412 lines
54 KiB
XML
412 lines
54 KiB
XML
<document id="440ED5AB10796662F84B0063BAAC6A60" ID-CLB-Dataset="272446" ID-DOI="10.1093/zoolinnean/zlad043" ID-GBIF-Dataset="81098796-8acf-4360-a625-ad76a281e99e" ID-ISSN="0024-4082" ID-Zenodo-Dep="8426263" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1696940821456" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan C. & Cerdeño, Esperanza" docDate="2023" docId="03B17549E92447689DAAFA02FB57FE65" docLanguage="en" docName="zlad043.pdf" docOrigin="Zoological Journal of the Linnean Society 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad043" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Protypotherium australe" docType="treatment" docVersion="9" lastPageNumber="429" masterDocId="FF880D31E92F47649E20FFB4FF9EFFC7" masterDocTitle="Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae" masterLastPageNumber="444" masterPageNumber="417" pageNumber="428" updateTime="1701999754155" updateUser="plazi">
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<mods:title id="EA680227B083853755CED16E8E9C9F65">Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae</mods:title>
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<mods:namePart id="CD656BBE9684FAA9E58DB528A367D29D">Fernández, Mercedes</mods:namePart>
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<mods:affiliation id="3CD7CDF9F6B5C20FFF0A9B08D72400FE">División Paleontología de Vertebrados, Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’, Avenida Ángel Gallardo 470, C 1405 DJR Ciudad Autónoma de Buenos Aires, Argentina & Laboratorio Anatomía y Evolución de los Vertebrados, Departamento de Ciencias Básicas, Universidad Nacional de Luján, Ruta 5 y Avenida Constitución 6700 Luján, Buenos Aires, Argentina & Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Juan Domingo Perón 2158, C 1040 AAH, Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="D94BC8CDCD0D086128FF3B3FF87CFE76">Fernicola, Juan C.</mods:namePart>
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<mods:affiliation id="66B4208EB3E9F45A4EBC96A23627F3C5">División Paleontología de Vertebrados, Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’, Avenida Ángel Gallardo 470, C 1405 DJR Ciudad Autónoma de Buenos Aires, Argentina & Laboratorio Anatomía y Evolución de los Vertebrados, Departamento de Ciencias Básicas, Universidad Nacional de Luján, Ruta 5 y Avenida Constitución 6700 Luján, Buenos Aires, Argentina & Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Juan Domingo Perón 2158, C 1040 AAH, Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="BAFE216D32C5E35D65CC2EB89F0184DE">Cerdeño, Esperanza</mods:namePart>
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<mods:affiliation id="39D9979C2AA6560DA866FB88BF01F748">Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Juan Domingo Perón 2158, C 1040 AAH, Buenos Aires, Argentina & Paleobiología y Paleoecología, Instituto de Nivología, Glaciología y Ciencias Ambientales, Centro Científico Tecnológico-CONICET-Mendoza, Avenida Ruiz Leal s / n, M 5500 Mendoza, Argentina</mods:affiliation>
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<mods:title id="B41D1379995E60A4CC95CF71E2E17019">Zoological Journal of the Linnean Society</mods:title>
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<mods:number id="DCCE89C6686592979C4CE404CEC8D49F">2023-07-11</mods:number>
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<treatment id="03B17549E92447689DAAFA02FB57FE65" ID-DOI="http://doi.org/10.5281/zenodo.10267832" ID-GBIF-Taxon="213996070" ID-Zenodo-Dep="10267832" LSID="urn:lsid:plazi:treatment:03B17549E92447689DAAFA02FB57FE65" httpUri="http://treatment.plazi.org/id/03B17549E92447689DAAFA02FB57FE65" lastPageId="12" lastPageNumber="429" pageId="11" pageNumber="428">
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<subSubSection id="C30297D4E924476F9DAAFA02FACDFA17" box="[906,1363,1462,1489]" pageId="11" pageNumber="428" type="nomenclature">
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<paragraph id="8BA7C45FE924476F9DAAFA02FACDFA17" blockId="11.[906,1363,1462,1489]" box="[906,1363,1462,1489]" pageId="11" pageNumber="428">
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<heading id="D0EF7333E924476F9DAAFA02FACDFA17" bold="true" box="[906,1363,1462,1489]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="428" reason="2">
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<taxonomicName id="4C18BFDCE924476F9DAAFA02FACDFA17" authority="(Ameghino, 1887a)" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[906,1363,1462,1489]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="australe">
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<emphasis id="B96C184DE924476F9DAAFA02FACDFA17" bold="true" box="[906,1363,1462,1489]" pageId="11" pageNumber="428">
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<emphasis id="B96C184DE924476F9DAAFA02FBE0FA16" bold="true" box="[906,1150,1462,1489]" italics="true" pageId="11" pageNumber="428">Protypotherium australe</emphasis>
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(
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<bibRefCitation id="EF89B9AEE924476F9AA9FA02FAD5FA17" author="Ameghino F" box="[1161,1355,1462,1488]" pageId="11" pageNumber="428" pagination="1 - 66" refId="ref21746" refString="Ameghino F. Observaciones generales sobre el orden de mamiferos extinguidos sub-americanos llamados Toxodontes (Toxodontia) y sinopsis de los generos y especies hasta ahora conocidos. Anales del Museo de La Plata (publicacion especial) 1887 a; 1: 1 - 66." type="journal article" year="1887">Ameghino, 1887a</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30297D4E924476F9D0AFA69FC49F917" pageId="11" pageNumber="428" type="description">
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<paragraph id="8BA7C45FE924476F9D0AFA69FB8AF9D3" blockId="11.[810,1459,1501,1963]" pageId="11" pageNumber="428">
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(
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<figureCitation id="1323D8DAE924476F9D15FA69FBBAFA33" box="[821,1060,1501,1525]" captionStart="Figure 2" captionStartId="5.[113,178,1779,1803]" captionTargetBox="[189,1384,147,1747]" captionTargetId="figure-7@5.[186,1387,144,1750]" captionTargetPageId="5" captionText="Figure 2. Protypotherium australe (D–E, G, I–J, L) and Pr. praerutilum (A–C, F, H, K, M), in occlusal view. (A) YPM-VPPU 15386, incomplete skull; (B) less I3 (alveolus)–C–dP1–P2–M3 of MACN-A 4049; (C) less I3–dP1–M3 of MACN-A 4278; (D) right I1 (alveolus)–dP1–P2–M3 of MACN-A 3882; (E) right I1–dP1–P2–M3 of MACN-A 9623–9633; (F) less I1 (alveolus)–dP1–P2–M3 of AMNH 9187; (G) right I1–2 (alveolus)–dP1–P2–M3 of YPM-VPPU 15643; (H) MLP 12-2886, incomplete skull; (I) right I1 (alveolus)–I2–dP1–P2–M3 of MACN-A 37; (J) MACN-A 3911, less maxillary fragment with C–dP1–P2–M3; (K) MACN-A 1778, right maxillary fragment with dP1–P2 (alveoli)– P3–M3; (L) right I1 (alveolus)–dP1–P2–M3 of AMNH 9149; (M) MACN-A 4016, less maxillary fragment with P4–M3.* indicates alveolus. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426270" httpUri="https://zenodo.org/record/8426270/files/figure.png" pageId="11" pageNumber="428">Figs 2D–E, G, I–J, L</figureCitation>
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,
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<figureCitation id="1323D8DAE924476F9A18FA69FB05FA33" box="[1080,1179,1501,1525]" captionStart="Figure 3" captionStartId="6.[129,194,1775,1799]" captionTargetBox="[200,1403,146,1745]" captionTargetId="figure-6@6.[198,1405,144,1747]" captionTargetPageId="6" captionText="Figure 3. Protypotherium australe (B, F–M) and Pr. praerutilum (A, C–E), in occlusal view. (A) MACN-A 3952, right mandibular fragment with p3–m3; (B) less dp1–p2–m3 of AMNH 9286; (C) MACN-A 11191a, right mandibular fragment with m1–3; (D) right p2–m3 of MACN-A 9614; (E) right i1–dp1–p2–m3 of MACN-A 1082; (F) less p4–m3 of AMNH 9565; (G) right p2–m3 of AMNH 9149; (H) less i1–dp1– p2–m3 of MPM-PV 3531; (I) MACN-A 532, right mandibular fragment with m2–3; (J) right p2–m3 of MPM-PV 3739; (K) MPM-PV 19612, right mandibular fragment with m2 (broken)–3; (L) MACN-A 3979, less mandibular fragment with p3–m3; (M) MACN-A 3955, less mandibular fragment with p3–m3. * indicates alveolus; ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426272" httpUri="https://zenodo.org/record/8426272/files/figure.png" pageId="11" pageNumber="428">3B, F–M</figureCitation>
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,
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<figureCitation id="1323D8DAE924476F9A8FFA69FB71FA33" box="[1199,1263,1501,1525]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="11" pageNumber="428">4C, F</figureCitation>
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,
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<figureCitation id="1323D8DAE924476F9B22FA69FAB4FA33" box="[1282,1322,1501,1525]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStart-2="Figure 7" captionStartId-0="8.[129,194,1753,1777]" captionStartId-1="9.[112,177,1757,1781]" captionStartId-2="10.[128,193,598,622]" captionTargetBox-0="[257,1345,147,1723]" captionTargetBox-1="[242,1330,147,1727]" captionTargetBox-2="[164,1435,147,567]" captionTargetId-0="figure-83@8.[254,1348,144,1726]" captionTargetId-1="figure-87@9.[239,1333,144,1730]" captionTargetId-2="figure-773@10.[161,1441,144,570]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." captionText-1="Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." captionText-2="Figure 7. Box-plot showing the length of I1–3 in: (A) Pr.praerutilum; (B) Pr.australe." figureDoi-0="http://doi.org/10.5281/zenodo.8426276" figureDoi-1="http://doi.org/10.5281/zenodo.8426278" figureDoi-2="http://doi.org/10.5281/zenodo.8426280" httpUri-0="https://zenodo.org/record/8426276/files/figure.png" httpUri-1="https://zenodo.org/record/8426278/files/figure.png" httpUri-2="https://zenodo.org/record/8426280/files/figure.png" pageId="11" pageNumber="428">5–7</figureCitation>
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; Supporting Information, Table S2)
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</paragraph>
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<paragraph id="8BA7C45FE924476F9D66F9A8FC1EF994" blockId="11.[810,1459,1501,1963]" pageId="11" pageNumber="428">
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1882
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<taxonomicName id="4C18BFDCE924476F9D5AF9AFFB66F9F4" authority="Moreno" authorityName="Moreno" box="[890,1272,1563,1587]" class="Mammalia" family="Interatheriidae" genus="Toxodontophanus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="australis">
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<emphasis id="B96C184DE924476F9D5AF9AFFB0EF9F4" box="[890,1168,1563,1587]" italics="true" pageId="11" pageNumber="428">Toxodontophanus australis</emphasis>
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Moreno
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</taxonomicName>
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, p. 23 (
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<emphasis id="B96C184DE924476F9B52F9A8FCF1F994" italics="true" pageId="11" pageNumber="428">nomen nudum</emphasis>
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).
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</paragraph>
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<paragraph id="8BA7C45FE924476F9D66F9EEFC30F956" blockId="11.[810,1459,1501,1963]" pageId="11" pageNumber="428">
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1887a
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<taxonomicName id="4C18BFDCE924476F9DA5F9EEFA97F9B5" authority="Ameghino" authorityName="Ameghino" box="[901,1289,1626,1650]" class="Mammalia" family="Interatheriidae" genus="Toxodontophanus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="australis">
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<emphasis id="B96C184DE924476F9DA5F9EEFB0FF9B5" box="[901,1169,1626,1650]" italics="true" pageId="11" pageNumber="428">Toxodontophanus australis</emphasis>
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Ameghino
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</taxonomicName>
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, p. 64 (original description).
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</paragraph>
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<paragraph id="8BA7C45FE924476F9D66F92DFC49F917" blockId="11.[810,1459,1501,1963]" pageId="11" pageNumber="428">
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1887b
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<taxonomicName id="4C18BFDCE924476F9DA7F92DFA31F976" authority="(Ameghino, 1887a)" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[903,1455,1689,1713]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="australe">
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<emphasis id="B96C184DE924476F9DA7F92DFB08F976" box="[903,1174,1689,1713]" italics="true" pageId="11" pageNumber="428">Protypotherium australe</emphasis>
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(
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<bibRefCitation id="EF89B9AEE924476F9AE6F92DFA38F976" author="Ameghino F" box="[1222,1446,1689,1713]" pageId="11" pageNumber="428" pagination="1 - 66" refId="ref21746" refString="Ameghino F. Observaciones generales sobre el orden de mamiferos extinguidos sub-americanos llamados Toxodontes (Toxodontia) y sinopsis de los generos y especies hasta ahora conocidos. Anales del Museo de La Plata (publicacion especial) 1887 a; 1: 1 - 66." type="journal article" year="1887">Ameghino, 1887a</bibRefCitation>
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)
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</taxonomicName>
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; Ameghino, p. 63.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30297D4E924476F9D66F963FCECF86C" pageId="11" pageNumber="428" type="reference_group">
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<paragraph id="8BA7C45FE924476F9D66F963FA3BF928" blockId="11.[810,1459,1501,1963]" box="[838,1445,1751,1775]" pageId="11" pageNumber="428">
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1909
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<taxonomicName id="4C18BFDCE924476F9D5AF963FA90F928" authority="Ameghino, 1891 b" authorityName="Ameghino" authorityYear="1891" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[890,1294,1751,1775]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="furculosus">
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<emphasis id="B96C184DE924476F9D5AF963FBD0F928" box="[890,1102,1751,1775]" italics="true" pageId="11" pageNumber="428">Patriarchus furculosus</emphasis>
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<bibRefCitation id="EF89B9AEE924476F9A74F963FA90F928" author="Ameghino F" box="[1108,1294,1751,1775]" pageId="11" pageNumber="428" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino, 1891b</bibRefCitation>
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</taxonomicName>
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; Sinclair, p. 33.
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</paragraph>
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<paragraph id="8BA7C45FE924476F9D66F943FA07F8C8" blockId="11.[810,1459,1501,1963]" box="[838,1433,1783,1807]" pageId="11" pageNumber="428">
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1909
|
||
<taxonomicName id="4C18BFDCE924476F9D5AF943FA9CF8C8" authority="Ameghino, 1891 b" authorityName="Ameghino" authorityYear="1891" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[890,1282,1783,1807]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="distortus">
|
||
<emphasis id="B96C184DE924476F9D5AF943FBDCF8C8" box="[890,1090,1783,1807]" italics="true" pageId="11" pageNumber="428">Patriarchus distortus</emphasis>
|
||
<bibRefCitation id="EF89B9AEE924476F9A68F943FA9CF8C8" author="Ameghino F" box="[1096,1282,1783,1807]" pageId="11" pageNumber="428" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino, 1891b</bibRefCitation>
|
||
</taxonomicName>
|
||
; Sinclair, p. 33.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE924476F9D66F8A2FAEAF8E9" blockId="11.[810,1459,1501,1963]" box="[838,1396,1814,1838]" pageId="11" pageNumber="428">
|
||
1909
|
||
<taxonomicName id="4C18BFDCE924476F9D5AF8A2FB42F8E9" authority="Ameghino, 1891 b" authorityName="Ameghino" authorityYear="1891" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[890,1244,1814,1838]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="altus">
|
||
<emphasis id="B96C184DE924476F9D5AF8A2FB83F8E9" box="[890,1053,1814,1838]" italics="true" pageId="11" pageNumber="428">Patriarchus altus</emphasis>
|
||
<bibRefCitation id="EF89B9AEE924476F9A02F8A2FB42F8E9" author="Ameghino F" box="[1058,1244,1814,1838]" pageId="11" pageNumber="428" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino, 1891b</bibRefCitation>
|
||
</taxonomicName>
|
||
; Sinclair, p. 34.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE924476F9D66F881FCECF8AA" blockId="11.[810,1459,1501,1963]" pageId="11" pageNumber="428">
|
||
<taxonomicName id="4C18BFDCE924476F9D66F881FB4AF88A" authority="Ameghino, 1904" authorityName="Ameghino" authorityYear="1904" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[838,1236,1845,1869]" class="Mammalia" family="Interatheriidae" genus="Epipatriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="bifidens">
|
||
<emphasis id="B96C184DE924476F9D66F881FBBAF88A" box="[838,1060,1845,1869]" italics="true" pageId="11" pageNumber="428">Epipatriarchus bifidens</emphasis>
|
||
<bibRefCitation id="EF89B9AEE924476F9A09F881FB4AF88A" author="Ameghino F" box="[1065,1236,1845,1869]" pageId="11" pageNumber="428" pagination="225 - 291" refId="ref22125" refString="Ameghino F. Nuevas especies de mamiferos cretaceos y terciarios de la Republica Argentina. Anales de la Sociedad Cientifica Argentina 1904; 56: 162 - 175, 327 - 341, 58: 35 - 42, 56 - 71, 182 - 192, 225 - 291." type="journal article" year="1904">Ameghino, 1904</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE924476F9D66F8C0FCECF86C" blockId="11.[810,1459,1501,1963]" pageId="11" pageNumber="428">
|
||
<taxonomicName id="4C18BFDCE924476F9D66F8C0FB23F84B" authority="Lane, 1927" authorityName="Lane" authorityYear="1927" box="[838,1213,1908,1932]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="martini">
|
||
<emphasis id="B96C184DE924476F9D66F8C0FBABF84B" box="[838,1077,1908,1932]" italics="true" pageId="11" pageNumber="428">Protypotherium martini</emphasis>
|
||
<bibRefCitation id="EF89B9AEE924476F9A61F8C0FB23F84B" author="Lane HH" box="[1089,1213,1908,1932]" pageId="11" pageNumber="428" pagination="313 - 7" refId="ref24387" refString="Lane HH. A new protypothere from the Santa Cruz Formation of Patagonia. University of Kansas. Science Bulletin 1927; 17: 313 - 7." type="journal article" year="1927">Lane, 1927</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30297D4E92347689EA1FF24FB57FE65" pageId="12" pageNumber="429" type="materials_examined">
|
||
<paragraph id="8BA7C45FE92347689EA1FF24FD90FEC1" blockId="12.[129,778,144,262]" pageId="12" pageNumber="429">
|
||
<materialsCitation id="3B70CE02E92347689EA1FF24FD90FEC1" ID-GBIF-Occurrence="4425895304" collectionCode="MACN-A" country="Argentina" location="Early - Middle Miocene" pageId="12" pageNumber="429" specimenCode="MACN-A 1338, MACN-A 1345-1346" specimenCount="2" stateProvince="Santa Cruz" typeStatus="holotype">
|
||
<emphasis id="B96C184DE92347689EA1FF24FF7FFF6F" box="[129,225,144,168]" italics="true" pageId="12" pageNumber="429">
|
||
<typeStatus id="54A37AFDE92347689EA1FF24FF42FF6F" box="[129,220,144,168]" pageId="12" pageNumber="429" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="DBBE6C24E92347689ECCFF24FE12FF6F" box="[236,396,144,168]" collectionCode="MACN-A" pageId="12" pageNumber="429">MACN-A 1338</specimenCode>
|
||
and
|
||
<specimenCode id="DBBE6C24E92347689F9DFF24FD00FF6F" box="[445,670,144,168]" collectionCode="MACN-A" pageId="12" pageNumber="429">MACN-A 1345–1346</specimenCode>
|
||
(
|
||
<figureCitation id="1323D8DAE92347689C8FFF24FD66FF6F" box="[687,760,144,168]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="12" pageNumber="429">Fig. 4C</figureCitation>
|
||
), casts of the missing holotype with less maxillary fragment with P3–M3 (see
|
||
<bibRefCitation id="EF89B9AEE92347689F2EFF7BFE68FF20" author="Fernandez M & Fernicola JC & Cerdeno E" box="[270,502,207,231]" pageId="12" pageNumber="429" pagination="473 - 98" refId="ref23394" refString="Fernandez M, Fernicola JC, Cerdeno E. Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino. Zootaxa 2018; 4387: 473 - 98. hups: // doi. org / 10.11646 / ZOOTAXA. 4387.3.4" type="journal article" year="2018">
|
||
Fernández
|
||
<emphasis id="B96C184DE92347689FA4FF7BFE27FF20" box="[388,441,207,231]" italics="true" pageId="12" pageNumber="429">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
),
|
||
<location id="8EC79284E92347689C32FF7BFC98FF20" LSID="urn:lsid:plazi:treatment:03B17549E92447689DAAFA02FB57FE65:8EC79284E92347689C32FF7BFC98FF20" box="[530,774,207,231]" country="Argentina" name="Early - Middle Miocene" pageId="12" pageNumber="429" stateProvince="Santa Cruz">Early–Middle Miocene</location>
|
||
, SCF,
|
||
<collectingRegion id="49DC0ABDE92347689E97FF5AFE16FEC1" box="[183,392,238,262]" country="Argentina" name="Santa Cruz" pageId="12" pageNumber="429">Santa Cruz Province</collectingRegion>
|
||
(
|
||
<collectingCountry id="F30F84CFE92347689FB4FF5AFD9CFEC1" box="[404,514,238,262]" name="Argentina" pageId="12" pageNumber="429">Argentina</collectingCountry>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689EA1FE93FD3CFEF8" blockId="12.[129,674,295,319]" box="[129,674,295,319]" pageId="12" pageNumber="429">
|
||
<emphasis id="B96C184DE92347689EA1FE93FEA7FEF8" box="[129,313,295,319]" italics="true" pageId="12" pageNumber="429">Referred materials:</emphasis>
|
||
Supporting Information,
|
||
<tableCitation id="C69AF1E4E92347689C64FE93FD02FEF8" box="[580,668,295,319]" captionStart="Table 1" captionStartId="21.[113,168,143,167]" captionTargetPageId="21" captionText="Table 1. Synthesis of the taxonomic scheme regarding the Santacrucian Interatheriinae genera Protypotherium, Interatherium (asser Fernández et al. 2023), Patriarchus (asser Fernández et al.2019), and Neoicochilus (asser Fernández et al.2021b). Species indicated in bold are the synonyms proposed in this contribution" httpUri="http://table.plazi.org/id/DF6794D7E93A47719E51FF3BFE73FF18" pageId="12" pageNumber="429" tableUuid="DF6794D7E93A47719E51FF3BFE73FF18">Table S1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689EA1FED5FD82FC89" blockId="12.[129,780,353,847]" pageId="12" pageNumber="429">
|
||
<emphasis id="B96C184DE92347689EA1FED5FDD1FEBE" box="[129,591,353,377]" italics="true" pageId="12" pageNumber="429">
|
||
Extended diagnosis:
|
||
<taxonomicName id="4C18BFDCE92347689F7AFED5FDD1FEBE" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[346,591,353,377]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="australe">Protypotherium australe</taxonomicName>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C18BFDCE92347689CCFFED5FF6BFE5F" authorityName="Ameghino" authorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE92347689CCFFED5FF6BFE5F" italics="true" pageId="12" pageNumber="429">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
by being larger (~20%) and I1 longer than I2–3; from
|
||
<taxonomicName id="4C18BFDCE92347689E9BFE14FEA3FE70" authorityName="Ameghino" authorityYear="1885" box="[187,317,416,439]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="antiquum">
|
||
<emphasis id="B96C184DE92347689E9BFE14FEA3FE70" box="[187,317,416,439]" italics="true" pageId="12" pageNumber="429">Pr. antiquum</emphasis>
|
||
</taxonomicName>
|
||
by the talonid of m3 with a shallower (and even absent) labial groove, and its mesial lobe with its distal region concave; from
|
||
<taxonomicName id="4C18BFDCE92347689F75FE6BFE62FE31" box="[341,508,478,502]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="colloncurensis">
|
||
<emphasis id="B96C184DE92347689F75FE6BFE62FE31" box="[341,508,478,502]" italics="true" pageId="12" pageNumber="429">Pr. colloncurensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE92347689C14FE6BFD20FE31" authorityName="Ameghino" authorityYear="1902" box="[564,702,478,502]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="columnifer">
|
||
<emphasis id="B96C184DE92347689C14FE6BFD20FE31" box="[564,702,478,502]" italics="true" pageId="12" pageNumber="429">Pr. columnifer</emphasis>
|
||
</taxonomicName>
|
||
by the talonid of p2–4 sub-triangular in outline; from
|
||
<taxonomicName id="4C18BFDCE92347689C41FE4AFC97FDD2" authorityName="Ameghino" authorityYear="1891" box="[609,777,509,533]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="compressidens">
|
||
<emphasis id="B96C184DE92347689C41FE4AFC97FDD2" box="[609,777,509,533]" italics="true" pageId="12" pageNumber="429">Pr. compressidens</emphasis>
|
||
</taxonomicName>
|
||
by exhibiting wider cheek teeth, in particular P2–M3; from
|
||
<taxonomicName id="4C18BFDCE92347689EA1FD89FEB9FD93" box="[129,295,572,596]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="colloncurensis">
|
||
<emphasis id="B96C184DE92347689EA1FD89FEB9FD93" box="[129,295,572,596]" italics="true" pageId="12" pageNumber="429">Pr. colloncurensis</emphasis>
|
||
</taxonomicName>
|
||
by a descending process of the maxilla with a moderate development; from
|
||
<taxonomicName id="4C18BFDCE92347689F97FDE8FDB3FDB4" authorityName="Ameghino" authorityYear="1889" box="[439,557,603,627]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="claudum">
|
||
<emphasis id="B96C184DE92347689F97FDE8FDB3FDB4" box="[439,557,603,627]" italics="true" pageId="12" pageNumber="429">Pr. claudum</emphasis>
|
||
</taxonomicName>
|
||
by having p2–3 with a distinctive entoflexid, p4 with a transversely deeper, narrower and more distally-placed ectoflexid, and m1–3 with deeper entoflexids; from
|
||
<taxonomicName id="4C18BFDCE92347689F19FD0EFE2BFD16" authorityName="Cabrera & Kraglievich" authorityYear="1931" box="[313,437,698,721]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="minutum">
|
||
<emphasis id="B96C184DE92347689F19FD0EFE2BFD16" box="[313,437,698,721]" italics="true" pageId="12" pageNumber="429">Pr. minutum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE92347689FCAFD0EFDECFD16" authorityName="Cabrera and Kraglievich" authorityYear="1931" box="[490,626,697,721]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="distinctum">
|
||
<emphasis id="B96C184DE92347689FCAFD0EFDECFD16" box="[490,626,697,721]" italics="true" pageId="12" pageNumber="429">Pr. distinctum</emphasis>
|
||
</taxonomicName>
|
||
by having less cement covering the cheek teeth, the P2–4 with an entoflexus located in a more distal position and slightly less expanded, and M1–3 with straighter ectolophs; and from
|
||
<taxonomicName id="4C18BFDCE92347689C1AFCACFD5CFCE8" authorityName="Ameghino" authorityYear="1902" box="[570,706,791,815]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="columnifer">
|
||
<emphasis id="B96C184DE92347689C1AFCACFD5CFCE8" box="[570,706,791,815]" italics="true" pageId="12" pageNumber="429">Pr. columnifer</emphasis>
|
||
</taxonomicName>
|
||
by the talonid of p4 narrower than the trigonid.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689EA1FCC4FD0BFBE2" blockId="12.[129,777,880,1061]" pageId="12" pageNumber="429">
|
||
<emphasis id="B96C184DE92347689EA1FCC4FD80FC4F" box="[129,542,880,904]" italics="true" pageId="12" pageNumber="429">Geographic and stratigraphic provenance:</emphasis>
|
||
<collectingRegion id="49DC0ABDE92347689C10FCC4FC97FC4F" box="[560,777,880,904]" country="Argentina" name="Santa Cruz" pageId="12" pageNumber="429">Santa Cruz Province</collectingRegion>
|
||
(
|
||
<collectingCountry id="F30F84CFE92347689EA8FC3BFF68FC60" box="[136,246,911,935]" name="Argentina" pageId="12" pageNumber="429">Argentina</collectingCountry>
|
||
), SCF,
|
||
<collectorName id="26EDA189E92347689F45FC3BFE02FC60" box="[357,412,911,935]" pageId="12" pageNumber="429">Early</collectorName>
|
||
–
|
||
<collectorName id="26EDA189E92347689F89FC3BFDF6FC60" box="[425,616,911,935]" pageId="12" pageNumber="429">Middle Miocene</collectorName>
|
||
,
|
||
<collectorName id="26EDA189E92347689CA5FC3BFC97FC60" box="[645,777,911,935]" pageId="12" pageNumber="429">Santacrucian</collectorName>
|
||
<collectionCode id="ED095C9AE92347689EA1FC1BFF45FC00" box="[129,219,943,967]" pageId="12" pageNumber="429">SALMA</collectionCode>
|
||
;
|
||
<location id="8EC79284E92347689ED4FC1BFEEEFC00" LSID="urn:lsid:plazi:treatment:03B17549E92447689DAAFA02FB57FE65:8EC79284E92347689ED4FC1BFEEEFC00" box="[244,368,943,967]" country="Argentina" name="Rio Negro" pageId="12" pageNumber="429" stateProvince="Santa Cruz">Río Negro</location>
|
||
,
|
||
<collectingRegion id="49DC0ABDE92347689FA9FC1AFE45FC01" box="[393,475,942,966]" country="Argentina" name="Chubut" pageId="12" pageNumber="429">Chubut</collectingRegion>
|
||
, and
|
||
<collectingRegion id="49DC0ABDE92347689C10FC1BFD0EFC00" box="[560,656,943,967]" country="Argentina" name="Neuquen" pageId="12" pageNumber="429">Neuquén</collectingRegion>
|
||
<location id="8EC79284E92347689C85FC1AFC97FC01" LSID="urn:lsid:plazi:treatment:03B17549E92447689DAAFA02FB57FE65:8EC79284E92347689C85FC1AFC97FC01" box="[677,777,942,966]" country="Argentina" name="Provinces" pageId="12" pageNumber="429" stateProvince="Neuquen">Provinces</location>
|
||
(
|
||
<collectingCountry id="F30F84CFE92347689EA8FC7AFF68FC21" box="[136,246,974,998]" name="Argentina" pageId="12" pageNumber="429">Argentina</collectingCountry>
|
||
),
|
||
<collectorName id="26EDA189E92347689F2AFC7AFE5EFC22" box="[266,448,974,998]" pageId="12" pageNumber="429">Collón Curá Fm.</collectorName>
|
||
,
|
||
<collectorName id="26EDA189E92347689FECFC7AFDE2FC21" box="[460,636,974,998]" pageId="12" pageNumber="429">Middle Miocene</collectorName>
|
||
,
|
||
<collectorName id="26EDA189E92347689CA8FC7AFC97FC21" box="[648,777,974,998]" pageId="12" pageNumber="429">Colloncuran</collectorName>
|
||
<collectionCode id="ED095C9AE92347689EA1FC59FF45FBC2" box="[129,219,1005,1029]" pageId="12" pageNumber="429">SALMA</collectionCode>
|
||
; south-western
|
||
<collectingRegion id="49DC0ABDE92347689F5DFC59FE53FBC2" box="[381,461,1005,1029]" country="Argentina" name="Chubut" pageId="12" pageNumber="429">Chubut</collectingRegion>
|
||
and north-western
|
||
<collectorName id="26EDA189E92347689CB7FC59FC97FBC2" box="[663,777,1005,1029]" pageId="12" pageNumber="429">Santa Cruz</collectorName>
|
||
provinces,
|
||
<collectorName id="26EDA189E92347689ECEFBB9FE53FBE3" box="[238,461,1036,1060]" pageId="12" pageNumber="429">Late Middle Miocene</collectorName>
|
||
, ‘
|
||
<collectorName id="26EDA189E92347689FF9FBB9FDACFBE2" box="[473,562,1037,1061]" pageId="12" pageNumber="429">Mayoan’</collectorName>
|
||
<collectionCode id="ED095C9AE92347689C18FBB9FD0CFBE2" box="[568,658,1037,1061]" pageId="12" pageNumber="429">SALMA</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689EA1FBF9FDE9FBA4" blockId="12.[129,631,1100,1124]" box="[129,631,1100,1124]" pageId="12" pageNumber="429">
|
||
<emphasis id="B96C184DE92347689EA1FBF9FE8DFBA3" box="[129,275,1101,1124]" italics="true" pageId="12" pageNumber="429">Measurements:</emphasis>
|
||
Supporting Information, Table S2.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689EA1FB31FDB4F994" blockId="12.[128,780,1157,1964]" pageId="12" pageNumber="429">
|
||
<emphasis id="B96C184DE92347689EA1FB31FF40FB5A" box="[129,222,1157,1181]" italics="true" pageId="12" pageNumber="429">Remarks:</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689ECEFB32FEE5FB5A" author="Lane HH" box="[238,379,1157,1181]" pageId="12" pageNumber="429" pagination="313 - 7" refId="ref24387" refString="Lane HH. A new protypothere from the Santa Cruz Formation of Patagonia. University of Kansas. Science Bulletin 1927; 17: 313 - 7." type="journal article" year="1927">Lane (1927)</bibRefCitation>
|
||
differentiated
|
||
<taxonomicName id="4C18BFDCE92347689C03FB32FD09FB5A" authorityName="Lane" authorityYear="1927" box="[547,663,1158,1181]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="martini">
|
||
<emphasis id="B96C184DE92347689C03FB32FD09FB5A" box="[547,663,1158,1181]" italics="true" pageId="12" pageNumber="429">Pr. martini</emphasis>
|
||
</taxonomicName>
|
||
from the other species of
|
||
<taxonomicName id="4C18BFDCE92347689F0CFB10FE58FB7B" authorityName="Ameghino" authorityYear="1885" box="[300,454,1188,1212]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96C184DE92347689F0CFB10FE58FB7B" box="[300,454,1188,1212]" italics="true" pageId="12" pageNumber="429">Protypotherium</emphasis>
|
||
</taxonomicName>
|
||
by the apparent narrowness of M1 (
|
||
<collectionCode id="ED095C9AE92347689E96FB70FF57FB1B" box="[182,201,1220,1244]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="429">R</collectionCode>
|
||
<subScript id="179CC61AE92347689EE9FB60FF79FB25" attach="left" box="[201,231,1236,1250]" fontSize="6" pageId="12" pageNumber="429">W/L</subScript>
|
||
= 0.56), the width of P2–4 (
|
||
<collectionCode id="ED095C9AE92347689C3FFB70FDACFB1B" box="[543,562,1220,1244]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="429">R</collectionCode>
|
||
<subScript id="179CC61AE92347689C12FB60FDCEFB25" attach="left" box="[562,592,1236,1250]" fontSize="6" pageId="12" pageNumber="429">W/L</subScript>
|
||
= 1.00, 1.25, and 1.11, respectively), and the presence of an acute disto-lingual margin. The author highlighted that the remaining dental characteristics, as well as the dental height, coincide with those of
|
||
<taxonomicName id="4C18BFDCE92347689EA1FAF6FF6EFA9E" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[129,240,1345,1369]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE92347689EA1FAF6FF6EFA9E" box="[129,240,1345,1369]" italics="true" pageId="12" pageNumber="429">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
. Nevertheless, not only was the sample studied by
|
||
<bibRefCitation id="EF89B9AEE92347689EA1FAD5FE94FABE" author="Lane HH" box="[129,266,1377,1401]" pageId="12" pageNumber="429" pagination="313 - 7" refId="ref24387" refString="Lane HH. A new protypothere from the Santa Cruz Formation of Patagonia. University of Kansas. Science Bulletin 1927; 17: 313 - 7." type="journal article" year="1927">Lane (1927)</bibRefCitation>
|
||
very small (
|
||
<specimenCount id="9D1E0FD6E92347689FB5FAD5FDA3FABE" box="[405,573,1377,1401]" pageId="12" pageNumber="429" type="generic">four specimens</specimenCount>
|
||
) in comparison to its abundance in the SCF, but also the values provided by the author fall within the range of variation obtained here for
|
||
<taxonomicName id="4C18BFDCE92347689CCFFA14FF53FA10" baseAuthorityName="Ameghino" baseAuthorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE92347689CCFFA14FF53FA10" italics="true" pageId="12" pageNumber="429">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
(M1, 0.55 ≤
|
||
<collectionCode id="ED095C9AE92347689F4BFA0BFEE5FA11" box="[363,379,1471,1494]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="429">R</collectionCode>
|
||
<subScript id="179CC61AE92347689F5BFA7BFE07FA1A" attach="left" box="[379,409,1487,1501]" fontSize="6" pageId="12" pageNumber="429">W/L</subScript>
|
||
≤ 0.8; P2, 0.6 <
|
||
<collectionCode id="ED095C9AE92347689C79FA0BFDF7FA11" box="[601,617,1471,1494]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="429">R</collectionCode>
|
||
<subScript id="179CC61AE92347689C49FA7BFD16FA1A" attach="left" box="[617,648,1487,1501]" fontSize="6" pageId="12" pageNumber="429">W/L</subScript>
|
||
≤ 1.37; P3, 0.72 ≤
|
||
<collectionCode id="ED095C9AE92347689EEAFA6AFF44FA32" box="[202,218,1502,1525]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="429">R</collectionCode>
|
||
<subScript id="179CC61AE92347689EFAFA5AFF66FA3B" attach="left" box="[218,248,1518,1532]" fontSize="6" pageId="12" pageNumber="429">W/L</subScript>
|
||
≤ 1.19; P4, 0.6 <
|
||
<collectionCode id="ED095C9AE92347689F96FA6AFE58FA32" box="[438,454,1502,1525]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="429">R</collectionCode>
|
||
<subScript id="179CC61AE92347689FE6FA5AFE7AFA3B" attach="left" box="[454,484,1518,1532]" fontSize="6" pageId="12" pageNumber="429">W/L</subScript>
|
||
<1.2). Regarding the acute disto-lingual margin, this is a variable feature not only for
|
||
<taxonomicName id="4C18BFDCE92347689CE2FA4AFF32F9F2" baseAuthorityName="Ameghino" baseAuthorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE92347689CE2FA4AFF32F9F2" italics="true" pageId="12" pageNumber="429">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
, but also for
|
||
<taxonomicName id="4C18BFDCE92347689F13F9A9FE56F9F2" authorityName="Ameghino" authorityYear="1887" box="[307,456,1565,1589]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE92347689F13F9A9FE56F9F2" box="[307,456,1565,1589]" italics="true" pageId="12" pageNumber="429">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
. Therefore,
|
||
<taxonomicName id="4C18BFDCE92347689C62F9A9FD32F9F3" authorityName="Lane" authorityYear="1927" box="[578,684,1565,1588]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="martini">
|
||
<emphasis id="B96C184DE92347689C62F9A9FD32F9F3" box="[578,684,1565,1588]" italics="true" pageId="12" pageNumber="429">Pr. martini</emphasis>
|
||
</taxonomicName>
|
||
is herein considered junior synonym of
|
||
<taxonomicName id="4C18BFDCE92347689F97F989FDBDF993" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[439,547,1596,1620]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE92347689F97F989FDBDF993" box="[439,547,1596,1620]" italics="true" pageId="12" pageNumber="429">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689EBCF9E8FB57FE65" blockId="12.[128,780,1157,1964]" lastBlockId="12.[825,1474,144,419]" pageId="12" pageNumber="429">
|
||
<bibRefCitation id="EF89B9AEE92347689EBCF9E8FED6F9B4" author="Vera B & Gonzalez-Ruiz L & Novo N" box="[156,328,1627,1651]" pageId="12" pageNumber="429" pagination="1143 - 63" refId="ref25496" refString="Vera B, Gonzalez-Ruiz L, Novo N, et al. The Interatheriinae (Mammalia, Notoungulata) of the Friasian sensu stricto and Mayoan (Middle to Late Miocene), and the fossils from Cerro Zeballos, Patagonia, Argentina. Journal of Systematic Palaeontology 2018; 17: 1143 - 63. hups: // doi. org / 10.1080 / 14772019.2018.1511387" type="journal article" year="2018">
|
||
Vera
|
||
<emphasis id="B96C184DE92347689EEDF9E8FF64F9B4" box="[205,250,1627,1651]" italics="true" pageId="12" pageNumber="429">et al.</emphasis>
|
||
(2018)
|
||
</bibRefCitation>
|
||
reviewed the genus
|
||
<taxonomicName id="4C18BFDCE92347689C2FF9EFFD01F9B4" authorityName="Ameghino" authorityYear="1904" box="[527,671,1627,1651]" class="Mammalia" family="Interatheriidae" genus="Epipatriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96C184DE92347689C2FF9EFFD01F9B4" box="[527,671,1627,1651]" italics="true" pageId="12" pageNumber="429">Epipatriarchus</emphasis>
|
||
</taxonomicName>
|
||
and stated that the
|
||
<typeStatus id="54A37AFDE92347689EF4F9CFFE9CF954" box="[212,258,1659,1683]" pageId="12" pageNumber="429">type</typeStatus>
|
||
materials of the
|
||
<typeStatus id="54A37AFDE92347689F89F9CFFE49F954" box="[425,471,1659,1683]" pageId="12" pageNumber="429">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C18BFDCE92347689C0FF9CFFD03F954" authorityName="Ameghino" authorityYear="1904" box="[559,669,1659,1683]" class="Mammalia" family="Interatheriidae" genus="Epipatriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="bifidens">
|
||
<emphasis id="B96C184DE92347689C0FF9CFFD03F954" box="[559,669,1659,1683]" italics="true" pageId="12" pageNumber="429">Ep. bifidens</emphasis>
|
||
</taxonomicName>
|
||
(currently missing, but figured by
|
||
<bibRefCitation id="EF89B9AEE92347689F51F92EFD97F975" author="Rovereto G" box="[369,521,1690,1714]" pageId="12" pageNumber="429" pagination="1 - 249" refId="ref24764" refString="Rovereto G. Los estratos araucanos y sus fosiles. Anales del Museo Nacional de Historia Natural de Buenos Aires 1914; 25: 1 - 249." type="journal article" year="1914">Rovereto 1914</bibRefCitation>
|
||
: figs 1–2), are assignable to
|
||
<taxonomicName id="4C18BFDCE92347689EBCF90DFE41F916" box="[156,479,1721,1745]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="colloncurensis">
|
||
<emphasis id="B96C184DE92347689EBCF90DFEA8F916" box="[156,310,1721,1745]" italics="true" pageId="12" pageNumber="429">Protypotherium</emphasis>
|
||
cf.
|
||
<emphasis id="B96C184DE92347689F7BF90DFE41F916" box="[347,479,1721,1745]" italics="true" pageId="12" pageNumber="429">colloncurensis</emphasis>
|
||
</taxonomicName>
|
||
. However, this identification was supported by the geographic and stratigraphic provenance of the
|
||
<typeStatus id="54A37AFDE92347689EEAF94CFF66F8D7" box="[202,248,1784,1808]" pageId="12" pageNumber="429">type</typeStatus>
|
||
materials of
|
||
<taxonomicName id="4C18BFDCE92347689FA7F94DFE64F8D7" authorityName="Ameghino" authorityYear="1904" box="[391,506,1784,1808]" class="Mammalia" family="Interatheriidae" genus="Epipatriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="bifidens">
|
||
<emphasis id="B96C184DE92347689FA7F94DFE64F8D7" box="[391,506,1784,1808]" italics="true" pageId="12" pageNumber="429">Ep. bifidens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE92347689C14F94DFD43F8D7" box="[564,733,1784,1808]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="colloncurensis">
|
||
<emphasis id="B96C184DE92347689C14F94DFD43F8D7" box="[564,733,1784,1808]" italics="true" pageId="12" pageNumber="429">Pr. colloncurensis</emphasis>
|
||
</taxonomicName>
|
||
, assumed to be the same by the authors. The provenance of these specimens is not entirely clear (and much less the stratigraphic provenance) as it happens with most materials housed at the old palaeontological collections, and it should not be used as a criterion for taxonomic identification. The illustrations in
|
||
<bibRefCitation id="EF89B9AEE92347689C8BF821FCE6FF6F" author="Rovereto G" box="[683,888,144,1964]" pageId="12" pageNumber="429" pagination="1 - 249" refId="ref24764" refString="Rovereto G. Los estratos araucanos y sus fosiles. Anales del Museo Nacional de Historia Natural de Buenos Aires 1914; 25: 1 - 249." type="journal article" year="1914">Rovereto (1914</bibRefCitation>
|
||
: figs 1–2) do allow us to agree with the generic assignment proposed by
|
||
<bibRefCitation id="EF89B9AEE92347689DD6FF04FB3AFF00" author="Vera B & Gonzalez-Ruiz L & Novo N" box="[1014,1188,175,199]" pageId="12" pageNumber="429" pagination="1143 - 63" refId="ref25496" refString="Vera B, Gonzalez-Ruiz L, Novo N, et al. The Interatheriinae (Mammalia, Notoungulata) of the Friasian sensu stricto and Mayoan (Middle to Late Miocene), and the fossils from Cerro Zeballos, Patagonia, Argentina. Journal of Systematic Palaeontology 2018; 17: 1143 - 63. hups: // doi. org / 10.1080 / 14772019.2018.1511387" type="journal article" year="2018">
|
||
Vera
|
||
<emphasis id="B96C184DE92347689A08FF04FBC8FF00" box="[1064,1110,175,199]" italics="true" pageId="12" pageNumber="429">et al.</emphasis>
|
||
(2018)
|
||
</bibRefCitation>
|
||
, but the dental morphology, particularly that of the lower teeth, and the dimensions (asser
|
||
<bibRefCitation id="EF89B9AEE92347689D19FF5AFC43FEC1" author="Ameghino F" box="[825,989,238,262]" pageId="12" pageNumber="429" pagination="225 - 291" refId="ref22125" refString="Ameghino F. Nuevas especies de mamiferos cretaceos y terciarios de la Republica Argentina. Anales de la Sociedad Cientifica Argentina 1904; 56: 162 - 175, 327 - 341, 58: 35 - 42, 56 - 71, 182 - 192, 225 - 291." type="journal article" year="1904">Ameghino 1904</bibRefCitation>
|
||
), align with those of
|
||
<taxonomicName id="4C18BFDCE92347689A90FF5BFA85FEC1" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[1200,1307,238,262]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE92347689A90FF5BFA85FEC1" box="[1200,1307,238,262]" italics="true" pageId="12" pageNumber="429">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
(see Supporting Information, Table S2). For instance,
|
||
<taxonomicName id="4C18BFDCE92347689A9BFEBAFAB5FEE2" authorityName="Ameghino" authorityYear="1904" box="[1211,1323,269,293]" class="Mammalia" family="Interatheriidae" genus="Epipatriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="bifidens">
|
||
<emphasis id="B96C184DE92347689A9BFEBAFAB5FEE2" box="[1211,1323,269,293]" italics="true" pageId="12" pageNumber="429">Ep. bifidens</emphasis>
|
||
</taxonomicName>
|
||
has lower premolars with sub-triangular talonids smaller than trigonids, as in
|
||
<taxonomicName id="4C18BFDCE92347689D19FEF9FC37FEA3" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[825,937,332,356]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE92347689D19FEF9FC37FEA3" box="[825,937,332,356]" italics="true" pageId="12" pageNumber="429">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
, whereas
|
||
<taxonomicName id="4C18BFDCE92347689A33FEF9FB25FEA3" box="[1043,1211,332,356]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="colloncurensis">
|
||
<emphasis id="B96C184DE92347689A33FEF9FB25FEA3" box="[1043,1211,332,356]" italics="true" pageId="12" pageNumber="429">Pr. colloncurensis</emphasis>
|
||
</taxonomicName>
|
||
has sub-circular talonids much smaller than trigonids. Therefore,
|
||
<taxonomicName id="4C18BFDCE92347689AEBFED8FAA7FE44" authorityName="Ameghino" authorityYear="1904" box="[1227,1337,363,387]" class="Mammalia" family="Interatheriidae" genus="Epipatriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="bifidens">
|
||
<emphasis id="B96C184DE92347689AEBFED8FAA7FE44" box="[1227,1337,363,387]" italics="true" pageId="12" pageNumber="429">Ep. bifidens</emphasis>
|
||
</taxonomicName>
|
||
is considered here as a junior synonym of
|
||
<taxonomicName id="4C18BFDCE92347689A77FE3FFB5DFE65" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[1111,1219,394,418]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE92347689A77FE3FFB5DFE65" box="[1111,1219,394,418]" italics="true" pageId="12" pageNumber="429">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |