802 lines
119 KiB
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802 lines
119 KiB
XML
<document id="43A67F9F33A0F846F4BF970DA7875E01" ID-CLB-Dataset="272446" ID-DOI="10.1093/zoolinnean/zlad043" ID-GBIF-Dataset="81098796-8acf-4360-a625-ad76a281e99e" ID-ISSN="0024-4082" ID-Zenodo-Dep="8426263" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1696940821456" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan C. & Cerdeño, Esperanza" docDate="2023" docId="03B17549E923476B9DAEFE69FE6CFA4A" docLanguage="en" docName="zlad043.pdf" docOrigin="Zoological Journal of the Linnean Society 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad043" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Protypotherium praerutilum Ameghino 1887" docType="treatment" docVersion="9" lastPageNumber="432" masterDocId="FF880D31E92F47649E20FFB4FF9EFFC7" masterDocTitle="Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae" masterLastPageNumber="444" masterPageNumber="417" pageNumber="429" updateTime="1701999754155" updateUser="plazi">
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<mods:title id="61AE94BD8CA89EDA54CE811CE4A0B754">Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae</mods:title>
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<mods:namePart id="B6A9B337A84007DEDD5EB956A65FBB06">Fernández, Mercedes</mods:namePart>
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<mods:affiliation id="9D73A10C5210A707752B97A3B83B295D">División Paleontología de Vertebrados, Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’, Avenida Ángel Gallardo 470, C 1405 DJR Ciudad Autónoma de Buenos Aires, Argentina & Laboratorio Anatomía y Evolución de los Vertebrados, Departamento de Ciencias Básicas, Universidad Nacional de Luján, Ruta 5 y Avenida Constitución 6700 Luján, Buenos Aires, Argentina & Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Juan Domingo Perón 2158, C 1040 AAH, Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="57B050A0FE58BE6E2033607146A667BA">Fernicola, Juan C.</mods:namePart>
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<mods:affiliation id="67BB7FE4021E6DEFC878163A82AE1B73">División Paleontología de Vertebrados, Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’, Avenida Ángel Gallardo 470, C 1405 DJR Ciudad Autónoma de Buenos Aires, Argentina & Laboratorio Anatomía y Evolución de los Vertebrados, Departamento de Ciencias Básicas, Universidad Nacional de Luján, Ruta 5 y Avenida Constitución 6700 Luján, Buenos Aires, Argentina & Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Juan Domingo Perón 2158, C 1040 AAH, Buenos Aires, Argentina</mods:affiliation>
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<mods:affiliation id="B2025D35C7D7A844BB769E31DC871983">Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Juan Domingo Perón 2158, C 1040 AAH, Buenos Aires, Argentina & Paleobiología y Paleoecología, Instituto de Nivología, Glaciología y Ciencias Ambientales, Centro Científico Tecnológico-CONICET-Mendoza, Avenida Ruiz Leal s / n, M 5500 Mendoza, Argentina</mods:affiliation>
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<mods:title id="A0D1E8734B507A9D92FD20DD8D946D8A">Zoological Journal of the Linnean Society</mods:title>
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<treatment id="03B17549E923476B9DAEFE69FE6CFA4A" ID-DOI="http://doi.org/10.5281/zenodo.10267834" ID-GBIF-Taxon="213996057" ID-Zenodo-Dep="10267834" LSID="urn:lsid:plazi:treatment:03B17549E923476B9DAEFE69FE6CFA4A" httpUri="http://treatment.plazi.org/id/03B17549E923476B9DAEFE69FE6CFA4A" lastPageId="15" lastPageNumber="432" pageId="12" pageNumber="429">
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<subSubSection id="C30297D4E92347689DAEFE69FAF3FE30" box="[910,1389,477,504]" pageId="12" pageNumber="429" type="nomenclature">
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<paragraph id="8BA7C45FE92347689DAEFE69FAF3FE30" blockId="12.[910,1389,477,504]" box="[910,1389,477,504]" pageId="12" pageNumber="429">
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<heading id="D0EF7333E92347689DAEFE69FAF3FE30" bold="true" box="[910,1389,477,504]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="429" reason="2">
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<taxonomicName id="4C18BFDCE92347689DAEFE69FAF3FE30" authority="Ameghino, 1887 b" authorityName="Ameghino" authorityYear="1887" box="[910,1389,477,504]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="praerutilum">
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<emphasis id="B96C184DE92347689DAEFE69FAF3FE30" bold="true" box="[910,1389,477,504]" pageId="12" pageNumber="429">
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<emphasis id="B96C184DE92347689DAEFE69FB34FE3F" bold="true" box="[910,1194,477,504]" italics="true" pageId="12" pageNumber="429">Protypotherium praerutilum</emphasis>
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<bibRefCitation id="EF89B9AEE92347689A8FFE69FAF3FE30" author="Ameghino F" box="[1199,1389,477,503]" pageId="12" pageNumber="429" pagination="1 - 26" refId="ref21793" refString="Ameghino F. Enumeracion sistematica de las especies de mamiferos fosiles coleccionados por Carlos Ameghino en los terrenos eocenos de Patagonia Austral y depositados en el Museo de La Plata. Boletin del Museo La Plata 1887 b; 1: 1 - 26." type="journal article" year="1887">Ameghino, 1887b</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C30297D4E92347689D19FDB0FAEEFDBD" pageId="12" pageNumber="429" type="description">
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<paragraph id="8BA7C45FE92347689D19FDB0FBBDFDFC" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
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(
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<figureCitation id="1323D8DAE92347689D64FDB0FBAEFDDC" box="[836,1072,516,540]" captionStart="Figure 2" captionStartId="5.[113,178,1779,1803]" captionTargetBox="[189,1384,147,1747]" captionTargetId="figure-7@5.[186,1387,144,1750]" captionTargetPageId="5" captionText="Figure 2. Protypotherium australe (D–E, G, I–J, L) and Pr. praerutilum (A–C, F, H, K, M), in occlusal view. (A) YPM-VPPU 15386, incomplete skull; (B) less I3 (alveolus)–C–dP1–P2–M3 of MACN-A 4049; (C) less I3–dP1–M3 of MACN-A 4278; (D) right I1 (alveolus)–dP1–P2–M3 of MACN-A 3882; (E) right I1–dP1–P2–M3 of MACN-A 9623–9633; (F) less I1 (alveolus)–dP1–P2–M3 of AMNH 9187; (G) right I1–2 (alveolus)–dP1–P2–M3 of YPM-VPPU 15643; (H) MLP 12-2886, incomplete skull; (I) right I1 (alveolus)–I2–dP1–P2–M3 of MACN-A 37; (J) MACN-A 3911, less maxillary fragment with C–dP1–P2–M3; (K) MACN-A 1778, right maxillary fragment with dP1–P2 (alveoli)– P3–M3; (L) right I1 (alveolus)–dP1–P2–M3 of AMNH 9149; (M) MACN-A 4016, less maxillary fragment with P4–M3.* indicates alveolus. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426270" httpUri="https://zenodo.org/record/8426270/files/figure.png" pageId="12" pageNumber="429">Figs 2A–C, F, H, K, M</figureCitation>
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,
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<figureCitation id="1323D8DAE92347689A1EFDB0FB06FDDB" box="[1086,1176,516,540]" captionStart="Figure 3" captionStartId="6.[129,194,1775,1799]" captionTargetBox="[200,1403,146,1745]" captionTargetId="figure-6@6.[198,1405,144,1747]" captionTargetPageId="6" captionText="Figure 3. Protypotherium australe (B, F–M) and Pr. praerutilum (A, C–E), in occlusal view. (A) MACN-A 3952, right mandibular fragment with p3–m3; (B) less dp1–p2–m3 of AMNH 9286; (C) MACN-A 11191a, right mandibular fragment with m1–3; (D) right p2–m3 of MACN-A 9614; (E) right i1–dp1–p2–m3 of MACN-A 1082; (F) less p4–m3 of AMNH 9565; (G) right p2–m3 of AMNH 9149; (H) less i1–dp1– p2–m3 of MPM-PV 3531; (I) MACN-A 532, right mandibular fragment with m2–3; (J) right p2–m3 of MPM-PV 3739; (K) MPM-PV 19612, right mandibular fragment with m2 (broken)–3; (L) MACN-A 3979, less mandibular fragment with p3–m3; (M) MACN-A 3955, less mandibular fragment with p3–m3. * indicates alveolus; ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426272" httpUri="https://zenodo.org/record/8426272/files/figure.png" pageId="12" pageNumber="429">3A, C–E</figureCitation>
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,
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<figureCitation id="1323D8DAE92347689A85FDB0FB41FDDB" box="[1189,1247,516,540]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="12" pageNumber="429">4D–F</figureCitation>
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,
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<figureCitation id="1323D8DAE92347689ACCFDB0FA8AFDDC" box="[1260,1300,516,540]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStart-2="Figure 7" captionStartId-0="8.[129,194,1753,1777]" captionStartId-1="9.[112,177,1757,1781]" captionStartId-2="10.[128,193,598,622]" captionTargetBox-0="[257,1345,147,1723]" captionTargetBox-1="[242,1330,147,1727]" captionTargetBox-2="[164,1435,147,567]" captionTargetId-0="figure-83@8.[254,1348,144,1726]" captionTargetId-1="figure-87@9.[239,1333,144,1730]" captionTargetId-2="figure-773@10.[161,1441,144,570]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." captionText-1="Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." captionText-2="Figure 7. Box-plot showing the length of I1–3 in: (A) Pr.praerutilum; (B) Pr.australe." figureDoi-0="http://doi.org/10.5281/zenodo.8426276" figureDoi-1="http://doi.org/10.5281/zenodo.8426278" figureDoi-2="http://doi.org/10.5281/zenodo.8426280" httpUri-0="https://zenodo.org/record/8426276/files/figure.png" httpUri-1="https://zenodo.org/record/8426278/files/figure.png" httpUri-2="https://zenodo.org/record/8426280/files/figure.png" pageId="12" pageNumber="429">5–7</figureCitation>
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,
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<figureCitation id="1323D8DAE92347689B01FDB0FAA0FDDB" box="[1313,1342,516,540]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)–p3–m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)–m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1–dp1–p2–p4, and right i1 and dp1–p2–m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3–M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)–m3; (G) MACN-A 3867, right mandibular fragment with dp1–2–p3–4 (trigonid); (H) MACN-A 9716–9732?g, right mandibular fragment with m1 (talonid)–3; (I) MPM-PV 19761, right maxillary fragment with M1–2; (J) MPM-PV 19781, right mandibular fragment with p2–m1; (K) MPM-PV 19782, less maxillary fragment with P2–M3. Abbreviation: ’, indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="12" pageNumber="429">8A</figureCitation>
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; Supporting Information, Table S2)
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</paragraph>
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<paragraph id="8BA7C45FE92347689D74FDF7FAEEFDBD" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
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1887b
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<taxonomicName id="4C18BFDCE92347689DB6FDF7FABFFD9C" authority="Ameghino" authorityName="Ameghino" authorityYear="1887" box="[918,1313,579,603]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="praerutilum">
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<emphasis id="B96C184DE92347689DB6FDF7FB32FD9C" box="[918,1196,579,603]" italics="true" pageId="12" pageNumber="429">Protypotherium praerutilum</emphasis>
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Ameghino
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</taxonomicName>
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, p. 15 (original description);
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<bibRefCitation id="EF89B9AEE92347689DE3FDD6FBF5FDBD" author="Ameghino F" box="[963,1131,610,634]" pageId="12" pageNumber="429" pagination="1 - 1028" refId="ref21871" refString="Ameghino F. Contribucion al conocimiento de los mamiferos fosiles de la Republica Argentina. Actas de la Academia Nacional de Ciencias Cordoba 1889; 6: 1 - 1028, atlas, 98 pls." type="journal article" year="1889">Ameghino 1889</bibRefCitation>
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, pl. 14, figs 6–7, 8, 12–13.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30297D4E92347689D74FD35FA34FACB" pageId="12" pageNumber="429" type="reference_group">
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<paragraph id="8BA7C45FE92347689D74FD35FCECFD7E" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
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1909
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<taxonomicName id="4C18BFDCE92347689DAAFD35FAC1FD5E" authority="Ameghino, 1891 b" authorityName="Ameghino" authorityYear="1891" box="[906,1375,641,665]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="leptocephalus">
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<emphasis id="B96C184DE92347689DAAFD35FB11FD5E" box="[906,1167,641,665]" italics="true" pageId="12" pageNumber="429">Patriarchus leptocephalus</emphasis>
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<bibRefCitation id="EF89B9AEE92347689ABAFD35FAC1FD5E" author="Ameghino F" box="[1178,1375,641,665]" pageId="12" pageNumber="429" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino, 1891b</bibRefCitation>
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</taxonomicName>
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; Sinclair, p. 40.
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</paragraph>
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<paragraph id="8BA7C45FE92347689D74FD74FCEEFD30" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
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1909
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<taxonomicName id="4C18BFDCE92347689DA8FD74FAFAFD1F" authority="Ameghino, 1891 b" authorityName="Ameghino" authorityYear="1891" box="[904,1380,704,728]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="convexidens">
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<emphasis id="B96C184DE92347689DA8FD74FB00FD1F" box="[904,1182,704,728]" italics="true" pageId="12" pageNumber="429">Protypotherium convexidens</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689A86FD74FAFAFD1F" author="Ameghino F" box="[1190,1380,704,728]" pageId="12" pageNumber="429" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino, 1891b</bibRefCitation>
|
||
</taxonomicName>
|
||
; Sinclair, p. 40.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689D74FD4BFC7BFCF1" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
|
||
<taxonomicName id="4C18BFDCE92347689D74FD4BFAA6FCD0" authority="Ameghino, 1887 b" authorityName="Ameghino" authorityYear="1887" box="[852,1336,767,791]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="attenuatum">
|
||
<emphasis id="B96C184DE92347689D74FD4BFBF5FCD1" box="[852,1131,767,791]" italics="true" pageId="12" pageNumber="429">Protypotherium attenuatum</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689A57FD4BFAA6FCD0" author="Ameghino F" box="[1143,1336,767,791]" pageId="12" pageNumber="429" pagination="1 - 26" refId="ref21793" refString="Ameghino F. Enumeracion sistematica de las especies de mamiferos fosiles coleccionados por Carlos Ameghino en los terrenos eocenos de Patagonia Austral y depositados en el Museo de La Plata. Boletin del Museo La Plata 1887 b; 1: 1 - 26." type="journal article" year="1887">Ameghino, 1887b</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689D74FC89FC1FFCB2" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
|
||
<taxonomicName id="4C18BFDCE92347689D74FC89FB4FFC92" authority="Ameghino, 1891 b" authorityName="Ameghino" authorityYear="1891" box="[852,1233,829,853]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="rectus">
|
||
<emphasis id="B96C184DE92347689D74FC89FB99FC92" box="[852,1031,829,853]" italics="true" pageId="12" pageNumber="429">Patriarchus rectus</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689A31FC89FB4FFC92" author="Ameghino F" box="[1041,1233,829,853]" pageId="12" pageNumber="429" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino, 1891b</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689D74FCC8FC7BFC74" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
|
||
<taxonomicName id="4C18BFDCE92347689D74FCC8FAB6FC53" authority="Ameghino, 1891 b" authorityName="Ameghino" authorityYear="1891" box="[852,1320,892,916]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="diastematus">
|
||
<emphasis id="B96C184DE92347689D74FCC8FBD2FC53" box="[852,1100,892,916]" italics="true" pageId="12" pageNumber="429">Patriarchus diastematus</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689A7FFCC8FAB6FC53" author="Ameghino F" box="[1119,1320,892,916]" pageId="12" pageNumber="429" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino, 1891b</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689D74FC0EFC1FFC35" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
|
||
<taxonomicName id="4C18BFDCE92347689D74FC0EFB7BFC14" authority="Ameghino, 1894" authorityName="Ameghino" authorityYear="1894" box="[852,1253,954,979]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="icochiloides">
|
||
<emphasis id="B96C184DE92347689D74FC0EFBABFC15" box="[852,1077,954,978]" italics="true" pageId="12" pageNumber="429">Patriarchus icochiloides</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689A19FC0EFB7BFC14" author="Ameghino F" box="[1081,1253,954,979]" pageId="12" pageNumber="429" refId="ref21976" refString="Ameghino F. Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Buenos Aires: Imprenta Coni e Hijos, 1894." type="book" year="1894">Ameghino, 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689D74FC4DFC7BFBF7" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
|
||
<taxonomicName id="4C18BFDCE92347689D74FC4DFAABFBD6" authority="Ameghino, 1891 b" authorityName="Ameghino" authorityYear="1891" box="[852,1333,1017,1041]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="diversidens">
|
||
<emphasis id="B96C184DE92347689D74FC4DFBFBFBD6" box="[852,1125,1017,1041]" italics="true" pageId="12" pageNumber="429">Protypotherium diversidens</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689A53FC4DFAABFBD6" author="Ameghino F" box="[1139,1333,1017,1041]" pageId="12" pageNumber="429" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino, 1891b</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689D74FB8CFC1FFBA8" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
|
||
<taxonomicName id="4C18BFDCE92347689D74FB8CFB7AFB97" authority="Ameghino, 1891 a" authorityName="Ameghino" authorityYear="1891" box="[852,1252,1080,1104]" class="Mammalia" family="Interatheriidae" genus="Eudiastatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="lingulatus">
|
||
<emphasis id="B96C184DE92347689D74FB8CFBB4FB97" box="[852,1066,1080,1104]" italics="true" pageId="12" pageNumber="429">Eudiastatus lingulatus</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689A0EFB8CFB7AFB97" author="Ameghino F" box="[1070,1252,1080,1104]" pageId="12" pageNumber="429" pagination="393 - 5" refId="ref21907" refString="Ameghino F. Los monos fosiles del Eoceno de la Republica Argentina. Revista Argentina de Historia Natural 1891 a; 1: 393 - 5." type="journal article" year="1891">Ameghino, 1891a</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689D74FBC2FC1FFB69" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
|
||
<taxonomicName id="4C18BFDCE92347689D74FBC2FB7AFB48" authority="Ameghino, 1894" authorityName="Ameghino" authorityYear="1894" box="[852,1252,1142,1167]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="lineare">
|
||
<emphasis id="B96C184DE92347689D74FBC2FBAAFB49" box="[852,1076,1142,1166]" italics="true" pageId="12" pageNumber="429">Protypotherium lineare</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689A18FBC2FB7AFB48" author="Ameghino F" box="[1080,1252,1142,1167]" pageId="12" pageNumber="429" refId="ref21976" refString="Ameghino F. Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Buenos Aires: Imprenta Coni e Hijos, 1894." type="book" year="1894">Ameghino, 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689D74FB01FC7BFB2B" blockId="12.[825,1474,516,1292]" pageId="12" pageNumber="429">
|
||
<taxonomicName id="4C18BFDCE92347689D74FB01FAB1FB0A" authority="Ameghino, 1891 b" authorityName="Ameghino" authorityYear="1891" box="[852,1327,1205,1229]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="globosum">
|
||
<emphasis id="B96C184DE92347689D74FB01FBC4FB0A" box="[852,1114,1205,1229]" italics="true" pageId="12" pageNumber="429">Protypotherium globosum</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689A4AFB01FAB1FB0A" author="Ameghino F" box="[1130,1327,1205,1229]" pageId="12" pageNumber="429" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino, 1891b</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689D74FB40FA34FACB" blockId="12.[825,1474,516,1292]" box="[852,1450,1268,1292]" pageId="12" pageNumber="429">
|
||
<taxonomicName id="4C18BFDCE92347689D74FB40FBE3FACB" authority="Roth, 1899" authorityName="Roth" authorityYear="1899" box="[852,1149,1268,1292]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="endiadys">
|
||
<emphasis id="B96C184DE92347689D74FB40FB9FFACB" box="[852,1025,1268,1292]" italics="true" pageId="12" pageNumber="429">Icochilus endiadys</emphasis>
|
||
<bibRefCitation id="EF89B9AEE92347689A27FB40FBE3FACB" author="Roth S" box="[1031,1149,1268,1292]" pageId="12" pageNumber="429" pagination="141 - 97" refId="ref24723" refString="Roth S. Apuntes sobre la geologia y paleontologia de los territorios de Rio Negro y Neuquen (diciembre de 1895 a junio de 1896). Revista del Museo de La Plata 1899; 9: 141 - 97." type="journal article" year="1899">Roth, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
; regarded as synonym herein.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30297D4E923476A9D19FA99FAC1F883" lastPageId="14" lastPageNumber="431" pageId="12" pageNumber="429" type="materials_examined">
|
||
<paragraph id="8BA7C45FE92347689D19FA99FAB5FA05" blockId="12.[825,1474,1325,1475]" pageId="12" pageNumber="429">
|
||
<materialsCitation id="3B70CE02E92347689D19FA99FAB5FA05" ID-GBIF-Occurrence="4425895303" collectionCode="MACN-A" country="Argentina" location="Early - Middle Miocene" pageId="12" pageNumber="429" specimenCode="MACN-A 1081, MACN-A 1082" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
|
||
<emphasis id="B96C184DE92347689D19FA99FC07FA82" box="[825,921,1325,1349]" italics="true" pageId="12" pageNumber="429">
|
||
<typeStatus id="54A37AFDE92347689D19FA99FC0AFA82" box="[825,916,1325,1349]" pageId="12" pageNumber="429" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="DBBE6C24E92347689D87FA99FBD0FA82" box="[935,1102,1325,1349]" collectionCode="MACN-A" pageId="12" pageNumber="429">MACN-A 1081</specimenCode>
|
||
(
|
||
<figureCitation id="1323D8DAE92347689A46FA99FB27FA82" box="[1126,1209,1325,1349]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="12" pageNumber="429">Fig. 4D</figureCitation>
|
||
), incomplete skull with less I1–M3 and right I1–2 (alveoli)–I3–C–dP1(alveoli)–P2– M3, and
|
||
<specimenCode id="DBBE6C24E92347689DB9FAD8FBA2FA43" box="[921,1084,1388,1412]" collectionCode="MACN-A" pageId="12" pageNumber="429">MACN-A 1082</specimenCode>
|
||
(
|
||
<figureCitation id="1323D8DAE92347689A6EFAD8FB09FA43" box="[1102,1175,1388,1412]" captionStart="Figure 3" captionStartId="6.[129,194,1775,1799]" captionTargetBox="[200,1403,146,1745]" captionTargetId="figure-6@6.[198,1405,144,1747]" captionTargetPageId="6" captionText="Figure 3. Protypotherium australe (B, F–M) and Pr. praerutilum (A, C–E), in occlusal view. (A) MACN-A 3952, right mandibular fragment with p3–m3; (B) less dp1–p2–m3 of AMNH 9286; (C) MACN-A 11191a, right mandibular fragment with m1–3; (D) right p2–m3 of MACN-A 9614; (E) right i1–dp1–p2–m3 of MACN-A 1082; (F) less p4–m3 of AMNH 9565; (G) right p2–m3 of AMNH 9149; (H) less i1–dp1– p2–m3 of MPM-PV 3531; (I) MACN-A 532, right mandibular fragment with m2–3; (J) right p2–m3 of MPM-PV 3739; (K) MPM-PV 19612, right mandibular fragment with m2 (broken)–3; (L) MACN-A 3979, less mandibular fragment with p3–m3; (M) MACN-A 3955, less mandibular fragment with p3–m3. * indicates alveolus; ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426272" httpUri="https://zenodo.org/record/8426272/files/figure.png" pageId="12" pageNumber="429">Fig. 3E</figureCitation>
|
||
), incomplete mandible with complete dentition (see
|
||
<bibRefCitation id="EF89B9AEE92347689A1AFA3FFA82FA64" author="Fernandez M & Fernicola JC & Cerdeno E" box="[1082,1308,1419,1443]" pageId="12" pageNumber="429" pagination="473 - 98" refId="ref23394" refString="Fernandez M, Fernicola JC, Cerdeno E. Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino. Zootaxa 2018; 4387: 473 - 98. hups: // doi. org / 10.11646 / ZOOTAXA. 4387.3.4" type="journal article" year="2018">
|
||
Fernández
|
||
<emphasis id="B96C184DE92347689A8EFA38FB7EFA64" box="[1198,1248,1419,1443]" italics="true" pageId="12" pageNumber="429">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
),
|
||
<location id="8EC79284E92347689B15FA3FFC0BFA05" LSID="urn:lsid:plazi:treatment:03B17549E923476B9DAEFE69FE6CFA4A:8EC79284E92347689B15FA3FFC0BFA05" country="Argentina" name="Early - Middle Miocene" pageId="12" pageNumber="429" stateProvince="Santa Cruz">Early–Middle Miocene</location>
|
||
, SCF,
|
||
<collectingRegion id="49DC0ABDE92347689DF4FA1FFB3BFA05" box="[980,1189,1450,1475]" country="Argentina" name="Santa Cruz" pageId="12" pageNumber="429">Santa Cruz Province</collectingRegion>
|
||
(
|
||
<collectingCountry id="F30F84CFE92347689A92FA1EFABEFA05" box="[1202,1312,1450,1474]" name="Argentina" pageId="12" pageNumber="429">Argentina</collectingCountry>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE92347689D19FA50FACBFA3C" blockId="12.[825,1365,1508,1532]" box="[825,1365,1508,1532]" pageId="12" pageNumber="429">
|
||
<emphasis id="B96C184DE92347689D19FA50FC6FFA3B" box="[825,1009,1508,1532]" italics="true" pageId="12" pageNumber="429">Referred materials:</emphasis>
|
||
Supporting Information,
|
||
<tableCitation id="C69AF1E4E92347689AD7FA50FAD1FA3B" box="[1271,1359,1508,1532]" captionStart="Table 1" captionStartId="21.[113,168,143,167]" captionTargetPageId="21" captionText="Table 1. Synthesis of the taxonomic scheme regarding the Santacrucian Interatheriinae genera Protypotherium, Interatherium (asser Fernández et al. 2023), Patriarchus (asser Fernández et al.2019), and Neoicochilus (asser Fernández et al.2021b). Species indicated in bold are the synonyms proposed in this contribution" httpUri="http://table.plazi.org/id/DF6794D7E93A47719E51FF3BFE73FF18" pageId="12" pageNumber="429" tableUuid="DF6794D7E93A47719E51FF3BFE73FF18">Table S1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE923476A9D19F9A9FD52FF20" blockId="12.[825,1477,1565,1965]" lastBlockId="14.[129,780,144,231]" lastPageId="14" lastPageNumber="431" pageId="12" pageNumber="429">
|
||
<emphasis id="B96C184DE92347689D19F9A9FABEF9F2" box="[825,1312,1565,1589]" italics="true" pageId="12" pageNumber="429">
|
||
Extended diagnosis:
|
||
<taxonomicName id="4C18BFDCE92347689A2BF9A9FABEF9F2" authorityName="Ameghino" authorityYear="1887" box="[1035,1312,1565,1589]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="praerutilum">Protypotherium praerutilum</taxonomicName>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C18BFDCE92347689B87F9AAFC1BF993" baseAuthorityName="Ameghino" baseAuthorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE92347689B87F9AAFC1BF993" italics="true" pageId="12" pageNumber="429">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
by exhibiting a smaller size (~20%) and a proportionally shorter I1 compared to I2–3; from
|
||
<taxonomicName id="4C18BFDCE92347689A89F9E8FAB4F9B4" authorityName="Ameghino" authorityYear="1885" box="[1193,1322,1628,1651]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="antiquum">
|
||
<emphasis id="B96C184DE92347689A89F9E8FAB4F9B4" box="[1193,1322,1628,1651]" italics="true" pageId="12" pageNumber="429">Pr. antiquum</emphasis>
|
||
</taxonomicName>
|
||
by the talonid of m3 with a shallower (and even absent) labial groove, and its mesial lobe with a concave distal region; from
|
||
<taxonomicName id="4C18BFDCE92347689B3CF92FFA5CF975" box="[1308,1474,1690,1714]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="colloncurensis">
|
||
<emphasis id="B96C184DE92347689B3CF92FFA5CF975" box="[1308,1474,1690,1714]" italics="true" pageId="12" pageNumber="429">Pr. colloncurensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE92347689D46F90EFC70F915" authorityName="Ameghino" authorityYear="1902" box="[870,1006,1722,1746]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="columnifer">
|
||
<emphasis id="B96C184DE92347689D46F90EFC70F915" box="[870,1006,1722,1746]" italics="true" pageId="12" pageNumber="429">Pr. columnifer</emphasis>
|
||
</taxonomicName>
|
||
by the talonid of p2–4 sub-triangular in outline; from
|
||
<taxonomicName id="4C18BFDCE92347689D85F96EFBD4F936" box="[933,1098,1753,1777]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="colloncurensis">
|
||
<emphasis id="B96C184DE92347689D85F96EFBD4F936" box="[933,1098,1753,1777]" italics="true" pageId="12" pageNumber="429">Pr. colloncurensis</emphasis>
|
||
</taxonomicName>
|
||
by a descending process of the maxilla with a moderate development; from
|
||
<taxonomicName id="4C18BFDCE92347689AC0F94DFAC8F8D7" authorityName="Ameghino" authorityYear="1889" box="[1248,1366,1784,1808]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="claudum">
|
||
<emphasis id="B96C184DE92347689AC0F94DFAC8F8D7" box="[1248,1366,1784,1808]" italics="true" pageId="12" pageNumber="429">Pr. claudum</emphasis>
|
||
</taxonomicName>
|
||
by having p2–3 with a differentiated entoflexid, p4 with a transversely deeper, narrower and more distally placed ectoflexid, and m1–3 with deeper entoflexids; from
|
||
<taxonomicName id="4C18BFDCE92347689AA0F8E3FAB3F8A9" authorityName="Ameghino" authorityYear="1891" box="[1152,1325,1878,1902]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="compressidens">
|
||
<emphasis id="B96C184DE92347689AA0F8E3FAB3F8A9" box="[1152,1325,1878,1902]" italics="true" pageId="12" pageNumber="429">Pr. compressidens</emphasis>
|
||
</taxonomicName>
|
||
by exhibiting wider cheek teeth, particularly P2–M3; from
|
||
<taxonomicName id="4C18BFDCE92347689B37F8C2FA0AF84A" authorityName="Cabrera & Kraglievich" authorityYear="1931" box="[1303,1428,1910,1933]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="minutum">
|
||
<emphasis id="B96C184DE92347689B37F8C2FA0AF84A" box="[1303,1428,1910,1933]" italics="true" pageId="12" pageNumber="429">Pr. minutum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE92347689D19F822FC21F86A" authorityName="Cabrera and Kraglievich" authorityYear="1931" box="[825,959,1941,1965]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="429" phylum="Chordata" rank="species" species="distinctum">
|
||
<emphasis id="B96C184DE92347689D19F822FC21F86A" box="[825,959,1941,1965]" italics="true" pageId="12" pageNumber="429">Pr. distinctum</emphasis>
|
||
</taxonomicName>
|
||
by less cement covering each tooth, the P2–4 with an entoflexus located in a more distal position and slightly less expanded, and M1–3 with straighter ectolophs; and from
|
||
<taxonomicName id="4C18BFDCE921476A9CCFFF04FF79FF20" authorityName="Ameghino" authorityYear="1902" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="columnifer">
|
||
<emphasis id="B96C184DE921476A9CCFFF04FF79FF20" italics="true" pageId="14" pageNumber="431">Pr. columnifer</emphasis>
|
||
</taxonomicName>
|
||
by the talonid of p4 narrower than the trigonid.
|
||
</paragraph>
|
||
<caption id="DF6794D7E92247699E51F940FB3CF873" ID-DOI="http://doi.org/10.5281/zenodo.8426282" ID-Zenodo-Dep="8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="13" pageNumber="430" startId="13.[113,178,1780,1804]" targetBox="[188,1387,151,1749]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8BA7C45FE92247699E51F940FB3CF873" blockId="13.[113,1450,1780,1973]" pageId="13" pageNumber="430">
|
||
<emphasis id="B96C184DE92247699E51F940FF57F8CB" bold="true" box="[113,201,1780,1804]" pageId="13" pageNumber="430">Figure 8.</emphasis>
|
||
<taxonomicName id="4C18BFDCE92247699EEEF940FE52F8CB" authorityName="Ameghino" authorityYear="1887" box="[206,460,1780,1804]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="13" pageNumber="430" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE92247699EEEF940FE52F8CB" box="[206,460,1780,1804]" italics="true" pageId="13" pageNumber="430">Protypotherium praerutilum</emphasis>
|
||
</taxonomicName>
|
||
, in occlusal view. A, MACN-A 4044, incomplete palate.
|
||
<taxonomicName id="4C18BFDCE92247699DFDF940FB20F8CB" authorityName="Ameghino" authorityYear="1889" box="[989,1214,1780,1804]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="13" pageNumber="430" phylum="Chordata" rank="species" species="claudum">
|
||
<emphasis id="B96C184DE92247699DFDF940FB20F8CB" box="[989,1214,1780,1804]" italics="true" pageId="13" pageNumber="430">Protypotherium claudum</emphasis>
|
||
</taxonomicName>
|
||
, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)–p3–m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)–m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1–dp1–p2–p4, and right i1 and dp1–p2–m3.
|
||
<taxonomicName id="4C18BFDCE92247699A5CF898FA13F883" authorityName="Ameghino" authorityYear="1891" box="[1148,1421,1836,1860]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="13" pageNumber="430" phylum="Chordata" rank="species" species="compressidens">
|
||
<emphasis id="B96C184DE92247699A5CF898FA13F883" box="[1148,1421,1836,1860]" italics="true" pageId="13" pageNumber="430">Protypotherium compressidens</emphasis>
|
||
</taxonomicName>
|
||
, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3–M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)–m3; (G) MACN-A 3867, right mandibular fragment with dp1–2–p3–4 (trigonid); (H) MACN-A 9716–9732?g, right mandibular fragment with m1 (talonid)–3; (I) MPM-PV 19761, right maxillary fragment with M1–2; (J) MPM-PV 19781, right mandibular fragment with p2–m1; (K) MPM-PV 19782, less maxillary fragment with P2–M3. Abbreviation: ’, indicates broken tooth. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA7C45FE921476A9EA1FEBCFD4EFEB9" blockId="14.[129,777,264,382]" pageId="14" pageNumber="431">
|
||
<emphasis id="B96C184DE921476A9EA1FEBCFD80FEE7" box="[129,542,264,288]" italics="true" pageId="14" pageNumber="431">Geographic and stratigraphic provenance:</emphasis>
|
||
<collectingRegion id="49DC0ABDE921476A9C10FEBCFC97FEE7" box="[560,777,264,288]" country="Argentina" name="Santa Cruz" pageId="14" pageNumber="431">Santa Cruz Province</collectingRegion>
|
||
(
|
||
<collectingCountry id="F30F84CFE921476A9EA8FE93FF68FEF8" box="[136,246,295,319]" name="Argentina" pageId="14" pageNumber="431">Argentina</collectingCountry>
|
||
), SCF,
|
||
<collectorName id="26EDA189E921476A9F45FE93FE02FEF8" box="[357,412,295,319]" pageId="14" pageNumber="431">Early</collectorName>
|
||
–
|
||
<collectorName id="26EDA189E921476A9F89FE93FDF6FEF8" box="[425,616,295,319]" pageId="14" pageNumber="431">Middle Miocene</collectorName>
|
||
,
|
||
<collectorName id="26EDA189E921476A9CA5FE93FC97FEF8" box="[645,777,295,319]" pageId="14" pageNumber="431">Santacrucian</collectorName>
|
||
<collectionCode id="ED095C9AE921476A9EA1FEF3FF45FE98" box="[129,219,327,351]" pageId="14" pageNumber="431">SALMA</collectionCode>
|
||
;
|
||
<location id="8EC79284E921476A9ECEFEF3FEFDFE98" LSID="urn:lsid:plazi:treatment:03B17549E923476B9DAEFE69FE6CFA4A:8EC79284E921476A9ECEFEF3FEFDFE98" box="[238,355,327,351]" country="Argentina" name="Rio Negro" pageId="14" pageNumber="431" stateProvince="Santa Cruz Province">Río Negro</location>
|
||
and
|
||
<collectingRegion id="49DC0ABDE921476A9F88FEF3FD96FE98" box="[424,520,327,351]" country="Argentina" name="Neuquen" pageId="14" pageNumber="431">Neuquén</collectingRegion>
|
||
provinces (
|
||
<collectingCountry id="F30F84CFE921476A9CB0FEF3FD60FE98" box="[656,766,327,351]" name="Argentina" pageId="14" pageNumber="431">Argentina</collectingCountry>
|
||
),
|
||
<collectorName id="26EDA189E921476A9EA1FED2FEB1FEBA" box="[129,303,358,382]" pageId="14" pageNumber="431">Collón Curá Fm.</collectorName>
|
||
,
|
||
<collectorName id="26EDA189E921476A9F18FED2FE7AFEB9" box="[312,484,358,382]" pageId="14" pageNumber="431">Middle Miocene</collectorName>
|
||
,
|
||
<collectorName id="26EDA189E921476A9FCCFED2FDF3FEB9" box="[492,621,358,382]" pageId="14" pageNumber="431">Colloncuran</collectorName>
|
||
<collectionCode id="ED095C9AE921476A9C53FED2FD53FEB9" box="[627,717,358,382]" pageId="14" pageNumber="431">SALMA</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE921476A9EA1FE1AFDE9FE02" blockId="14.[129,631,429,453]" box="[129,631,429,453]" pageId="14" pageNumber="431">
|
||
<emphasis id="B96C184DE921476A9EA1FE1AFE8DFE02" box="[129,275,430,453]" italics="true" pageId="14" pageNumber="431">Measurements:</emphasis>
|
||
Supporting Information, Table S2.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE921476A9EA1FE53FEC8FB95" blockId="14.[128,780,487,1983]" pageId="14" pageNumber="431">
|
||
<emphasis id="B96C184DE921476A9EA1FE53FF40FE38" box="[129,222,487,511]" italics="true" pageId="14" pageNumber="431">Remarks:</emphasis>
|
||
Regarding the upper permanent dentition,
|
||
<taxonomicName id="4C18BFDCE921476A9CCFFE53FF6BFDD9" authorityName="Ameghino" authorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9CCFFE53FF6BFDD9" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
exhibits I1 (x L =
|
||
<quantity id="4CE069BAE921476A9FFEFDB2FDDFFDDA" box="[478,577,518,542]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="14" pageNumber="431" unit="mm" value="4.4">4.4 mm</quantity>
|
||
; see Supporting Information, Table S2) with a greater mesio-distal length than that of I2–3 (I2, x
|
||
<subScript id="179CC61AE921476A9F1AFDE1FEDDFDA4" attach="left" box="[314,323,597,611]" fontSize="6" pageId="14" pageNumber="431">L</subScript>
|
||
= 4.0 mm; I3, x
|
||
<subScript id="179CC61AE921476A9FCBFDE1FE6AFDA4" attach="left" box="[491,500,597,611]" fontSize="6" pageId="14" pageNumber="431">L</subScript>
|
||
=
|
||
<quantity id="4CE069BAE921476A9C30FDF1FDFDFD9B" box="[528,611,581,605]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="431" unit="mm" value="3.7">3.7 mm</quantity>
|
||
; see Supporting Information, Table S2), but the differences between these lengths are less than those in
|
||
<taxonomicName id="4C18BFDCE921476A9F97FD30FDB8FD5C" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[439,550,643,667]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE921476A9F97FD30FDB8FD5C" box="[439,550,643,667]" italics="true" pageId="14" pageNumber="431">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
(I1, x L =
|
||
<quantity id="4CE069BAE921476A9CAEFD37FD7CFD5C" box="[654,738,643,667]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="14" pageNumber="431" unit="mm" value="5.8">5.8 mm</quantity>
|
||
; I2, x
|
||
<subScript id="179CC61AE921476A9EABFD07FF0AFD06" attach="left" box="[139,148,691,705]" fontSize="6" pageId="14" pageNumber="431">L</subScript>
|
||
≈
|
||
<quantity id="4CE069BAE921476A9E93FD17FE94FD7D" box="[179,266,675,699]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="14" pageNumber="431" unit="mm" value="4.7">4.7 mm</quantity>
|
||
; I3, x
|
||
<subScript id="179CC61AE921476A9F6DFD07FEC8FD06" attach="left" box="[333,342,691,705]" fontSize="6" pageId="14" pageNumber="431">L</subScript>
|
||
≈
|
||
<quantity id="4CE069BAE921476A9F54FD17FE55FD7D" box="[372,459,675,699]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="431" unit="mm" value="4.8">4.8 mm</quantity>
|
||
; see Supporting Information, Table S2) (see
|
||
<figureCitation id="1323D8DAE921476A9F3FFD76FEC4FD1E" box="[287,346,706,730]" captionStart="Figure 7" captionStartId="10.[128,193,598,622]" captionTargetBox="[164,1435,147,567]" captionTargetId="figure-773@10.[161,1441,144,570]" captionTargetPageId="10" captionText="Figure 7. Box-plot showing the length of I1–3 in: (A) Pr.praerutilum; (B) Pr.australe." figureDoi="http://doi.org/10.5281/zenodo.8426280" httpUri="https://zenodo.org/record/8426280/files/figure.png" pageId="14" pageNumber="431">Fig. 7</figureCitation>
|
||
).
|
||
<bibRefCitation id="EF89B9AEE921476A9F52FD76FD92FD1D" author="Tauber AA" box="[370,524,706,730]" pageId="14" pageNumber="431" pagination="1 - 28" refId="ref25150" refString="Tauber AA. Los representantes del genero Protypotherium (Mammalia, Notoungulata, Interatheriidae) del Mioceno Temprano del Sudeste de la Provincia de Santa Cruz, Republica Argentina. Academia Nacional de Ciencias, Cordoba, Misc 1996; 95: 1 - 28." type="journal article" year="1996">Tauber (1996)</bibRefCitation>
|
||
mentioned the high degree of imbrication between I1–3 as a diagnostic characteristic of
|
||
<taxonomicName id="4C18BFDCE921476A9EBAFCB5FEB3FCDE" authorityName="Ameghino" authorityYear="1887" box="[154,301,769,793]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9EBAFCB5FEB3FCDE" box="[154,301,769,793]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
, but we do not consider it diagnostic due to the following reasons. First, this is a variable characteristic within the analysed sample; in fact, some asymmetries concerning the degree of imbrication when comparing the less and right premaxilla have been identified in some specimens of this species (e.g.
|
||
<collectionCode id="ED095C9AE921476A9EC4FC29FED5FC72" box="[228,331,925,949]" pageId="14" pageNumber="431">MACN-A</collectionCode>
|
||
3920,
|
||
<collectionCode id="ED095C9AE921476A9FB5FC29FE78FC72" box="[405,486,925,949]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="14" pageNumber="431">AMNH</collectionCode>
|
||
9565,
|
||
<collectionCode id="ED095C9AE921476A9C10FC29FD09FC72" box="[560,663,925,949]" pageId="14" pageNumber="431">MACN-A</collectionCode>
|
||
9620;
|
||
<figureCitation id="1323D8DAE921476A9CC3FC29FF26FC12" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="14" pageNumber="431">Fig. 4E–F</figureCitation>
|
||
), as well as in
|
||
<taxonomicName id="4C18BFDCE921476A9F72FC09FE5EFC12" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[338,448,957,981]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE921476A9F72FC09FE5EFC12" box="[338,448,957,981]" italics="true" pageId="14" pageNumber="431">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<collectionCode id="ED095C9AE921476A9FD9FC09FDFEFC12" box="[505,608,957,981]" pageId="14" pageNumber="431">MACN-A</collectionCode>
|
||
9623–9633 and
|
||
<collectionCode id="ED095C9AE921476A9EA1FC68FF76FC33" box="[129,232,988,1012]" pageId="14" pageNumber="431">MACN-A</collectionCode>
|
||
9644;
|
||
<figureCitation id="1323D8DAE921476A9F0EFC68FEE4FC33" box="[302,378,988,1012]" captionStart="Figure 2" captionStartId="5.[113,178,1779,1803]" captionTargetBox="[189,1384,147,1747]" captionTargetId="figure-7@5.[186,1387,144,1750]" captionTargetPageId="5" captionText="Figure 2. Protypotherium australe (D–E, G, I–J, L) and Pr. praerutilum (A–C, F, H, K, M), in occlusal view. (A) YPM-VPPU 15386, incomplete skull; (B) less I3 (alveolus)–C–dP1–P2–M3 of MACN-A 4049; (C) less I3–dP1–M3 of MACN-A 4278; (D) right I1 (alveolus)–dP1–P2–M3 of MACN-A 3882; (E) right I1–dP1–P2–M3 of MACN-A 9623–9633; (F) less I1 (alveolus)–dP1–P2–M3 of AMNH 9187; (G) right I1–2 (alveolus)–dP1–P2–M3 of YPM-VPPU 15643; (H) MLP 12-2886, incomplete skull; (I) right I1 (alveolus)–I2–dP1–P2–M3 of MACN-A 37; (J) MACN-A 3911, less maxillary fragment with C–dP1–P2–M3; (K) MACN-A 1778, right maxillary fragment with dP1–P2 (alveoli)– P3–M3; (L) right I1 (alveolus)–dP1–P2–M3 of AMNH 9149; (M) MACN-A 4016, less maxillary fragment with P4–M3.* indicates alveolus. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426270" httpUri="https://zenodo.org/record/8426270/files/figure.png" pageId="14" pageNumber="431">Figs 2E</figureCitation>
|
||
,
|
||
<figureCitation id="1323D8DAE921476A9FA6FC68FE38FC33" box="[390,422,988,1012]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="14" pageNumber="431">4G</figureCitation>
|
||
). Second, the
|
||
<typeStatus id="54A37AFDE921476A9C1BFC68FD09FC33" box="[571,663,988,1012]" pageId="14" pageNumber="431" type="holotype">holotype</typeStatus>
|
||
of the species (
|
||
<collectionCode id="ED095C9AE921476A9E96FC4FFEBFFBD4" box="[182,289,1019,1043]" pageId="14" pageNumber="431">MACN-A</collectionCode>
|
||
1081–1082;
|
||
<figureCitation id="1323D8DAE921476A9F8FFC4FFE63FBD4" box="[431,509,1019,1043]" captionStart="Figure 3" captionStartId="6.[129,194,1775,1799]" captionTargetBox="[200,1403,146,1745]" captionTargetId="figure-6@6.[198,1405,144,1747]" captionTargetPageId="6" captionText="Figure 3. Protypotherium australe (B, F–M) and Pr. praerutilum (A, C–E), in occlusal view. (A) MACN-A 3952, right mandibular fragment with p3–m3; (B) less dp1–p2–m3 of AMNH 9286; (C) MACN-A 11191a, right mandibular fragment with m1–3; (D) right p2–m3 of MACN-A 9614; (E) right i1–dp1–p2–m3 of MACN-A 1082; (F) less p4–m3 of AMNH 9565; (G) right p2–m3 of AMNH 9149; (H) less i1–dp1– p2–m3 of MPM-PV 3531; (I) MACN-A 532, right mandibular fragment with m2–3; (J) right p2–m3 of MPM-PV 3739; (K) MPM-PV 19612, right mandibular fragment with m2 (broken)–3; (L) MACN-A 3979, less mandibular fragment with p3–m3; (M) MACN-A 3955, less mandibular fragment with p3–m3. * indicates alveolus; ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426272" httpUri="https://zenodo.org/record/8426272/files/figure.png" pageId="14" pageNumber="431">Figs 3E</figureCitation>
|
||
,
|
||
<figureCitation id="1323D8DAE921476A9C2BFC4FFDB5FBD4" box="[523,555,1019,1043]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="14" pageNumber="431">4D</figureCitation>
|
||
; see
|
||
<bibRefCitation id="EF89B9AEE921476A9C44FC4FFF2BFBF4" author="Fernandez M & Fernicola JC & Cerdeno E" pageId="14" pageNumber="431" pagination="473 - 98" refId="ref23394" refString="Fernandez M, Fernicola JC, Cerdeno E. Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino. Zootaxa 2018; 4387: 473 - 98. hups: // doi. org / 10.11646 / ZOOTAXA. 4387.3.4" type="journal article" year="2018">
|
||
Fernández
|
||
<emphasis id="B96C184DE921476A9CF8FC48FC94FBD4" box="[728,778,1019,1043]" italics="true" pageId="14" pageNumber="431">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
) does not show this sort of imbrication; in fact, its degree of imbrication is low.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE921476A9EBCFBEDFABAFE64" blockId="14.[128,780,487,1983]" lastBlockId="14.[825,1477,144,1985]" pageId="14" pageNumber="431">
|
||
Several species are here considered, for the first time, as junior synonyms of
|
||
<emphasis id="B96C184DE921476A9F6DFBCDFE79FB57" box="[333,487,1144,1168]" italics="true" pageId="14" pageNumber="431">
|
||
<taxonomicName id="4C18BFDCE921476A9F6DFBCDFE7DFB57" authorityName="Ameghino" authorityYear="1887" box="[333,483,1144,1168]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">Pr. praerutilum</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Traditionally,
|
||
<taxonomicName id="4C18BFDCE921476A9C59FBCDFC97FB57" authorityName="Ameghino" authorityYear="1887" box="[633,777,1145,1168]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="attenuatum">
|
||
<emphasis id="B96C184DE921476A9C59FBCDFC97FB57" box="[633,777,1145,1168]" italics="true" pageId="14" pageNumber="431">Pr. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE921476A9E91FB2DFED4FB77" authorityName="Ameghino" authorityYear="1887" box="[177,330,1176,1200]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9E91FB2DFED4FB77" box="[177,330,1176,1200]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
were considered separate taxa. According to Sinclair’s (1909) proposal, supported by
|
||
<bibRefCitation id="EF89B9AEE921476A9C68FB03FD7CFB08" author="Tauber AA" box="[584,738,1207,1231]" pageId="14" pageNumber="431" pagination="1 - 28" refId="ref25150" refString="Tauber AA. Los representantes del genero Protypotherium (Mammalia, Notoungulata, Interatheriidae) del Mioceno Temprano del Sudeste de la Provincia de Santa Cruz, Republica Argentina. Academia Nacional de Ciencias, Cordoba, Misc 1996; 95: 1 - 28." type="journal article" year="1996">Tauber (1996)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C18BFDCE921476A9CCFFB0CFF6FFB29" authorityName="Ameghino" authorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="attenuatum">
|
||
<emphasis id="B96C184DE921476A9CCFFB0CFF6FFB29" italics="true" pageId="14" pageNumber="431">Pr. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C18BFDCE921476A9F51FB63FD9BFB29" authorityName="Ameghino" authorityYear="1887" box="[369,517,1238,1262]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9F51FB63FD9BFB29" box="[369,517,1238,1262]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
mainly by its smaller size. However, according to the measurements provided by Sinclair (1909), the
|
||
<specimenCount id="9D1E0FD6E921476A9F20FAA1FE39FAEA" box="[256,423,1301,1325]" pageId="14" pageNumber="431" type="generic">three specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="4C18BFDCE921476A9FEBFAA2FDFFFAEA" authorityName="Ameghino" authorityYear="1887" box="[459,609,1301,1325]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9FEBFAA2FDFFFAEA" box="[459,609,1301,1325]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
(e.g. L M1 = 6.0–
|
||
<quantity id="4CE069BAE921476A9EA1FA81FF4EFA8B" box="[129,208,1333,1357]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="14" pageNumber="431" unit="mm" value="6.6">6.6 mm</quantity>
|
||
; W
|
||
<subScript id="179CC61AE921476A9ED3FAF1FE96FA94" attach="left" box="[243,264,1349,1363]" fontSize="6" pageId="14" pageNumber="431">M1</subScript>
|
||
=
|
||
<quantity id="4CE069BAE921476A9F05FA80FEEBFA8B" box="[293,373,1332,1356]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="431" unit="mm" value="4.5">4.5 mm</quantity>
|
||
; L
|
||
<subScript id="179CC61AE921476A9FADFAF1FE3EFA94" attach="left" box="[397,416,1349,1363]" fontSize="6" pageId="14" pageNumber="431">m1</subScript>
|
||
=
|
||
<quantity id="4CE069BAE921476A9F9DFA80FD93FA8B" box="[445,525,1332,1356]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="14" pageNumber="431" unit="mm" value="5.7">5.7 mm</quantity>
|
||
; W
|
||
<subScript id="179CC61AE921476A9C0DFAF1FDDEFA94" attach="left" box="[557,576,1349,1363]" fontSize="6" pageId="14" pageNumber="431">m1</subScript>
|
||
= 3.0 mm) and two of
|
||
<taxonomicName id="4C18BFDCE921476A9EBDFAE1FEB1FAAB" authorityName="Ameghino" authorityYear="1887" box="[157,303,1365,1388]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="attenuatum">
|
||
<emphasis id="B96C184DE921476A9EBDFAE1FEB1FAAB" box="[157,303,1365,1388]" italics="true" pageId="14" pageNumber="431">Pr. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
(e.g. L
|
||
<subScript id="179CC61AE921476A9F59FAD0FE13FAB5" attach="left" box="[377,397,1380,1394]" fontSize="6" pageId="14" pageNumber="431">M1</subScript>
|
||
= 6.0–
|
||
<quantity id="4CE069BAE921476A9FF8FAE0FDB5FAAC" box="[472,555,1364,1388]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="14" pageNumber="431" unit="mm" value="6.5">6.5 mm</quantity>
|
||
; W
|
||
<subScript id="179CC61AE921476A9C73FAD0FDF6FAB5" attach="left" box="[595,616,1380,1394]" fontSize="6" pageId="14" pageNumber="431">M1</subScript>
|
||
= 4.0–
|
||
<quantity id="4CE069BAE921476A9C92FAE0FC9BFAAC" box="[690,773,1364,1388]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="14" pageNumber="431" unit="mm" value="4.6">4.6 mm</quantity>
|
||
; L m1 =
|
||
<quantity id="4CE069BAE921476A9EE0FAC7FE8DFA4C" box="[192,275,1395,1419]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="431" unit="mm" value="5.5">5.5 mm</quantity>
|
||
; W m1 = 3.0 mm) completely overlap. In order to resolve this issue, we analysed the dental dimensions of M1–2 and m1–2 of the Santacrucian specimens of
|
||
<taxonomicName id="4C18BFDCE921476A9C4FFA06FC97FA0D" authorityName="Ameghino" authorityYear="1885" box="[623,777,1458,1482]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96C184DE921476A9C4FFA06FC97FA0D" box="[623,777,1458,1482]" italics="true" pageId="14" pageNumber="431">Protypotherium</emphasis>
|
||
</taxonomicName>
|
||
(note that
|
||
<taxonomicName id="4C18BFDCE921476A9ECBFA66FEC0FA2E" authorityName="Ameghino" authorityYear="1889" box="[235,350,1489,1513]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="claudum">
|
||
<emphasis id="B96C184DE921476A9ECBFA66FEC0FA2E" box="[235,350,1489,1513]" italics="true" pageId="14" pageNumber="431">Pr. claudum</emphasis>
|
||
</taxonomicName>
|
||
was excluded due to the existence of morphological differences that allow the species characterization; see below). When comparing M1–2 (
|
||
<figureCitation id="1323D8DAE921476A9FFEF9A4FDB5F9EF" box="[478,555,1552,1576]" captionStart="Figure 5" captionStartId="8.[129,194,1753,1777]" captionTargetBox="[257,1345,147,1723]" captionTargetId="figure-83@8.[254,1348,144,1726]" captionTargetPageId="8" captionText="Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426276" httpUri="https://zenodo.org/record/8426276/files/figure.png" pageId="14" pageNumber="431">Figs 5A</figureCitation>
|
||
,
|
||
<figureCitation id="1323D8DAE921476A9C18F9A4FDCBF9EF" box="[568,597,1552,1576]" captionStart="Figure 6" captionStartId="9.[112,177,1757,1781]" captionTargetBox="[242,1330,147,1727]" captionTargetId="figure-87@9.[239,1333,144,1730]" captionTargetPageId="9" captionText="Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426278" httpUri="https://zenodo.org/record/8426278/files/figure.png" pageId="14" pageNumber="431">6A</figureCitation>
|
||
) and m1–2 (
|
||
<figureCitation id="1323D8DAE921476A9CC0F9A4FF02F980" captionStart="Figure 5" captionStartId="8.[129,194,1753,1777]" captionTargetBox="[257,1345,147,1723]" captionTargetId="figure-83@8.[254,1348,144,1726]" captionTargetPageId="8" captionText="Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426276" httpUri="https://zenodo.org/record/8426276/files/figure.png" pageId="14" pageNumber="431">Figs 5B</figureCitation>
|
||
,
|
||
<figureCitation id="1323D8DAE921476A9E8BF99BFF58F980" box="[171,198,1583,1607]" captionStart="Figure 6" captionStartId="9.[112,177,1757,1781]" captionTargetBox="[242,1330,147,1727]" captionTargetId="figure-87@9.[239,1333,144,1730]" captionTargetPageId="9" captionText="Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426278" httpUri="https://zenodo.org/record/8426278/files/figure.png" pageId="14" pageNumber="431">6B</figureCitation>
|
||
) dimensions, the scaưer diagrams show three different groups (
|
||
<figureCitation id="1323D8DAE921476A9EF8F9FBFE8BF9A1" box="[216,277,1614,1639]" captionStart="Figure 5" captionStartId="8.[129,194,1753,1777]" captionTargetBox="[257,1345,147,1723]" captionTargetId="figure-83@8.[254,1348,144,1726]" captionTargetPageId="8" captionText="Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426276" httpUri="https://zenodo.org/record/8426276/files/figure.png" pageId="14" pageNumber="431">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="1323D8DAE921476A9F01F9FBFEB0F9A0" box="[289,302,1615,1639]" captionStart="Figure 6" captionStartId="9.[112,177,1757,1781]" captionTargetBox="[242,1330,147,1727]" captionTargetId="figure-87@9.[239,1333,144,1730]" captionTargetPageId="9" captionText="Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426278" httpUri="https://zenodo.org/record/8426278/files/figure.png" pageId="14" pageNumber="431">6</figureCitation>
|
||
), particularly more evident in M/m1, that are named asser the valid name of the species: (1) Morphotype A or
|
||
<taxonomicName id="4C18BFDCE921476A9EBFF93AFE90F962" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[159,270,1677,1701]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE921476A9EBFF93AFE90F962" box="[159,270,1677,1701]" italics="true" pageId="14" pageNumber="431">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
, which includes the
|
||
<typeStatus id="54A37AFDE921476A9FC8F939FDDAF962" box="[488,580,1677,1701]" pageId="14" pageNumber="431" type="holotype">holotype</typeStatus>
|
||
of the species and the referred materials, including the type specimens of the considered junior synonyms of
|
||
<taxonomicName id="4C18BFDCE921476A9F89F979FD86F923" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[425,536,1740,1764]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE921476A9F89F979FD86F923" box="[425,536,1740,1764]" italics="true" pageId="14" pageNumber="431">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
(see below for further justifications); (2) Morphotype B or
|
||
<taxonomicName id="4C18BFDCE921476A9FDAF958FD10F8C4" authorityName="Ameghino" authorityYear="1887" box="[506,654,1771,1795]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9FDAF958FD10F8C4" box="[506,654,1771,1795]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
, which contains the
|
||
<typeStatus id="54A37AFDE921476A9EFDF8BEFEA7F8E5" box="[221,313,1802,1826]" pageId="14" pageNumber="431" type="holotype">holotype</typeStatus>
|
||
of the species, the
|
||
<typeStatus id="54A37AFDE921476A9FD9F8BFFDCDF8E4" box="[505,595,1803,1827]" pageId="14" pageNumber="431" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C18BFDCE921476A9C53F8BFFC9DF8E5" authorityName="Ameghino" authorityYear="1887" box="[627,771,1803,1826]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="attenuatum">
|
||
<emphasis id="B96C184DE921476A9C53F8BFFC9DF8E5" box="[627,771,1803,1826]" italics="true" pageId="14" pageNumber="431">Pr. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
, and the referred materials of
|
||
<taxonomicName id="4C18BFDCE921476A9F8BF89FFDDFF885" authorityName="Ameghino" authorityYear="1887" box="[427,577,1834,1858]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9F8BF89FFDDFF885" box="[427,577,1834,1858]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
, including the type specimens of the junior synonyms (see below for further justifications); and (3) Morphotype C or
|
||
<taxonomicName id="4C18BFDCE921476A9C0AF8DDFD49F847" authorityName="Ameghino" authorityYear="1891" box="[554,727,1896,1920]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="compressidens">
|
||
<emphasis id="B96C184DE921476A9C0AF8DDFD49F847" box="[554,727,1896,1920]" italics="true" pageId="14" pageNumber="431">Pr. compressidens</emphasis>
|
||
</taxonomicName>
|
||
that only includes the
|
||
<typeStatus id="54A37AFDE921476A9F16F83CFE0CF867" box="[310,402,1928,1952]" pageId="14" pageNumber="431" type="holotype">holotype</typeStatus>
|
||
and a few referred specimens herein considered. It is worth mentioning that in the case of the third species, the grouping regarding m1 is not as conspicuous as in the other two groups, due to the number of specimens assignable to
|
||
<taxonomicName id="4C18BFDCE921476A9DA6FF7BFBB1FF21" authorityName="Ameghino" authorityYear="1891" box="[902,1071,206,230]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="compressidens">
|
||
<emphasis id="B96C184DE921476A9DA6FF7BFBB1FF21" box="[902,1071,206,230]" italics="true" pageId="14" pageNumber="431">Pr. compressidens</emphasis>
|
||
</taxonomicName>
|
||
(a minimum of three is required to establish the grouping). Nevertheless, the values do not fall within the other two groups (
|
||
<figureCitation id="1323D8DAE921476A9A14FEB9FB1EFEE2" box="[1076,1152,269,293]" captionStart="Figure 5" captionStartId="8.[129,194,1753,1777]" captionTargetBox="[257,1345,147,1723]" captionTargetId="figure-83@8.[254,1348,144,1726]" captionTargetPageId="8" captionText="Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426276" httpUri="https://zenodo.org/record/8426276/files/figure.png" pageId="14" pageNumber="431">Fig. 5B</figureCitation>
|
||
), showing a differentiation in terms of length and width of m1. In the case of m2, this differentiation is less evident, because only the
|
||
<typeStatus id="54A37AFDE921476A9AC3FEF8FAA1FEA3" box="[1251,1343,332,356]" pageId="14" pageNumber="431" type="holotype">holotype</typeStatus>
|
||
does not fall within the other two groups, leaving the remaining specimens within the size range of
|
||
<taxonomicName id="4C18BFDCE921476A9A09FE3FFB23FE65" authorityName="Ameghino" authorityYear="1887" box="[1065,1213,394,418]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9A09FE3FFB23FE65" box="[1065,1213,394,418]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1323D8DAE921476A9AEDFE3EFA8AFE65" box="[1229,1300,394,418]" captionStart="Figure 6" captionStartId="9.[112,177,1757,1781]" captionTargetBox="[242,1330,147,1727]" captionTargetId="figure-87@9.[239,1333,144,1730]" captionTargetPageId="9" captionText="Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426278" httpUri="https://zenodo.org/record/8426278/files/figure.png" pageId="14" pageNumber="431">Fig. 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE921476A9D75FE1EFBEAFB76" blockId="14.[825,1477,144,1985]" pageId="14" pageNumber="431">
|
||
Excluding
|
||
<taxonomicName id="4C18BFDCE921476A9DE7FE1EFBE8FE05" authorityName="Ameghino" authorityYear="1891" box="[967,1142,426,450]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="compressidens">
|
||
<emphasis id="B96C184DE921476A9DE7FE1EFBE8FE05" box="[967,1142,426,450]" italics="true" pageId="14" pageNumber="431">Pr. compressidens</emphasis>
|
||
</taxonomicName>
|
||
from the analysis, due to the presence of morphological differences with
|
||
<taxonomicName id="4C18BFDCE921476A9B22FE7EFAEFFE26" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[1282,1393,457,481]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE921476A9B22FE7EFAEFFE26" box="[1282,1393,457,481]" italics="true" pageId="14" pageNumber="431">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE921476A9B87FE7EFC33FDC7" authorityName="Ameghino" authorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9B87FE7EFC33FDC7" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
(see below), Morphotypes A and B were statistically tested in order to prove if they were statistically different in terms of the dental dimensions. All the statistical details are shown in Supporting Information, Appendix S4, and summarized as follows. In general terms, the requirement of normal distribution was not met for the length and/or width of M/m1–2 of
|
||
<taxonomicName id="4C18BFDCE921476A9D74FD11FC5FFD7B" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[852,961,676,700]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE921476A9D74FD11FC5FFD7B" box="[852,961,676,700]" italics="true" pageId="14" pageNumber="431">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE921476A9DD5FD11FB14FD7B" authorityName="Ameghino" authorityYear="1887" box="[1013,1162,676,700]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9DD5FD11FB14FD7B" box="[1013,1162,676,700]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
(Shapiro–Wilk’s test,
|
||
<emphasis id="B96C184DE921476A9B4CFD11FAE4FD7B" box="[1388,1402,677,700]" italics="true" pageId="14" pageNumber="431">P</emphasis>
|
||
<.01) and all datasets show a non-homogeneous variance (Levene’s test,
|
||
<emphasis id="B96C184DE921476A9D4BFD50FCE7FD3C" box="[875,889,740,763]" italics="true" pageId="14" pageNumber="431">P</emphasis>
|
||
<.05). Given the non-compliance of both necessary requirements to carry out a Student’s
|
||
<emphasis id="B96C184DE921476A9AE8FCB7FB4EFCDD" box="[1224,1232,771,794]" italics="true" pageId="14" pageNumber="431">t</emphasis>
|
||
-test, a non-parametric study was used, in this case a Mann–Whitney U Test, in order to compare the tooth dimensions between the two groups. All samples analysed with a Mann-Whitney U test (α = 5%) showed a significant difference between dental dimensions (M/m1–2) for Morphotypes A and B (U = 0,
|
||
<emphasis id="B96C184DE921476A9AA0FC14FB10FC70" box="[1152,1166,928,951]" italics="true" pageId="14" pageNumber="431">P</emphasis>
|
||
<.05). Therefore,
|
||
<taxonomicName id="4C18BFDCE921476A9B73FC14FA5CFC70" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[1363,1474,927,951]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE921476A9B73FC14FA5CFC70" box="[1363,1474,927,951]" italics="true" pageId="14" pageNumber="431">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE921476A9D48FC0BFB9EFC11" authorityName="Ameghino" authorityYear="1887" box="[872,1024,958,982]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9D48FC0BFB9EFC11" box="[872,1024,958,982]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
statistically differ based on the length and width of M/m1–2. P. In this framework,
|
||
<taxonomicName id="4C18BFDCE921476A9AECFC6AFAA8FC31" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[1228,1334,990,1014]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE921476A9AECFC6AFAA8FC31" box="[1228,1334,990,1014]" italics="true" pageId="14" pageNumber="431">Pr.australe</emphasis>
|
||
</taxonomicName>
|
||
involves large individuals (L M1 ≥
|
||
<quantity id="4CE069BAE921476A9DDFFC49FBCAFBD2" box="[1023,1108,1021,1045]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="14" pageNumber="431" unit="mm" value="7.1">7.1 mm</quantity>
|
||
; W M1 ≥
|
||
<quantity id="4CE069BAE921476A9A8CFC49FA90FBD2" box="[1196,1294,1021,1045]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" pageId="14" pageNumber="431" unit="mm" value="4.15">4.15 mm</quantity>
|
||
; L m1 ≥
|
||
<quantity id="4CE069BAE921476A9B7CFC49FA20FBD2" box="[1372,1470,1021,1045]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" pageId="14" pageNumber="431" unit="mm" value="7.15">7.15 mm</quantity>
|
||
; W
|
||
<subScript id="179CC61AE921476A9D70FB99FCFDFBFC" attach="left" box="[848,867,1069,1083]" fontSize="6" pageId="14" pageNumber="431">m1</subScript>
|
||
≥
|
||
<quantity id="4CE069BAE921476A9DA0FBA8FC4DFBF2" box="[896,979,1052,1077]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="431" unit="mm" value="3.5">3.5 mm</quantity>
|
||
) (
|
||
<figureCitation id="1323D8DAE921476A9DCDFBA9FBB2FBF3" box="[1005,1068,1052,1077]" captionStart="Figure 5" captionStartId="8.[129,194,1753,1777]" captionTargetBox="[257,1345,147,1723]" captionTargetId="figure-83@8.[254,1348,144,1726]" captionTargetPageId="8" captionText="Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426276" httpUri="https://zenodo.org/record/8426276/files/figure.png" pageId="14" pageNumber="431">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="1323D8DAE921476A9A1AFBA9FBD9FBF2" box="[1082,1095,1053,1077]" captionStart="Figure 6" captionStartId="9.[112,177,1757,1781]" captionTargetBox="[242,1330,147,1727]" captionTargetId="figure-87@9.[239,1333,144,1730]" captionTargetPageId="9" captionText="Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426278" httpUri="https://zenodo.org/record/8426278/files/figure.png" pageId="14" pageNumber="431">6</figureCitation>
|
||
), whereas
|
||
<taxonomicName id="4C18BFDCE921476A9A9BFBA9FACFFBF2" authorityName="Ameghino" authorityYear="1887" box="[1211,1361,1053,1077]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9A9BFBA9FACFFBF2" box="[1211,1361,1053,1077]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
represents small to medium individuals (L
|
||
<subScript id="179CC61AE921476A9AAAFBF8FB01FB9D" attach="left" box="[1162,1183,1100,1114]" fontSize="6" pageId="14" pageNumber="431">M1</subScript>
|
||
≤
|
||
<quantity id="4CE069BAE921476A9A9CFB88FA8FFB94" box="[1212,1297,1084,1108]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="14" pageNumber="431" unit="mm" value="6.8">6.8 mm</quantity>
|
||
; W
|
||
<subScript id="179CC61AE921476A9B17FBF8FAD2FB9D" attach="left" box="[1335,1356,1100,1114]" fontSize="6" pageId="14" pageNumber="431">M1</subScript>
|
||
≤ 5.0 mm; L
|
||
<subScript id="179CC61AE921476A9D69FBD8FCC2FBBD" attach="left" box="[841,860,1132,1146]" fontSize="6" pageId="14" pageNumber="431">m1</subScript>
|
||
≤
|
||
<quantity id="4CE069BAE921476A9D59FBEFFC50FBB4" box="[889,974,1115,1139]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="14" pageNumber="431" unit="mm" value="6.8">6.8 mm</quantity>
|
||
; W
|
||
<subScript id="179CC61AE921476A9DD3FBD8FB98FBBD" attach="left" box="[1011,1030,1132,1146]" fontSize="6" pageId="14" pageNumber="431">m1</subScript>
|
||
≤ 4.0 mm) (
|
||
<figureCitation id="1323D8DAE921476A9AB3FBEFFB4AFBB4" box="[1171,1236,1115,1139]" captionStart="Figure 5" captionStartId="8.[129,194,1753,1777]" captionTargetBox="[257,1345,147,1723]" captionTargetId="figure-83@8.[254,1348,144,1726]" captionTargetPageId="8" captionText="Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426276" httpUri="https://zenodo.org/record/8426276/files/figure.png" pageId="14" pageNumber="431">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="1323D8DAE921476A9AC3FBEFFB6EFBB4" box="[1251,1264,1115,1139]" captionStart="Figure 6" captionStartId="9.[112,177,1757,1781]" captionTargetBox="[242,1330,147,1727]" captionTargetId="figure-87@9.[239,1333,144,1730]" captionTargetPageId="9" captionText="Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426278" httpUri="https://zenodo.org/record/8426278/files/figure.png" pageId="14" pageNumber="431">6</figureCitation>
|
||
). Our analysis suggests no significant difference in the size of M/m1–2 between
|
||
<taxonomicName id="4C18BFDCE921476A9B87FBCFFC33FB75" authorityName="Ameghino" authorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9B87FBCFFC33FB75" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE921476A9DFEFB2FFBF3FB75" authorityName="Ameghino" authorityYear="1887" box="[990,1133,1179,1202]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="attenuatum">
|
||
<emphasis id="B96C184DE921476A9DFEFB2FFBF3FB75" box="[990,1133,1179,1202]" italics="true" pageId="14" pageNumber="431">Pr. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE921476A9D75FB0DFCE0F94F" blockId="14.[825,1477,144,1985]" pageId="14" pageNumber="431">
|
||
Besides size, another character traditionally used to differentiate
|
||
<taxonomicName id="4C18BFDCE921476A9D4EFB6DFB9BFB36" authorityName="Ameghino" authorityYear="1887" box="[878,1029,1241,1265]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9D4EFB6DFB9BFB36" box="[878,1029,1241,1265]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE921476A9A1CFB6DFB51FB37" authorityName="Ameghino" authorityYear="1887" box="[1084,1231,1241,1264]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="attenuatum">
|
||
<emphasis id="B96C184DE921476A9A1CFB6DFB51FB37" box="[1084,1231,1241,1264]" italics="true" pageId="14" pageNumber="431">Pr. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
is the curvature of the cheek tooth series, but when comparing the curvature of the
|
||
<typeStatus id="54A37AFDE921476A9D19FAA3FC01FAE8" box="[825,927,1303,1327]" pageId="14" pageNumber="431" type="holotype">holotypes</typeStatus>
|
||
of both species, independent from size and taking into consideration that
|
||
<taxonomicName id="4C18BFDCE921476A9A18FA83FB50FA89" authorityName="Ameghino" authorityYear="1887" box="[1080,1230,1335,1358]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="attenuatum">
|
||
<emphasis id="B96C184DE921476A9A18FA83FB50FA89" box="[1080,1230,1335,1358]" italics="true" pageId="14" pageNumber="431">Pr. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
involves a fragmented maxilla, this feature does not differ. We argue that the morphological differences between
|
||
<taxonomicName id="4C18BFDCE921476A9A7DFAC2FB6BFA4A" authorityName="Ameghino" authorityYear="1887" box="[1117,1269,1397,1421]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9A7DFAC2FB6BFA4A" box="[1117,1269,1397,1421]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE921476A9B0EFAC2FA5CFA4A" authorityName="Ameghino" authorityYear="1887" box="[1326,1474,1398,1421]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="attenuatum">
|
||
<emphasis id="B96C184DE921476A9B0EFAC2FA5CFA4A" box="[1326,1474,1398,1421]" italics="true" pageId="14" pageNumber="431">Pr. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
can be explained by ontogenetic and/or intraspecific variation, and because our statistical analyses demonstrate that size is not a diagnostic feature in order to separate both species, we consider them synonyms. As the name priority is determined by the reviewer who takes notice of this and not by the page priority (
|
||
<collectionCode id="ED095C9AE921476A9D63F985FC1CF98E" box="[835,898,1585,1609]" pageId="14" pageNumber="431">CINZ</collectionCode>
|
||
2000), we select
|
||
<taxonomicName id="4C18BFDCE921476A9A1BF986FB4FF98E" authorityName="Ameghino" authorityYear="1887" box="[1083,1233,1585,1609]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9A1BF986FB4FF98E" box="[1083,1233,1585,1609]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
as the senior synonym because its
|
||
<typeStatus id="54A37AFDE921476A9D97F9E4FB8DF9AF" box="[951,1043,1616,1640]" pageId="14" pageNumber="431" type="holotype">holotype</typeStatus>
|
||
is more complete (see
|
||
<bibRefCitation id="EF89B9AEE921476A9B38F9E4FCF3F94F" author="Fernandez M & Fernicola JC & Cerdeno E" pageId="14" pageNumber="431" pagination="473 - 98" refId="ref23394" refString="Fernandez M, Fernicola JC, Cerdeno E. Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino. Zootaxa 2018; 4387: 473 - 98. hups: // doi. org / 10.11646 / ZOOTAXA. 4387.3.4" type="journal article" year="2018">
|
||
Fernández
|
||
<emphasis id="B96C184DE921476A9BAFF9E5FA5DF9AF" box="[1423,1475,1616,1640]" italics="true" pageId="14" pageNumber="431">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE921476A9D75F924FAC1F883" blockId="14.[825,1477,144,1985]" pageId="14" pageNumber="431">
|
||
In contrast to Sinclair (1909), who proposed that
|
||
<taxonomicName id="4C18BFDCE921476A9B7FF924FA5CF960" authorityName="Ameghino" authorityYear="1894" box="[1375,1474,1679,1703]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="lineare">
|
||
<emphasis id="B96C184DE921476A9B7FF924FA5CF960" box="[1375,1474,1679,1703]" italics="true" pageId="14" pageNumber="431">Pr. lineare</emphasis>
|
||
</taxonomicName>
|
||
was the junior synonym of
|
||
<taxonomicName id="4C18BFDCE921476A9A69F91BFB2AF901" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[1097,1204,1710,1734]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE921476A9A69F91BFB2AF901" box="[1097,1204,1710,1734]" italics="true" pageId="14" pageNumber="431">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
based on its size (followed by
|
||
<bibRefCitation id="EF89B9AEE921476A9D77F97AFC47F921" author="Tauber AA" box="[855,985,1742,1766]" pageId="14" pageNumber="431" pagination="1 - 28" refId="ref25150" refString="Tauber AA. Los representantes del genero Protypotherium (Mammalia, Notoungulata, Interatheriidae) del Mioceno Temprano del Sudeste de la Provincia de Santa Cruz, Republica Argentina. Academia Nacional de Ciencias, Cordoba, Misc 1996; 95: 1 - 28." type="journal article" year="1996">Tauber 1996</bibRefCitation>
|
||
), we consider the former as the junior synonym of
|
||
<taxonomicName id="4C18BFDCE921476A9D77F95AFC70F8C2" authorityName="Ameghino" authorityYear="1887" box="[855,1006,1773,1797]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9D77F95AFC70F8C2" box="[855,1006,1773,1797]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
, because the size of its
|
||
<typeStatus id="54A37AFDE921476A9ACBF959FAD9F8C2" box="[1259,1351,1773,1797]" pageId="14" pageNumber="431" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED095C9AE921476A9B77F959FA5CF8C2" box="[1367,1474,1773,1797]" pageId="14" pageNumber="431">MACN-A</collectionCode>
|
||
4038–4039; see
|
||
<bibRefCitation id="EF89B9AEE921476A9DC0F8B8FB28F8E2" author="Fernandez M & Fernicola JC & Cerdeno E" box="[992,1206,1804,1829]" pageId="14" pageNumber="431" pagination="473 - 98" refId="ref23394" refString="Fernandez M, Fernicola JC, Cerdeno E. Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino. Zootaxa 2018; 4387: 473 - 98. hups: // doi. org / 10.11646 / ZOOTAXA. 4387.3.4" type="journal article" year="2018">
|
||
Fernández
|
||
<emphasis id="B96C184DE921476A9A70F8B9FBE0F8E3" box="[1104,1150,1804,1828]" italics="true" pageId="14" pageNumber="431">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
) falls within the size range of
|
||
<taxonomicName id="4C18BFDCE921476A9D73F899FC79F883" authorityName="Ameghino" authorityYear="1887" box="[851,999,1836,1860]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE921476A9D73F899FC79F883" box="[851,999,1836,1860]" italics="true" pageId="14" pageNumber="431">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
(Supporting Information, Table S2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30297D4E921476B9D75F8FFFE6CFA4A" lastPageId="15" lastPageNumber="432" pageId="14" pageNumber="431" type="reference_group">
|
||
<paragraph id="8BA7C45FE921476B9D75F8FFFD45FC5F" blockId="14.[825,1477,144,1985]" lastBlockId="15.[113,765,144,1421]" lastPageId="15" lastPageNumber="432" pageId="14" pageNumber="431">
|
||
<taxonomicName id="4C18BFDCE921476A9D75F8FFFB94F8A4" authorityName="Roth" authorityYear="1899" box="[853,1034,1867,1891]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="species" species="endiadys">
|
||
<emphasis id="B96C184DE921476A9D75F8FFFB94F8A4" box="[853,1034,1867,1891]" italics="true" pageId="14" pageNumber="431">Icochilus endiadys</emphasis>
|
||
</taxonomicName>
|
||
, from the Collon Curá Fm. (
|
||
<collectingRegion id="49DC0ABDE921476A9B7FF8FFFC06F845" country="Argentina" name="Neuquen" pageId="14" pageNumber="431">Neuquén Province</collectingRegion>
|
||
;
|
||
<bibRefCitation id="EF89B9AEE921476A9D82F8DEFB8FF844" author="Roth S" box="[930,1041,1898,1923]" pageId="14" pageNumber="431" pagination="141 - 97" refId="ref24723" refString="Roth S. Apuntes sobre la geologia y paleontologia de los territorios de Rio Negro y Neuquen (diciembre de 1895 a junio de 1896). Revista del Museo de La Plata 1899; 9: 141 - 97." type="journal article" year="1899">Roth 1899</bibRefCitation>
|
||
), was recently included in
|
||
<taxonomicName id="4C18BFDCE921476A9B08F8DEFA5CF845" authorityName="Ameghino" authorityYear="1885" box="[1320,1474,1898,1922]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="431" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96C184DE921476A9B08F8DEFA5CF845" box="[1320,1474,1898,1922]" italics="true" pageId="14" pageNumber="431">Protypotherium</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF89B9AEE921476A9D7CF83EFB87F865" author="Vera B & Reguero MA & Gonzalez-Ruiz L" box="[860,1049,1930,1954]" pageId="14" pageNumber="431" pagination="845 - 63" refId="ref25562" refString="Vera B, Reguero MA, Gonzalez-Ruiz L. The Interatheriinae notoungulates from the Middle Miocene Collon Cura Formation in Argentina. Acta Palaeontologica Polonica 2017; 62: 845 - 63. hups: // dx. doi. org / 10.4202 / app. 00373.2017" type="journal article" year="2017">
|
||
Vera
|
||
<emphasis id="B96C184DE921476A9DB3F83EFC58F865" box="[915,966,1930,1954]" italics="true" pageId="14" pageNumber="431">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
. These authors mentioned the presence of a reduced first upper premolar that was sub-circular in cross section as a diagnostic feature of
|
||
<taxonomicName id="4C18BFDCE920476B9FF4FF24FD55FF6F" baseAuthorityName="Roth" baseAuthorityYear="1898" box="[468,715,144,168]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="endiadys">
|
||
<emphasis id="B96C184DE920476B9FF4FF24FD55FF6F" box="[468,715,144,168]" italics="true" pageId="15" pageNumber="432">Protypotherium endiadys</emphasis>
|
||
</taxonomicName>
|
||
and stated that this condition is seen in the
|
||
<typeStatus id="54A37AFDE920476B9C2BFF1BFDF9FF00" box="[523,615,175,199]" pageId="15" pageNumber="432" type="holotype">holotype</typeStatus>
|
||
of the species MLP 12-2886 (
|
||
<figureCitation id="1323D8DAE920476B9F36FF7BFEFBFF20" box="[278,357,207,231]" captionStart="Figure 2" captionStartId="5.[113,178,1779,1803]" captionTargetBox="[189,1384,147,1747]" captionTargetId="figure-7@5.[186,1387,144,1750]" captionTargetPageId="5" captionText="Figure 2. Protypotherium australe (D–E, G, I–J, L) and Pr. praerutilum (A–C, F, H, K, M), in occlusal view. (A) YPM-VPPU 15386, incomplete skull; (B) less I3 (alveolus)–C–dP1–P2–M3 of MACN-A 4049; (C) less I3–dP1–M3 of MACN-A 4278; (D) right I1 (alveolus)–dP1–P2–M3 of MACN-A 3882; (E) right I1–dP1–P2–M3 of MACN-A 9623–9633; (F) less I1 (alveolus)–dP1–P2–M3 of AMNH 9187; (G) right I1–2 (alveolus)–dP1–P2–M3 of YPM-VPPU 15643; (H) MLP 12-2886, incomplete skull; (I) right I1 (alveolus)–I2–dP1–P2–M3 of MACN-A 37; (J) MACN-A 3911, less maxillary fragment with C–dP1–P2–M3; (K) MACN-A 1778, right maxillary fragment with dP1–P2 (alveoli)– P3–M3; (L) right I1 (alveolus)–dP1–P2–M3 of AMNH 9149; (M) MACN-A 4016, less maxillary fragment with P4–M3.* indicates alveolus. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426270" httpUri="https://zenodo.org/record/8426270/files/figure.png" pageId="15" pageNumber="432">Fig. 2H</figureCitation>
|
||
). Although the less dP1 (P1 according to
|
||
<bibRefCitation id="EF89B9AEE920476B9EACFF5AFEB8FEC1" author="Vera B & Reguero MA & Gonzalez-Ruiz L" box="[140,294,238,262]" pageId="15" pageNumber="432" pagination="845 - 63" refId="ref25562" refString="Vera B, Reguero MA, Gonzalez-Ruiz L. The Interatheriinae notoungulates from the Middle Miocene Collon Cura Formation in Argentina. Acta Palaeontologica Polonica 2017; 62: 845 - 63. hups: // dx. doi. org / 10.4202 / app. 00373.2017" type="journal article" year="2017">
|
||
Vera
|
||
<emphasis id="B96C184DE920476B9E9FFF5BFF70FEC1" box="[191,238,238,262]" italics="true" pageId="15" pageNumber="432">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
) of this
|
||
<typeStatus id="54A37AFDE920476B9F5BFF5AFE49FEC1" box="[379,471,238,262]" pageId="15" pageNumber="432" type="holotype">holotype</typeStatus>
|
||
is broken at the base, it does show a sub-circular outline. However, the base of right dP1, albeit very badly preserved, appears sub-elliptical. There is possibly an internal asymmetry regarding the morphology of the alveolus of dP1, but the only completely preserved base of dP
|
||
<quantity id="4CE069BAE920476B9CD1FEDFFF1BFE64" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="432" unit="in" value="1.0">1 in</quantity>
|
||
this specimen is sub-circular. We consider this assessment indeterminate and that the morphology of this tooth is not diagnostic for
|
||
<taxonomicName id="4C18BFDCE920476B9EC0FE7EFEC6FE26" baseAuthorityName="Roth" baseAuthorityYear="1898" box="[224,344,457,481]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="endiadys">
|
||
<emphasis id="B96C184DE920476B9EC0FE7EFEC6FE26" box="[224,344,457,481]" italics="true" pageId="15" pageNumber="432">Pr. endiadys</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF89B9AEE920476B9F47FE7EFDBAFE26" author="Vera B & Reguero MA & Gonzalez-Ruiz L" box="[359,548,457,481]" pageId="15" pageNumber="432" pagination="845 - 63" refId="ref25562" refString="Vera B, Reguero MA, Gonzalez-Ruiz L. The Interatheriinae notoungulates from the Middle Miocene Collon Cura Formation in Argentina. Acta Palaeontologica Polonica 2017; 62: 845 - 63. hups: // dx. doi. org / 10.4202 / app. 00373.2017" type="journal article" year="2017">
|
||
Vera
|
||
<emphasis id="B96C184DE920476B9FBFFE7EFE4CFE26" box="[415,466,457,481]" italics="true" pageId="15" pageNumber="432">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
also mentioned the reduction of dP1 and dp1 as diagnostic features of the species, but according to the measurements provided by these authors, these teeth do not differ from those observed in the other species of
|
||
<taxonomicName id="4C18BFDCE920476B9E9AFDF2FECAFD99" authorityName="Ameghino" authorityYear="1885" box="[186,340,582,606]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96C184DE920476B9E9AFDF2FECAFD99" box="[186,340,582,606]" italics="true" pageId="15" pageNumber="432">Protypotherium</emphasis>
|
||
</taxonomicName>
|
||
, considered here as valid, particularly
|
||
<taxonomicName id="4C18BFDCE920476B9CFFFDF3FF23FDB9" baseAuthorityName="Ameghino" baseAuthorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="australe">
|
||
<emphasis id="B96C184DE920476B9CFFFDF3FF23FDB9" italics="true" pageId="15" pageNumber="432">Pr. australe</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18BFDCE920476B9ED0FDD3FE1BFDB9" authorityName="Ameghino" authorityYear="1887" box="[240,389,614,638]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE920476B9ED0FDD3FE1BFDB9" box="[240,389,614,638]" italics="true" pageId="15" pageNumber="432">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
, which have the largest samples (see Supporting Information, Table S2). Therefore, we consider this character indeterminate too. As mentioned above, the M3 of
|
||
<taxonomicName id="4C18BFDCE920476B9CFFFD11FF7BFD1B" authorityName="Ameghino" authorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE920476B9CFFFD11FF7BFD1B" italics="true" pageId="15" pageNumber="432">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
presents a distally expanded metaloph whose length varies from poorly to highly developed, which is one of the characteristics mentioned by
|
||
<bibRefCitation id="EF89B9AEE920476B9F5CFCB7FDA6FCDC" author="Vera B & Reguero MA & Gonzalez-Ruiz L" box="[380,568,770,795]" pageId="15" pageNumber="432" pagination="845 - 63" refId="ref25562" refString="Vera B, Reguero MA, Gonzalez-Ruiz L. The Interatheriinae notoungulates from the Middle Miocene Collon Cura Formation in Argentina. Acta Palaeontologica Polonica 2017; 62: 845 - 63. hups: // dx. doi. org / 10.4202 / app. 00373.2017" type="journal article" year="2017">
|
||
Vera
|
||
<emphasis id="B96C184DE920476B9F93FCB7FE7BFCDD" box="[435,485,770,794]" italics="true" pageId="15" pageNumber="432">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
to re-diagnose
|
||
<taxonomicName id="4C18BFDCE920476B9CFFFCB7FF5BFCFD" baseAuthorityName="Roth" baseAuthorityYear="1898" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="endiadys">
|
||
<emphasis id="B96C184DE920476B9CFFFCB7FF5BFCFD" italics="true" pageId="15" pageNumber="432">Pr. endiadys</emphasis>
|
||
</taxonomicName>
|
||
; however, due to its variability, we do not support this character as diagnostic. Therefore, in the absence of any other characteristics that differentiate
|
||
<taxonomicName id="4C18BFDCE920476B9F94FCD5FDB9FCBF" baseAuthorityName="Roth" baseAuthorityYear="1898" box="[436,551,864,888]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="endiadys">
|
||
<emphasis id="B96C184DE920476B9F94FCD5FDB9FCBF" box="[436,551,864,888]" italics="true" pageId="15" pageNumber="432">Pr. endiadys</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C18BFDCE920476B9C42FCD5FD6BFCBF" authorityName="Ameghino" authorityYear="1887" box="[610,757,864,888]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE920476B9C42FCD5FD6BFCBF" box="[610,757,864,888]" italics="true" pageId="15" pageNumber="432">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
, the former is proposed here as a junior synonym of the laưer.
|
||
</paragraph>
|
||
<paragraph id="8BA7C45FE920476B9EADFC2BFE6CFA4A" blockId="15.[113,765,144,1421]" pageId="15" pageNumber="432">
|
||
Finally,
|
||
<taxonomicName id="4C18BFDCE920476B9EC3FC2BFE4DFC70" authorityName="Ameghino" authorityYear="1891" baseAuthorityName="Roth" baseAuthorityYear="1898" box="[227,467,927,951]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="diastematus">
|
||
<emphasis id="B96C184DE920476B9EC3FC2BFE4DFC70" box="[227,467,927,951]" italics="true" pageId="15" pageNumber="432">Patriarchus diastematus</emphasis>
|
||
</taxonomicName>
|
||
was treated as
|
||
<taxonomicName id="4C18BFDCE920476B9CA7FC2BFD64FC70" authorityName="Ziflel" authorityYear="1893" box="[647,762,927,951]" class="Mammalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="421" phylum="Chordata" rank="subOrder" subOrder="Typotheria">Typotheria</taxonomicName>
|
||
<emphasis id="B96C184DE920476B9E51FC0BFF6FFC11" box="[113,241,958,982]" italics="true" pageId="15" pageNumber="432">incertae sedis</emphasis>
|
||
by Sinclair (1909). Originally, it was differentiated from the remaining species of
|
||
<taxonomicName id="4C18BFDCE920476B9F90FC6AFDBEFC31" authorityName="Ameghino" authorityYear="1889" baseAuthorityName="Roth" baseAuthorityYear="1898" box="[432,544,990,1014]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96C184DE920476B9F90FC6AFDBEFC31" box="[432,544,990,1014]" italics="true" pageId="15" pageNumber="432">Patriarchus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a sub-cylindrical first upper premolar and a diastema between this tooth and P2 (
|
||
<bibRefCitation id="EF89B9AEE920476B9F2DFBA8FE5EFBF3" author="Ameghino F" box="[269,448,1052,1076]" pageId="15" pageNumber="432" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino 1891b</bibRefCitation>
|
||
). However, the maxilla of the
|
||
<typeStatus id="54A37AFDE920476B9E51FB88FF53FB93" box="[113,205,1084,1108]" pageId="15" pageNumber="432" type="holotype">holotype</typeStatus>
|
||
(MACN-A 4044;
|
||
<figureCitation id="1323D8DAE920476B9FB0FB88FE45FB93" box="[400,475,1084,1108]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)–p3–m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)–m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1–dp1–p2–p4, and right i1 and dp1–p2–m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3–M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)–m3; (G) MACN-A 3867, right mandibular fragment with dp1–2–p3–4 (trigonid); (H) MACN-A 9716–9732?g, right mandibular fragment with m1 (talonid)–3; (I) MPM-PV 19761, right maxillary fragment with M1–2; (J) MPM-PV 19781, right mandibular fragment with p2–m1; (K) MPM-PV 19782, less maxillary fragment with P2–M3. Abbreviation: ’, indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8A</figureCitation>
|
||
; see
|
||
<bibRefCitation id="EF89B9AEE920476B9C31FB88FD71FB93" author="Fernandez M & Fernicola JC & Cerdeno E" box="[529,751,1084,1108]" pageId="15" pageNumber="432" pagination="473 - 98" refId="ref23394" refString="Fernandez M, Fernicola JC, Cerdeno E. Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino. Zootaxa 2018; 4387: 473 - 98. hups: // doi. org / 10.11646 / ZOOTAXA. 4387.3.4" type="journal article" year="2018">
|
||
Fernández
|
||
<emphasis id="B96C184DE920476B9CA4FB88FD2BFB93" box="[644,693,1084,1108]" italics="true" pageId="15" pageNumber="432">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
) shows a sub-elliptic alveolus of dP1, as do the majority of the alveoli of dP1 of
|
||
<taxonomicName id="4C18BFDCE920476B9F23FBCEFE03FB55" authorityName="Ameghino" authorityYear="1885" box="[259,413,1146,1170]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96C184DE920476B9F23FBCEFE03FB55" box="[259,413,1146,1170]" italics="true" pageId="15" pageNumber="432">Protypotherium</emphasis>
|
||
</taxonomicName>
|
||
(but they can also be sub-circular). Additionally, the ‘diastema’ between dP1 and P2 is in fact a small space because dP1 is slightly obliquely implanted and lays on P2, as observed in almost all specimens assigned to
|
||
<taxonomicName id="4C18BFDCE920476B9C7BFB6CFD6BFB37" authorityName="Ameghino" authorityYear="1885" box="[603,757,1240,1264]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96C184DE920476B9C7BFB6CFD6BFB37" box="[603,757,1240,1264]" italics="true" pageId="15" pageNumber="432">Protypotherium</emphasis>
|
||
</taxonomicName>
|
||
, including the
|
||
<typeStatus id="54A37AFDE920476B9F20FB4CFEC2FAD7" box="[256,348,1272,1296]" pageId="15" pageNumber="432" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C18BFDCE920476B9F5EFB4CFD8AFAD7" authorityName="Ameghino" authorityYear="1887" box="[382,532,1272,1296]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE920476B9F5EFB4CFD8AFAD7" box="[382,532,1272,1296]" italics="true" pageId="15" pageNumber="432">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
(MACN-A 1081;
|
||
<figureCitation id="1323D8DAE920476B9CF4FB4CFF0CFAE8" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="15" pageNumber="432">Fig. 4D</figureCitation>
|
||
; see
|
||
<bibRefCitation id="EF89B9AEE920476B9EE9FAA3FE36FAE8" author="Fernandez M & Fernicola JC & Cerdeno E" box="[201,424,1303,1327]" pageId="15" pageNumber="432" pagination="473 - 98" refId="ref23394" refString="Fernandez M, Fernicola JC, Cerdeno E. Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino. Zootaxa 2018; 4387: 473 - 98. hups: // doi. org / 10.11646 / ZOOTAXA. 4387.3.4" type="journal article" year="2018">
|
||
Fernández
|
||
<emphasis id="B96C184DE920476B9F1CFAACFEF3FAE8" box="[316,365,1303,1327]" italics="true" pageId="15" pageNumber="432">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). Therefore, both characteristics are discarded as diagnostic; as there is no other feature that can contrast
|
||
<taxonomicName id="4C18BFDCE920476B9EEBFAE2FEFAFAA9" authorityName="Ameghino" authorityYear="1891" box="[203,356,1366,1390]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="diastematus">
|
||
<emphasis id="B96C184DE920476B9EEBFAE2FEFAFAA9" box="[203,356,1366,1390]" italics="true" pageId="15" pageNumber="432">Pa. diastematus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C18BFDCE920476B9F82FAE2FDA9FAA9" authorityName="Ameghino" authorityYear="1887" box="[418,567,1366,1390]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="praerutilum">
|
||
<emphasis id="B96C184DE920476B9F82FAE2FDA9FAA9" box="[418,567,1366,1390]" italics="true" pageId="15" pageNumber="432">Pr. praerutilum</emphasis>
|
||
</taxonomicName>
|
||
, the former is considered a junior synonym of the laưer.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |