villa Cisneros’ [= Dakhla],
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<collectingCountry id="F37F5E9A8315FFF7FD583E6A16E0FA7B" name="Western Sahara" pageId="29" pageNumber="146">Western Sahara</collectingCountry>
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. Type locality: originally
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<collectingCountry id="F37F5E9A8315FFF7FDF53E94146FFA7B" box="[521,639,1422,1444]" name="Western Sahara" pageId="29" pageNumber="146">Rio de Oro</collectingCountry>
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and Southern
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<collectingCountry id="F37F5E9A8315FFF7FF353EB7170FFA1C" box="[201,287,1453,1475]" name="Algeria" pageId="29" pageNumber="146">Algeria</collectingCountry>
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, restricted to ‘Villa Cisneros’ [= Dakhla],
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<collectingCountry id="F37F5E9A8315FFF7FF673ED61747FA3D" box="[155,343,1484,1506]" name="Western Sahara" pageId="29" pageNumber="146">Western Sahara</collectingCountry>
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, by
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<typeStatus id="54D3A0A88315FFF7FE7A3ED617FCFA3D" box="[390,492,1484,1506]" pageId="29" pageNumber="146" type="lectotype">lectotype</typeStatus>
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designation.
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</treatmentCitationGroup>
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</paragraph>
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<caption id="DF174E828315FFF7FF673F131458FBE6" ID-DOI="http://doi.org/10.5281/zenodo.5437193" ID-Zenodo-Dep="5437193" httpUri="https://zenodo.org/record/5437193/files/figure.png" pageId="29" pageNumber="146" startId="29.[155,233,1033,1052]" targetBox="[157,1431,198,996]" targetPageId="29">
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<paragraph id="8BD71E0A8315FFF7FF673F131458FBE6" blockId="29.[155,1434,1033,1082]" pageId="29" pageNumber="146">
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<emphasis id="B91CC2188315FFF7FF673F131700FBC3" bold="true" box="[155,272,1033,1052]" pageId="29" pageNumber="146">Figure 19.</emphasis>
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Geographical distribution of
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<taxonomicName id="4C6865898315FFF7FDBB3F131540FBC3" authorityName="Boulenger" authorityYear="1918" box="[583,848,1033,1052]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="146" phylum="Chordata" rank="species" species="longipes">
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<emphasis id="B91CC2188315FFF7FDBB3F131540FBC3" box="[583,848,1033,1052]" italics="true" pageId="29" pageNumber="146">Acanthodactylus longipes</emphasis>
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</taxonomicName>
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. Data from Salvador (1982),
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<bibRefCitation id="EFF963FB8315FFF7FB873F131355FBC3" author="Baha El Din SM" box="[1147,1349,1033,1052]" pageId="29" pageNumber="146" pagination="18 - 27" refId="ref21524" refString="Baha El Din SM. 1994. A contribution to the herpetology of Sinai. British Herpetological Society Bulletin 48: 18 - 27." type="journal article" year="1994">Baha El Din (1994)</bibRefCitation>
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,
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<bibRefCitation id="EFF963FB8315FFF7FAB33F13173EFBE6" author="Bons J & Geniez Ph" pageId="29" pageNumber="146" refId="ref21662" refString="Bons J, Geniez Ph. 1996. Amphibiens et Reptiles du Maroc, Sahara Occidental compris. Atlas biogeographique. Barcelona: Asociacion Herpetologica Espanola." type="book" year="1996">Bons & Geniez (1996)</bibRefCitation>
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,
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<bibRefCitation id="EFF963FB8315FFF7FEC53F3D17DDFBE6" author="Nouira S" box="[313,461,1062,1082]" pageId="29" pageNumber="146" refId="ref22685" refString="Nouira S. 1996. Systematique, ecologie et biogeographie evolutive des Lacertidae (Reptilia, Sauria). Importance dans l'herpetofaune tunisienne. DPhil Thesis, University of Tunis." type="book" year="1996">Nouïra (1996)</bibRefCitation>
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, this study.
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</paragraph>
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||
</caption>
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<paragraph id="8BD71E0A8315FFF7FF483EE017A6F9B3" blockId="29.[155,770,1530,1644]" pageId="29" pageNumber="146">
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||
<emphasis id="B91CC2188315FFF7FF483EE0172EF9D0" box="[180,318,1530,1551]" italics="true" pageId="29" pageNumber="146">Chresonyms</emphasis>
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.
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<taxonomicName id="4C6865898315FFF7FEA73EE01752F9F1" authority="Gunther, 1903" authorityName="Gunther" authorityYear="1903" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="146" phylum="Chordata" rank="subSpecies" species="inornatus" subSpecies="aureus">
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<emphasis id="B91CC2188315FFF7FEA73EE01512F9CF" box="[347,770,1530,1552]" italics="true" pageId="29" pageNumber="146">Acanthodactylus inornatus aureus</emphasis>
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<bibRefCitation id="EFF963FB8315FFF7FF673D021752F9F1" author="Gunther A" box="[155,322,1560,1582]" pageId="29" pageNumber="146" pagination="298 - 299" refId="ref22138" refString="Gunther A. 1903. Reptiles from Rio de Oro, Western Sahara. Novitates Zoologicae 10: 298 - 299." type="journal article" year="1903">Günther, 1903</bibRefCitation>
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</taxonomicName>
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:
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<bibRefCitation id="EFF963FB8315FFF7FEB23D031479F9F1" author="Bons J & Girot B" box="[334,617,1560,1582]" pageId="29" pageNumber="146" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons & Girot, 1964: 330</bibRefCitation>
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.
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<taxonomicName id="4C6865898315FFF7FD883D0217E3F993" authority="Gunther, 1903" authorityName="Gunther" authorityYear="1903" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="146" phylum="Chordata" rank="species" species="aureus">
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||
<emphasis id="B91CC2188315FFF7FD883D021721F992" italics="true" pageId="29" pageNumber="146">Acanthodactylus aureus</emphasis>
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||
<bibRefCitation id="EFF963FB8315FFF7FEBF3D2D17E3F993" author="Gunther A" box="[323,499,1591,1613]" pageId="29" pageNumber="146" pagination="298 - 299" refId="ref22138" refString="Gunther A. 1903. Reptiles from Rio de Oro, Western Sahara. Novitates Zoologicae 10: 298 - 299." type="journal article" year="1903">Günther, 1903</bibRefCitation>
|
||
</taxonomicName>
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||
:
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<collectingCountry id="F37F5E9A8315FFF7FDF53D2D147EF992" box="[521,622,1591,1613]" name="El Salvador" pageId="29" pageNumber="146">Salvador</collectingCountry>
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, 1982: 122;
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<bibRefCitation id="EFF963FB8315FFF7FF673D4C1776F9B4" author="Arnold EN" box="[155,358,1622,1644]" pageId="29" pageNumber="146" pagination="291 - 339" refId="ref21308" refString="Arnold EN. 1983. Osteology, genitalia and the relationships of Acanthodactylus (Reptilia: Lacertidae). Bulletin of the British Museum (Natural History) Zoology 44: 291 - 339." type="journal article" year="1983">Arnold, 1983: 328</bibRefCitation>
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(part).
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</paragraph>
|
||
</subSubSection>
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<subSubSection id="C3724D818315FFF5FF483D9E14B1F9B1" lastPageId="31" lastPageNumber="148" pageId="29" pageNumber="146" type="distribution">
|
||
<paragraph id="8BD71E0A8315FFF7FF483D9E136DFAF5" blockId="29.[155,771,1668,1904]" lastBlockId="29.[819,1435,1177,1322]" pageId="29" pageNumber="146">
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||
<emphasis id="B91CC2188315FFF7FF483D9E172FF946" box="[180,319,1668,1689]" italics="true" pageId="29" pageNumber="146">Distribution</emphasis>
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||
(
|
||
<figureCitation id="1353028F8315FFF7FEAF3D9E17B2F945" box="[339,418,1668,1690]" captionStart="Figure 21" captionStartId="31.[155,233,1033,1052]" captionTargetBox="[157,1431,195,999]" captionTargetId="figure-416@31.[155,1432,191,1000]" captionTargetPageId="31" captionText="Figure 21. Geographical distribution of Acanthodactylus aureus (circles) and Acanthodactylus taghitensis (triangles). Data from Böhme (1978), Salvador (1982), Geniez & Foucart (1995), Bons & Geniez (1996), Hasi et al. (1998), Donaire et al. (2000), this study." figureDoi="http://doi.org/10.5281/zenodo.5437197" httpUri="https://zenodo.org/record/5437197/files/figure.png" pageId="29" pageNumber="146">Fig. 21</figureCitation>
|
||
). Saharan oceanic fringe from
|
||
<collectingCountry id="F37F5E9A8315FFF7FF673DB816F9F967" box="[155,233,1698,1720]" name="Morocco" pageId="29" pageNumber="146">Agadir</collectingCountry>
|
||
(
|
||
<collectingCountry id="F37F5E9A8315FFF7FF093DB91745F966" box="[245,341,1699,1721]" name="Morocco" pageId="29" pageNumber="146">Morocco</collectingCountry>
|
||
) to the Cap Blanc Peninsula (
|
||
<collectingCountry id="F37F5E9A8315FFF7FD5F3DB916FCF908" name="Western Sahara" pageId="29" pageNumber="146">Western Sahara</collectingCountry>
|
||
/
|
||
<collectingCountry id="F37F5E9A8315FFF7FF093DDB176AF908" box="[245,378,1729,1751]" name="Mauritania" pageId="29" pageNumber="146">Mauritania</collectingCountry>
|
||
). The species is further encountered at two points in
|
||
<collectingCountry id="F37F5E9A8315FFF7FE6D3DFA1404F929" box="[401,532,1760,1782]" name="Mauritania" pageId="29" pageNumber="146">Mauritania</collectingCountry>
|
||
(
|
||
<collectingCountry id="F37F5E9A8315FFF7FDDE3DFA1499F929" box="[546,649,1760,1782]" name="El Salvador" pageId="29" pageNumber="146">Salvador</collectingCountry>
|
||
, 1982) and three stations of the
|
||
<collectingCountry id="F37F5E9A8315FFF7FE573DE41415F8CB" box="[427,517,1790,1812]" name="Senegal" pageId="29" pageNumber="146">Senegal</collectingCountry>
|
||
coast (
|
||
<bibRefCitation id="EFF963FB8315FFF7FD9A3DE414EFF8CB" author="Bohme W" box="[614,767,1790,1812]" pageId="29" pageNumber="146" pagination="360 - 416" refId="ref21607" refString="Bohme W. 1978. Zur Herpetofaunistik des Senegal. Bonner Zoologische Beitrage 29: 360 - 416." type="journal article" year="1978">Böhme, 1978</bibRefCitation>
|
||
;
|
||
<collectingCountry id="F37F5E9A8315FFF7FF673C071710F8EC" box="[155,256,1821,1843]" name="El Salvador" pageId="29" pageNumber="146">Salvador</collectingCountry>
|
||
, 1982). It reaches as far as
|
||
<quantity id="4C90B3EF8315FFF7FDC13C071483F8EC" box="[573,659,1821,1843]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="146" unit="km" value="200.0">200 km</quantity>
|
||
inland in
|
||
<collectingCountry id="F37F5E9A8315FFF7FF673C261744F88D" box="[155,340,1852,1874]" name="Western Sahara" pageId="29" pageNumber="146">Western Sahara</collectingCountry>
|
||
(pers. obs.). The animals from Fderik (
|
||
<collectingCountry id="F37F5E9A8315FFF7FF5F3C401737F8AF" box="[163,295,1882,1904]" name="Mauritania" pageId="29" pageNumber="146">Mauritania</collectingCountry>
|
||
) are referable to
|
||
<taxonomicName id="4C6865898315FFF7FE0D3C40157BFB70" authorityName="Geniez & Foucart" authorityYear="1995" box="[497,875,1178,1903]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="146" phylum="Chordata" rank="species" species="taghitensis">
|
||
<emphasis id="B91CC2188315FFF7FE0D3C40157BFB70" box="[497,875,1178,1903]" italics="true" pageId="29" pageNumber="146">Acanthodactylus taghitensis</emphasis>
|
||
</taxonomicName>
|
||
. The Atar locality (specimen BMNH 1982.294) is wrongly positioned (far inland) in
|
||
<collectingCountry id="F37F5E9A8315FFF7FB4A3FA2130BFB11" box="[1206,1307,1208,1230]" name="El Salvador" pageId="29" pageNumber="146">Salvador</collectingCountry>
|
||
(1982) and
|
||
<bibRefCitation id="EFF963FB8315FFF7FCCF3FCC15C3FB32" author="Arnold EN" box="[819,979,1238,1261]" pageId="29" pageNumber="146" pagination="291 - 339" refId="ref21308" refString="Arnold EN. 1983. Osteology, genitalia and the relationships of Acanthodactylus (Reptilia: Lacertidae). Bulletin of the British Museum (Natural History) Zoology 44: 291 - 339." type="journal article" year="1983">Arnold (1983)</bibRefCitation>
|
||
. It is in fact situated near Nouadhibou, i.e. near the coast.
|
||
<taxonomicName id="4C6865898315FFF7FBF23FEF126AFAD4" authorityName="Gunther" authorityYear="1903" box="[1038,1146,1269,1291]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="146" phylum="Chordata" rank="species" species="aureus">
|
||
<emphasis id="B91CC2188315FFF7FBF23FEF126AFAD4" box="[1038,1146,1269,1291]" italics="true" pageId="29" pageNumber="146">A. aureus</emphasis>
|
||
</taxonomicName>
|
||
is the most common lacertid on the whole oceanic side of
|
||
<collectingCountry id="F37F5E9A8315FFF7FB403E0E1369FAF5" box="[1212,1401,1300,1322]" name="Western Sahara" pageId="29" pageNumber="146">Western Sahara</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD71E0A8315FFF5FCB03E4816E2F9E6" blockId="29.[819,1435,1361,1904]" lastBlockId="31.[155,771,1172,1593]" lastPageId="31" lastPageNumber="148" pageId="29" pageNumber="146">
|
||
<emphasis id="B91CC2188315FFF7FCB03E4815ACFAB8" box="[844,956,1362,1383]" italics="true" pageId="29" pageNumber="146">Diagnosis</emphasis>
|
||
. A species of variable size according to populations (reaching a maximum snout-vent length of
|
||
<quantity id="4C90B3EF8315FFF7FCB33E94158FFA7B" box="[847,927,1422,1444]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="29" pageNumber="146" unit="mm" value="65.0">65 mm</quantity>
|
||
, mean: 53.65). Only two supralabials in contact with the subocular in 99% of the specimens, as a result of the fusion of the third and fourth (rarely the fourth and fifth) supralabials (
|
||
<tableCitation id="C6EA2BB18315FFF7FB6A3EF012FBF9DE" box="[1174,1259,1514,1537]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="29" pageNumber="146">Table 3</tableCitation>
|
||
). Dorsal scales relatively large, not very numerous (range: 38–59, mean: 46.7;
|
||
<tableCitation id="C6EA2BB18315FFF7FC343D32120CF9E1" box="[968,1052,1576,1598]" captionStart="Table 2" captionStartId="5.[155,219,196,215]" captionText="Table 2. Means, standard deviations (in parentheses) and ranges of the quantitative variables used in this study. See Table 1 for variable definitions" pageId="29" pageNumber="146">Table 2</tableCitation>
|
||
), slightly larger on the dorsum than flanks, pyramidal, and obtusely carinate (CARE code 3 or
|
||
<quantity id="4C90B3EF8315FFF7FC5F3D7F15C0F9A4" box="[931,976,1637,1659]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="29" pageNumber="146" unit="in" value="4.0">4 in</quantity>
|
||
99% of the individuals;
|
||
<tableCitation id="C6EA2BB18315FFF7FB173D7F132EF9A4" box="[1259,1342,1637,1659]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="29" pageNumber="146">Table 3</tableCitation>
|
||
). Fewer than two rows of supraciliary granules in 97% of the individuals (
|
||
<tableCitation id="C6EA2BB18315FFF7FC393DB8120AF966" box="[965,1050,1698,1721]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="29" pageNumber="146">Table 3</tableCitation>
|
||
). Generally 14 or (less often) 13 longitudinal rows of ventrals (in 77% of the specimens, range: 12–17; mean: 13.9;
|
||
<tableCitation id="C6EA2BB18315FFF7FB823DFA12C3F929" box="[1150,1235,1760,1782]" captionStart="Table 2" captionStartId="5.[155,219,196,215]" captionText="Table 2. Means, standard deviations (in parentheses) and ranges of the quantitative variables used in this study. See Table 1 for variable definitions" pageId="29" pageNumber="146">Table 2</tableCitation>
|
||
). Femoral pores rather numerous (range: 19–26, mean: 21.2;
|
||
<tableCitation id="C6EA2BB18315FFF7FAC43DE4139EF8CA" box="[1336,1422,1790,1813]" captionStart="Table 2" captionStartId="5.[155,219,196,215]" captionText="Table 2. Means, standard deviations (in parentheses) and ranges of the quantitative variables used in this study. See Table 1 for variable definitions" pageId="29" pageNumber="146">Table 2</tableCitation>
|
||
). Dorsal coloration distinctive, comprising dark rectangular blotches longitudinally aligned over light longitudinal lines. In old males, however, the dark blotches and the light lines can produce a mottled or even reticulate pattern. A golden yellow tone appears on the dorsum of males during reproduction. Further characterized by a concave forehead following a short but strongly acuminate snout. Supralabials sometimes uniformly greyish or purplish-blue. Easily separated from
|
||
<taxonomicName id="4C6865898317FFF5FF1F3E57176EFABE" baseAuthorityName="Audouin" baseAuthorityYear="1827" box="[227,382,1356,1378]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="148" phylum="Chordata" rank="species" species="scutellatus">
|
||
<emphasis id="B91CC2188317FFF5FF1F3E57176EFABE" box="[227,382,1356,1378]" italics="true" pageId="31" pageNumber="148">A. scutellatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6865898317FFF5FE723E57142BFABE" authorityName="Chabanaud" authorityYear="1918" box="[398,571,1356,1378]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="148" phylum="Chordata" rank="species" species="senegalensis">
|
||
<emphasis id="B91CC2188317FFF5FE723E57142BFABE" box="[398,571,1356,1378]" italics="true" pageId="31" pageNumber="148">A. senegalensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6865898317FFF5FDB73E5714DEFABE" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[587,718,1356,1378]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="148" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC2188317FFF5FDB73E5714DEFABE" box="[587,718,1356,1378]" italics="true" pageId="31" pageNumber="148">A. dumerili</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6865898317FFF5FF673E711732FA5F" authorityName="Boulenger" authorityYear="1918" box="[155,290,1387,1408]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="148" phylum="Chordata" rank="species" species="longipes">
|
||
<emphasis id="B91CC2188317FFF5FF673E711732FA5F" box="[155,290,1387,1408]" italics="true" pageId="31" pageNumber="148">A. longipes</emphasis>
|
||
</taxonomicName>
|
||
by the peculiar conformation of the supralabial scales bordering the subocular. The exceptional individuals lacking this feature can be recognized by the typical habitus of the species, resulting from its distinctive colour pattern and peculiar head profile. For distinction from
|
||
<taxonomicName id="4C6865898317FFF5FE0C3D1F1480F9C6" authorityName="Geniez & Foucart" authorityYear="1995" box="[496,656,1540,1562]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="148" phylum="Chordata" rank="species" species="taghitensis">
|
||
<emphasis id="B91CC2188317FFF5FE0C3D1F1480F9C6" box="[496,656,1540,1562]" italics="true" pageId="31" pageNumber="148">A. taghitensis</emphasis>
|
||
</taxonomicName>
|
||
, see that species.
|
||
</paragraph>
|
||
<caption id="DF174E828316FFF4FF723DC317CCF89B" ID-DOI="http://doi.org/10.5281/zenodo.5437195" ID-Zenodo-Dep="5437195" httpUri="https://zenodo.org/record/5437195/files/figure.png" pageId="30" pageNumber="147" startId="30.[142,220,1753,1772]" targetBox="[166,1398,199,1719]" targetPageId="30">
|
||
<paragraph id="8BD71E0A8316FFF4FF723DC317CCF89B" blockId="30.[142,1423,1753,1861]" pageId="30" pageNumber="147">
|
||
<emphasis id="B91CC2188316FFF4FF723DC31714F933" bold="true" box="[142,260,1753,1772]" pageId="30" pageNumber="147">Figure 20.</emphasis>
|
||
(a)
|
||
<taxonomicName id="4C6865898316FFF4FEC93DC31453F933" authorityName="Boulenger" authorityYear="1918" box="[309,579,1753,1772]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="147" phylum="Chordata" rank="species" species="longipes">
|
||
<emphasis id="B91CC2188316FFF4FEC93DC31453F933" box="[309,579,1753,1772]" italics="true" pageId="30" pageNumber="147">Acanthodactylus longipes</emphasis>
|
||
</taxonomicName>
|
||
, male, MNHN 1997.4771, Azzefâl (Mauritania). (b)
|
||
<taxonomicName id="4C6865898316FFF4FB863DC31398F933" authorityName="Boulenger" authorityYear="1918" box="[1146,1416,1753,1772]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="147" phylum="Chordata" rank="species" species="longipes">
|
||
<emphasis id="B91CC2188316FFF4FB863DC31398F933" box="[1146,1416,1753,1772]" italics="true" pageId="30" pageNumber="147">Acanthodactylus longipes</emphasis>
|
||
</taxonomicName>
|
||
, male, lectotype, BMNH 1946.8.4.31, Wargla (Algeria), close-up of head. (c)
|
||
<taxonomicName id="4C6865898316FFF4FC4E3DEC12A0F8D6" authorityName="Gunther" authorityYear="1903" box="[946,1200,1782,1801]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="147" phylum="Chordata" rank="species" species="aureus">
|
||
<emphasis id="B91CC2188316FFF4FC4E3DEC12A0F8D6" box="[946,1200,1782,1801]" italics="true" pageId="30" pageNumber="147">Acanthodactylus aureus</emphasis>
|
||
</taxonomicName>
|
||
, male, PHG 35, Sidi Ouassaï, near Massa (Morocco), picture by P. Geniez. (d)
|
||
<taxonomicName id="4C6865898316FFF4FD203C0E1212F8F8" authorityName="Geniez & Foucart" authorityYear="1995" box="[732,1026,1812,1831]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="147" phylum="Chordata" rank="species" species="taghitensis">
|
||
<emphasis id="B91CC2188316FFF4FD203C0E1212F8F8" box="[732,1026,1812,1831]" italics="true" pageId="30" pageNumber="147">Acanthodactylus taghitensis</emphasis>
|
||
</taxonomicName>
|
||
, female, PHG 28, 5 km SSW of Taghit (Algeria), picture by M. Geniez.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF174E828317FFF5FF673F131748FB89" ID-DOI="http://doi.org/10.5281/zenodo.5437197" ID-Zenodo-Dep="5437197" httpUri="https://zenodo.org/record/5437197/files/figure.png" pageId="31" pageNumber="148" startId="31.[155,233,1033,1052]" targetBox="[157,1431,195,999]" targetPageId="31">
|
||
<paragraph id="8BD71E0A8317FFF5FF673F131748FB89" blockId="31.[155,1435,1033,1111]" pageId="31" pageNumber="148">
|
||
<emphasis id="B91CC2188317FFF5FF673F131700FBC3" bold="true" box="[155,272,1033,1052]" pageId="31" pageNumber="148">Figure 21.</emphasis>
|
||
Geographical distribution of
|
||
<taxonomicName id="4C6865898317FFF5FDAB3F131547FBC4" authorityName="Gunther" authorityYear="1903" box="[599,855,1033,1052]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="148" phylum="Chordata" rank="species" species="aureus">
|
||
<emphasis id="B91CC2188317FFF5FDAB3F131547FBC4" box="[599,855,1033,1052]" italics="true" pageId="31" pageNumber="148">Acanthodactylus aureus</emphasis>
|
||
</taxonomicName>
|
||
(circles) and
|
||
<taxonomicName id="4C6865898317FFF5FC0D3F13130CFBC3" authorityName="Geniez & Foucart" authorityYear="1995" box="[1009,1308,1033,1052]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="148" phylum="Chordata" rank="species" species="taghitensis">
|
||
<emphasis id="B91CC2188317FFF5FC0D3F13130CFBC3" box="[1009,1308,1033,1052]" italics="true" pageId="31" pageNumber="148">Acanthodactylus taghitensis</emphasis>
|
||
</taxonomicName>
|
||
(triangles). Data from
|
||
<bibRefCitation id="EFF963FB8317FFF5FEF03F3C178EFBE6" author="Bohme W" box="[268,414,1062,1081]" pageId="31" pageNumber="148" pagination="360 - 416" refId="ref21607" refString="Bohme W. 1978. Zur Herpetofaunistik des Senegal. Bonner Zoologische Beitrage 29: 360 - 416." type="journal article" year="1978">Böhme (1978)</bibRefCitation>
|
||
, Salvador (1982),
|
||
<bibRefCitation id="EFF963FB8317FFF5FDA43F3C1549FBE6" author="Geniez P & Foucart A" box="[600,857,1062,1082]" pageId="31" pageNumber="148" pagination="3 - 9" refId="ref22027" refString="Geniez P, Foucart A. 1995. Un nouvel Acanthodactyle en Algerie: Acanthodactylus taghitensis n. sp. (Reptilia, Sauria, Lacertidae). Bulletin du Museum National d'Histoire Naturelle, Section A, Zoologie, Biologie et Ecologie Animales 17: 3 - 9." type="journal article" year="1995">Geniez & Foucart (1995)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF963FB8317FFF5FC9F3F3D1257FBE6" author="Bons J & Geniez Ph" box="[867,1095,1062,1082]" pageId="31" pageNumber="148" refId="ref21662" refString="Bons J, Geniez Ph. 1996. Amphibiens et Reptiles du Maroc, Sahara Occidental compris. Atlas biogeographique. Barcelona: Asociacion Herpetologica Espanola." type="book" year="1996">Bons & Geniez (1996)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF963FB8317FFF5FBAD3F3C1313FBE6" author="Hasi M & Lopez-Jurado LF & Mateo JA & Geniez P" box="[1105,1283,1062,1082]" pageId="31" pageNumber="148" pagination="33 - 38" refId="ref22283" refString="Hasi M, Lopez-Jurado LF, Mateo JA, Geniez P. 1998. Nouvelles observations herpetologiques au Sahara Occidental, 3. Bulletin de la Societe Herpetologique de France 84: 33 - 38." type="journal article" year="1998">
|
||
Hasi
|
||
<emphasis id="B91CC2188317FFF5FB743F3D12A5FBE6" box="[1160,1205,1062,1081]" italics="true" pageId="31" pageNumber="148">et al</emphasis>
|
||
. (1998)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF963FB8317FFF5FAF23F3C16CEFB88" author="Donaire D & Mateo JA & Hasi M & Geniez Ph" pageId="31" pageNumber="148" pagination="8 - 12" refId="ref21946" refString="Donaire D, Mateo JA, Hasi M, Geniez Ph. 2000. Nuevos datos sobre la fauna reptiliana de la Hamada de Tinduf (Argelia). Boletin de la Asociation Herpetologica Espanola 11: 8 - 12." type="journal article" year="2000">
|
||
Donaire
|
||
<emphasis id="B91CC2188317FFF5FA943F3D1385FBE6" box="[1384,1429,1062,1081]" italics="true" pageId="31" pageNumber="148">et al</emphasis>
|
||
. (2000)
|
||
</bibRefCitation>
|
||
, this study.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD71E0A8317FFF5FF483D4214B1F9B1" blockId="31.[180,673,1624,1646]" box="[180,673,1624,1646]" pageId="31" pageNumber="148">
|
||
<emphasis id="B91CC2188317FFF5FF483D4217ACF9B1" box="[180,444,1624,1646]" italics="true" pageId="31" pageNumber="148">Geographical variation</emphasis>
|
||
. None documented.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |