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<document id="E6E0B147DD4B06AE11BC7CBF433A3078" ID-CLB-Dataset="295393" ID-DOI="10.5281/zenodo.11105986" ID-GBIF-Dataset="4c7b4d3d-1fbe-4f48-8cb2-002e928eae42" ID-ISSN="0068-547X" ID-Zenodo-Dep="11105986" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1714051164034" checkinUser="felipe" docAuthor="Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Castroviejo-Fisher, Santiago &amp; De, Ignacio" docDate="2020" docId="03B1733AFFE0A144FFFBFE84EED9FAC0" docLanguage="en" docName="ProcCalifAcadSci.66.8.137-230.pdf" docOrigin="Proceedings of the California Academy of Sciences 66 (8)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.1:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="1" docTitle="Phlyctimantis leonardi Photo" docType="treatment" docVersion="4" lastPageNumber="164" masterDocId="FF880B42FFFBA15FFF80FF85EE63FFB7" masterDocTitle="Synopsis of the Amphibians of Equatorial Guinea based upon the Authors Field Work and Spanish Natural History Collections" masterLastPageNumber="230" masterPageNumber="137" pageNumber="164" updateTime="1714671758489" updateUser="ExternalLinkService">
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<mods:title id="0DF30D5CC6BCD3D92CFD43B01D78C754">Synopsis of the Amphibians of Equatorial Guinea based upon the Authors Field Work and Spanish Natural History Collections</mods:title>
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<mods:namePart id="5B6CB8867C1E1E6E2F35E810E87FFC64">Sánchez-Vialas, Alberto</mods:namePart>
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<mods:namePart id="D287B54AC123AB8DA83F50412FA9ED73">Calvo-Revuelta, Marta</mods:namePart>
<mods:affiliation id="A0A6B1EA6D56ADE41F26637A9B0A0E10">Museo Nacional de Ciencias Naturales (MNCN-CSIC), C / José Gutiérrez Abascal, 2. 28006, Madrid, Spain;</mods:affiliation>
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<mods:namePart id="C944F5E868FE601F888294E049BA0433">Castroviejo-Fisher, Santiago</mods:namePart>
<mods:affiliation id="FC42AB3732C5BBFF2A013E39B167F609">Laboratorio de Sistemática de Vertebrados, Pontifícia Universidade Católica do Rio Grande do Sul (PUCRS). Av / Ipiranga, 6681, Prédio 40, sala 110, 90619 - 900, Porto Alegre, Brazil; * Corresponding author: IDlR (iriva @ mncn. csic. es)</mods:affiliation>
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<mods:namePart id="2FD9229B85C61C447E0EF7027190D702">De, Ignacio</mods:namePart>
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<treatment id="03B1733AFFE0A144FFFBFE84EED9FAC0" ID-DOI="http://doi.org/10.5281/zenodo.11106219" ID-Zenodo-Dep="11106219" LSID="urn:lsid:plazi:treatment:03B1733AFFE0A144FFFBFE84EED9FAC0" httpUri="http://treatment.plazi.org/id/03B1733AFFE0A144FFFBFE84EED9FAC0" lastPageNumber="164" pageId="27" pageNumber="164">
<subSubSection id="C30291A7FFE0A144FFFBFE84EAECFEAE" box="[123,1167,257,281]" pageId="27" pageNumber="164" type="nomenclature">
<paragraph id="8BA7C22CFFE0A144FFFBFE84EAECFEAE" blockId="27.[123,1174,218,1030]" box="[123,1167,257,281]" pageId="27" pageNumber="164">
<heading id="D0EF7540FFE0A144FFFBFE84EAECFEAE" box="[123,1167,257,281]" centered="true" fontSize="10" level="2" pageId="27" pageNumber="164" reason="4">
<emphasis id="B96C1E3EFFE0A144FFFBFE84EAECFEAE" bold="true" box="[123,1167,257,281]" pageId="27" pageNumber="164">
<taxonomicName id="4C18B9AFFFE0A144FFFBFE84ED86FEAE" ID-CoL="4GJPF" authority="(Boulenger, 1906) Photo" authorityName="Photo" baseAuthorityName="Boulenger" baseAuthorityYear="1906" box="[123,997,257,281]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" isUncertain="true" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="B96C1E3EFFE0A144FFFBFE84EF71FEAE" bold="true" box="[123,274,257,281]" italics="true" pageId="27" pageNumber="164">Phlyctimantis</emphasis>
cf.
<emphasis id="B96C1E3EFFE0A144FEBCFE84EFFBFEAE" bold="true" box="[316,408,257,281]" italics="true" pageId="27" pageNumber="164">leonardi</emphasis>
(
<bibRefCitation id="EF89BFDDFFE0A144FE28FE84EC3EFEAE" author="BOULENGER, G. A." box="[424,605,257,281]" pageId="27" pageNumber="164" pagination="157 - 172" refId="ref29449" refString="BOULENGER, G. A. 1906 a. Report on the reptiles collected by the late L. Fea in West Africa. Annali del Museo Civico di Storia Naturale di Genova 3 (2): 157 - 172." type="journal article" year="1906">Boulenger, 1906</bibRefCitation>
) Photo
</taxonomicName>
figures 18AB
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30291A7FFE0A144FF2BFEA4EFCFFECD" pageId="27" pageNumber="164" type="materials_examined">
<paragraph id="8BA7C22CFFE0A144FF2BFEA4EFCFFECD" blockId="27.[123,1174,218,1030]" pageId="27" pageNumber="164">
<materialsCitation id="3B70C871FFE0A144FF2BFEA4ED8EFE8D" box="[171,1005,288,314]" collectionCode="TYPE, LOCALITY" country="Equatorial Guinea" location="Bioko" pageId="27" pageNumber="164" specimenCount="1">
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<collectionCode id="ED095AE9FFE0A144FF2BFEA4EE82FE8E" box="[171,225,289,313]" pageId="27" pageNumber="164">TYPE</collectionCode>
</smallCapsWord>
<smallCapsWord id="8D4154F0FFE0A144FF6DFEA1EF2DFE8F" baselines="307" box="[237,334,292,312]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="27" pageNumber="164">
<collectionCode id="ED095AE9FFE0A144FF6DFEA1EF2DFE8F" box="[237,334,292,312]" pageId="27" pageNumber="164">LOCALITY</collectionCode>
</smallCapsWord>
.— “Punta Frailes, Fernando Po”,
<location id="8EC794F7FFE0A144FD51FEA5ED71FE8D" LSID="urn:lsid:plazi:treatment:03B1733AFFE0A144FFFBFE84EED9FAC0:8EC794F7FFE0A144FD51FEA5ED71FE8D" box="[721,786,288,314]" country="Equatorial Guinea" name="Bioko" pageId="27" pageNumber="164">Bioko</location>
,
<collectingCountry id="F30F82BCFFE0A144FCA5FEA5ED8EFE8D" box="[805,1005,288,314]" name="Equatorial Guinea" pageId="27" pageNumber="164">Equatorial Guinea</collectingCountry>
</materialsCitation>
and “NDjolè, French Congo” Ndjolé,
<collectingCountry id="F30F82BCFFE0A144FEFFFEC5EFA5FEED" box="[383,454,320,346]" name="Gabon" pageId="27" pageNumber="164">Gabon</collectingCountry>
.
<materialsCitation id="3B70C871FFE0A144FE54FEC4EFC6FECC" collectionCode="LOCALITY" country="Gabon" location="Gabon" pageId="27" pageNumber="164" specimenCount="1" typeStatus="lectotype">
<bibRefCitation id="EF89BFDDFFE0A144FE54FEC4ECFCFEED" author="CAPOCACCIA, L." box="[468,671,320,346]" pageId="27" pageNumber="164" pagination="208 - 222" refId="ref29667" refString="CAPOCACCIA, L. 1957. Catalogo dei tipi di anfibi del Museo Civico di Storia Naturale di Genova. Annali del Museo Civico di Storia Naturale di Genova 69 (3): 208 - 222." type="journal article" year="1957">Capocaccia (1957)</bibRefCitation>
restricted the type locality by
<typeStatus id="54A37C8EFFE0A144FC6AFEC5EA2FFEED" box="[1002,1100,320,346]" pageId="27" pageNumber="164" type="lectotype">lectotype</typeStatus>
designation to “NDjolè”,
<collectingCountry id="F30F82BCFFE0A144FEDEFEE4EFC6FECC" box="[350,421,353,379]" name="Gabon" pageId="27" pageNumber="164">Gabon</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30291A7FFE0A144FF2BFE06EFAAFE4B" pageId="27" pageNumber="164" type="distribution">
<paragraph id="8BA7C22CFFE0A144FF2BFE06EFAAFE4B" blockId="27.[123,1174,218,1030]" pageId="27" pageNumber="164">
<smallCapsWord id="8D4154F0FFE0A144FF2BFE06EF5FFE2D" baselines="405,405" box="[171,316,387,411]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Distribution" pageId="27" pageNumber="164">DISTRIBUTION</smallCapsWord>
.— This species is distributed from
<collectingCountry id="F30F82BCFFE0A144FD38FE07ED44FE2C" box="[696,807,386,411]" name="Cameroon" pageId="27" pageNumber="164">Cameroon</collectingCountry>
to
<collectingCountry id="F30F82BCFFE0A144FCC9FE04EDF3FE2C" box="[841,912,385,411]" name="Gabon" pageId="27" pageNumber="164">Gabon</collectingCountry>
,
<collectingCountry id="F30F82BCFFE0A144FC1EFE04EA0BFE2C" box="[926,1128,385,411]" name="Republic of the Congo" pageId="27" pageNumber="164">Republic of Congo</collectingCountry>
and
<collectingCountry id="F30F82BCFFE0A144FFFBFE26EFABFE0B" box="[123,456,418,444]" name="Democratic Republic of the Congo" pageId="27" pageNumber="164">Democratic Republic of Congo</collectingCountry>
. In
<collectingCountry id="F30F82BCFFE0A144FE71FE27ECD7FE0B" box="[497,692,418,444]" name="Equatorial Guinea" pageId="27" pageNumber="164">Equatorial Guinea</collectingCountry>
, it has been cited in
<collectingCountry id="F30F82BCFFE0A144FC0EFE1AED96FE0C" box="[910,1013,415,444]" name="Equatorial Guinea" pageId="27" pageNumber="164">Río Muni</collectingCountry>
, over the coast of Bome, and Monte Alén (
<bibRefCitation id="EF89BFDDFFE0A144FE31FE41EC0AFE6B" author="DE LA RIVA" box="[433,617,450,476]" pageId="27" pageNumber="164" pagination="123 - 139" refId="ref29939" refString="DE LA RIVA, I. 1994. Anfibios anuros del Parque Nacional de Monte Alen, Rio Muni, Guinea Ecuatorial. Revista Espanola de Herpetologia 8: 123 - 139." type="journal article" year="1994">De la Riva 1994</bibRefCitation>
), and on Bioko at Punta Europa (= Punta Frailes) (
<bibRefCitation id="EF89BFDDFFE0A144FF04FE66EF58FE4B" author="BOULENGER, G. A." box="[132,315,483,509]" pageId="27" pageNumber="164" pagination="157 - 172" refId="ref29449" refString="BOULENGER, G. A. 1906 a. Report on the reptiles collected by the late L. Fea in West Africa. Annali del Museo Civico di Storia Naturale di Genova 3 (2): 157 - 172." type="journal article" year="1906">Boulenger 1906a</bibRefCitation>
) (
<figureCitation id="1323DEA9FFE0A144FED4FE61EFDAFE4B" box="[340,441,484,509]" captionStart="MAPS" captionStartId="84.[155,174,1667,1687]" captionTargetBox="[170,1125,242,1634]" captionTargetId="figure-51@84.[139,1156,216,1654]" captionTargetPageId="84" captionText="MAPS 19AC. Distribution maps for Equatorial Guinean records of (A) Opisthothylax immaculatus; (B) Phlyctimantis cf. leonardi; (C) Petropedetes cameronensis." figureDoi="http://doi.org/10.5281/zenodo.11106103" httpUri="https://zenodo.org/record/11106103/files/figure.png" pageId="27" pageNumber="164">Map 19B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30291A7FFE0A144FF2BFD81ED4AFBB2" pageId="27" pageNumber="164" type="discussion">
<paragraph id="8BA7C22CFFE0A144FF2BFD81EDFDFC53" blockId="27.[123,1174,218,1030]" pageId="27" pageNumber="164">
<smallCapsWord id="8D4154F0FFE0A144FF2BFD81EF43FDAB" baselines="535,535" box="[171,288,516,541]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Comments" pageId="27" pageNumber="164">COMMENTS</smallCapsWord>
.— There is a second described species of
<taxonomicName id="4C18B9AFFFE0A144FC80FD83EDF0FDA9" authorityName="Laurent and Combaz" authorityYear="1950" box="[768,915,518,542]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="genus">
<emphasis id="B96C1E3EFFE0A144FC80FD83EDF0FDA9" box="[768,915,518,542]" italics="true" pageId="27" pageNumber="164">Phlyctimantis</emphasis>
</taxonomicName>
inhabiting west-central Africa, namely
<taxonomicName id="4C18B9AFFFE0A144FEB0FDA3ECB8FD89" authority="Perret, 1986" authorityName="Perret" authorityYear="1986" box="[304,731,549,574]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B96C1E3EFFE0A144FEB0FDA3EC27FD89" box="[304,580,550,574]" italics="true" pageId="27" pageNumber="164">Phlyctimantis boulengeri</emphasis>
Perret, 1986
</taxonomicName>
(
<typeStatus id="54A37C8EFFE0A144FD72FDA0ED44FD8A" box="[754,807,549,573]" pageId="27" pageNumber="164">Type</typeStatus>
locality: “Fainchang, région de Mamfe,
<collectingCountry id="F30F82BCFFE0A144FF55FDC0EF27FDE9" box="[213,324,581,606]" name="Cameroon" pageId="27" pageNumber="164">Cameroon</collectingCountry>
occidental”), considered as a subspecies of
<taxonomicName id="4C18B9AFFFE0A144FC9AFDCDEDEEFDE8" authorityName="Photo" baseAuthorityName="Boulenger" baseAuthorityYear="1906" box="[794,909,583,607]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="B96C1E3EFFE0A144FC9AFDCDEDEEFDE8" box="[794,909,583,607]" italics="true" pageId="27" pageNumber="164">P. leonardi</emphasis>
</taxonomicName>
by some authors (
<bibRefCitation id="EF89BFDDFFE0A144FBD3FDC3EED3FDC8" author="AMIET, J. L." pageId="27" pageNumber="164" refId="ref28302" refString="AMIET, J. L. 2012. Les Rainettes du Cameroun (Amphibiens Anoures). Saint-Nazaire, France, La Nef des Livres. 591 pp." type="book" year="2012">Amiet 2012</bibRefCitation>
; Amiet and Goutte 2017). The identity of several populations from west and central-western Africa remains problematic (
<bibRefCitation id="EF89BFDDFFE0A144FE2EFD02EC4FFD28" author="AMIET, J. L." box="[430,556,646,671]" pageId="27" pageNumber="164" pagination="87 - 126" refId="ref28270" refString="AMIET, J. L. 2007. Les Phlyctimantis et Kassina du Cameroun (Amphibia, Anura, Hyperoliidae). Revue Suisse de Zoologie 114: 87 - 126." type="journal article" year="2007">Amiet 2007</bibRefCitation>
;
<bibRefCitation id="EF89BFDDFFE0A144FDBAFD00EC9AFD28" author="KOHLER, J. &amp; K. SCHEELKE &amp; S. SCHICK &amp; M. VEITH &amp; S. LOTTERS" box="[570,761,645,672]" pageId="27" pageNumber="164" pagination="127 - 142" refId="ref31252" refString="KOHLER, J., K. SCHEELKE, S. SCHICK, M. VEITH, AND S. LOTTERS. 2005. Contribution to the taxonomy of hyperoliid frogs (Amphibia: Anura: Hyperoliidae): advertisement calls of twelve species from East and Central Africa. African Zoology, 40 (1): 127 - 142." type="journal article" year="2005">Köhler et al. 2005</bibRefCitation>
). Although there are records identified as
<taxonomicName id="4C18B9AFFFE0A144FF1AFD2CEF6CFD76" authorityName="Photo" baseAuthorityName="Boulenger" baseAuthorityYear="1906" box="[154,271,681,705]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="B96C1E3EFFE0A144FF1AFD2CEF6CFD76" box="[154,271,681,705]" italics="true" pageId="27" pageNumber="164">P. leonardi</emphasis>
</taxonomicName>
from Bioko and
<collectingCountry id="F30F82BCFFE0A144FE4EFD26EC54FD08" box="[462,567,675,704]" name="Equatorial Guinea" pageId="27" pageNumber="164">Río Muni</collectingCountry>
, the taxonomic allocation of both mainland and insular populations needs to be revised.
<bibRefCitation id="EF89BFDDFFE0A144FE6FFD43ECACFD56" author="KOHLER, J. &amp; K. SCHEELKE &amp; S. SCHICK &amp; M. VEITH &amp; S. LOTTERS" box="[495,719,710,737]" pageId="27" pageNumber="164" pagination="127 - 142" refId="ref31252" refString="KOHLER, J., K. SCHEELKE, S. SCHICK, M. VEITH, AND S. LOTTERS. 2005. Contribution to the taxonomy of hyperoliid frogs (Amphibia: Anura: Hyperoliidae): advertisement calls of twelve species from East and Central Africa. African Zoology, 40 (1): 127 - 142." type="journal article" year="2005">Köhler et al. (2005)</bibRefCitation>
suggested that several populations from
<collectingCountry id="F30F82BCFFE0A144FFFBFD6DEE89FCB6" box="[123,234,744,769]" name="Cameroon" pageId="27" pageNumber="164">Cameroon</collectingCountry>
,
<collectingCountry id="F30F82BCFFE0A144FF7CFD62EF20FCB6" box="[252,323,743,769]" name="Gabon" pageId="27" pageNumber="164">Gabon</collectingCountry>
,
<collectingCountry id="F30F82BCFFE0A144FED5FD62EC7EFCB6" box="[341,541,743,769]" name="Equatorial Guinea" pageId="27" pageNumber="164">Equatorial Guinea</collectingCountry>
and
<collectingCountry id="F30F82BCFFE0A144FDDAFD6CEDD4FCB6" box="[602,951,743,769]" name="Democratic Republic of the Congo" pageId="27" pageNumber="164">Democratic Republic of Congo</collectingCountry>
probably belong to
<taxonomicName id="4C18B9AFFFE0A144FFFBFC8EEF69FC95" authorityName="Perret" authorityYear="1986" box="[123,266,778,802]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B96C1E3EFFE0A144FFFBFC8EEF69FC95" box="[123,266,778,802]" italics="true" pageId="27" pageNumber="164">P. boulengeri</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF89BFDDFFE0A144FE99FC8DEF8BFC95" author="BOSCH, J. &amp; I. DE LA RIVA &amp; R. MARQUEZ" box="[281,488,776,802]" pageId="27" pageNumber="164" pagination="246 - 255" refId="ref29223" refString="BOSCH, J., I. DE LA RIVA, AND R. MARQUEZ. 2000. Advertisement calls of seven species of hyperoliid frogs, from Equatorial Guinea. Amphibia-Reptilia 21 (2): 246 - 255." type="journal article" year="2000">Bosch et al. (2000)</bibRefCitation>
provided a call spectrogram of putative
<taxonomicName id="4C18B9AFFFE0A144FC21FC8FEAF6FC95" authorityName="Photo" baseAuthorityName="Boulenger" baseAuthorityYear="1906" box="[929,1173,778,802]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="B96C1E3EFFE0A144FC21FC8FEAF6FC95" box="[929,1173,778,802]" italics="true" pageId="27" pageNumber="164">Phlyctimantis leonardi</emphasis>
</taxonomicName>
from Bome,
<collectingCountry id="F30F82BCFFE0A144FE83FCA0EF08FCF5" box="[259,363,805,834]" name="Equatorial Guinea" pageId="27" pageNumber="164">Río Muni</collectingCountry>
, which are highly similar to those of
<taxonomicName id="4C18B9AFFFE0A144FD7DFCA9EDEFFCF4" authorityName="Perret" authorityYear="1986" box="[765,908,811,835]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B96C1E3EFFE0A144FD7DFCA9EDEFFCF4" box="[765,908,811,835]" italics="true" pageId="27" pageNumber="164">P. boulengeri</emphasis>
</taxonomicName>
from Barrage de Tchimbélé,
<collectingCountry id="F30F82BCFFE0A144FF35FCCCEE9FFCD4" box="[181,252,841,867]" name="Gabon" pageId="27" pageNumber="164">Gabon</collectingCountry>
(
<bibRefCitation id="EF89BFDDFFE0A144FE8BFCCDEFAFFCD4" author="KOHLER, J. &amp; K. SCHEELKE &amp; S. SCHICK &amp; M. VEITH &amp; S. LOTTERS" box="[267,460,840,867]" pageId="27" pageNumber="164" pagination="127 - 142" refId="ref31252" refString="KOHLER, J., K. SCHEELKE, S. SCHICK, M. VEITH, AND S. LOTTERS. 2005. Contribution to the taxonomy of hyperoliid frogs (Amphibia: Anura: Hyperoliidae): advertisement calls of twelve species from East and Central Africa. African Zoology, 40 (1): 127 - 142." type="journal article" year="2005">Köhler et al. 2005</bibRefCitation>
). However, both species names (
<taxonomicName id="4C18B9AFFFE0A144FCAFFCC9EDDDFCD4" authorityName="Perret" authorityYear="1986" box="[815,958,843,867]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B96C1E3EFFE0A144FCAFFCC9EDDDFCD4" box="[815,958,843,867]" italics="true" pageId="27" pageNumber="164">P. boulengeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18B9AFFFE0A144FC71FCC9EA06FCD4" authorityName="Photo" baseAuthorityName="Boulenger" baseAuthorityYear="1906" box="[1009,1125,843,867]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="B96C1E3EFFE0A144FC71FCC9EA06FCD4" box="[1009,1125,843,867]" italics="true" pageId="27" pageNumber="164">P. leonardi</emphasis>
</taxonomicName>
) are likely representing still more undescribed species (
<bibRefCitation id="EF89BFDDFFE0A144FD23FCEDED0BFC34" author="KOHLER, J. &amp; K. SCHEELKE &amp; S. SCHICK &amp; M. VEITH &amp; S. LOTTERS" box="[675,872,872,899]" pageId="27" pageNumber="164" pagination="127 - 142" refId="ref31252" refString="KOHLER, J., K. SCHEELKE, S. SCHICK, M. VEITH, AND S. LOTTERS. 2005. Contribution to the taxonomy of hyperoliid frogs (Amphibia: Anura: Hyperoliidae): advertisement calls of twelve species from East and Central Africa. African Zoology, 40 (1): 127 - 142." type="journal article" year="2005">Köhler et al. 2005</bibRefCitation>
;
<bibRefCitation id="EF89BFDDFFE0A144FCF6FCEEED95FC34" author="AMIET, J. L." box="[886,1014,874,899]" pageId="27" pageNumber="164" pagination="87 - 126" refId="ref28270" refString="AMIET, J. L. 2007. Les Phlyctimantis et Kassina du Cameroun (Amphibia, Anura, Hyperoliidae). Revue Suisse de Zoologie 114: 87 - 126." type="journal article" year="2007">Amiet 2007</bibRefCitation>
; Onadeko and Rödel 2009). The specimen in the EBD collection came from Bata region,
<collectingCountry id="F30F82BCFFE0A144FC6CFC02EA38FC14" box="[1004,1115,903,932]" name="Equatorial Guinea" pageId="27" pageNumber="164">Río Muni</collectingCountry>
, and morphologically resembles
<taxonomicName id="4C18B9AFFFE0A144FE24FC2BEC7BFC72" authorityName="Photo" baseAuthorityName="Boulenger" baseAuthorityYear="1906" box="[420,536,941,965]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="B96C1E3EFFE0A144FE24FC2BEC7BFC72" box="[420,536,941,965]" italics="true" pageId="27" pageNumber="164">P. leonardi</emphasis>
</taxonomicName>
. Thus, we tentatively keep the populations from
<collectingCountry id="F30F82BCFFE0A144FBA6FC2EEEABFC52" name="Equatorial Guinea" pageId="27" pageNumber="164">Equatorial Guinea</collectingCountry>
as
<taxonomicName id="4C18B9AFFFE0A144FF6DFC4BEFE4FC52" authorityName="Photo" baseAuthorityName="Boulenger" baseAuthorityYear="1906" box="[237,391,971,997]" class="Amphibia" family="Hyperoliidae" genus="Phlyctimantis" isUncertain="true" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="B96C1E3EFFE0A144FF6DFC4BEE9EFC52" box="[237,253,974,997]" italics="true" pageId="27" pageNumber="164">P</emphasis>
. cf.
<emphasis id="B96C1E3EFFE0A144FEAEFC48EFE4FC52" box="[302,391,973,997]" italics="true" pageId="27" pageNumber="164">leonardi</emphasis>
</taxonomicName>
until more studies help to clarify these questions.
</paragraph>
<paragraph id="8BA7C22CFFE0A144FF2BFC69ED4AFBB2" blockId="27.[123,1174,218,1030]" box="[171,809,1004,1030]" pageId="27" pageNumber="164">
<smallCapsWord id="8D4154F0FFE0A144FF2BFC69EF7FFBB3" baselines="1023,1023" box="[171,284,1004,1029]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Specimens" pageId="27" pageNumber="164">SPECIMENS</smallCapsWord>
<smallCapsWord id="8D4154F0FFE0A144FEA4FC74EFECFBB2" baselines="1024" box="[292,399,1009,1029]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="examined" pageId="27" pageNumber="164">EXAMINED</smallCapsWord>
.— Asonga, Bata, 1984 (EBD 18254).
</paragraph>
</subSubSection>
<subSubSection id="C30291A7FFE0A144FE6CFBABEC09FB23" pageId="27" pageNumber="164" type="description">
<paragraph id="8BA7C22CFFE0A144FE6CFBABED47FBF1" blockId="27.[123,1174,1070,1400]" box="[492,804,1070,1095]" pageId="27" pageNumber="164">
<taxonomicName id="4C18B9AFFFE0A144FE6CFBABED47FBF1" authority="Noble, 1931" authorityName="Noble" authorityYear="1931" box="[492,804,1070,1095]" class="Amphibia" family="Petropedetidae" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="family">
<emphasis id="B96C1E3EFFE0A144FE6CFBABED47FBF1" bold="true" box="[492,804,1070,1095]" pageId="27" pageNumber="164">Petropedetidae Noble, 1931</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BA7C22CFFE0A144FE60FBD0ED53FBDA" blockId="27.[123,1174,1070,1400]" box="[480,816,1109,1133]" pageId="27" pageNumber="164">
<taxonomicName id="4C18B9AFFFE0A144FE60FBD0ED53FBDA" authority="Reichenow, 1874" authorityName="Reichenow" authorityYear="1874" box="[480,816,1109,1133]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="genus">
<emphasis id="B96C1E3EFFE0A144FE60FBD0ED53FBDA" bold="true" box="[480,816,1109,1133]" pageId="27" pageNumber="164">
<emphasis id="B96C1E3EFFE0A144FE60FBD0EC08FBDA" bold="true" box="[480,619,1109,1133]" italics="true" pageId="27" pageNumber="164">Petropedetes</emphasis>
Reichenow, 1874
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BA7C22CFFE0A144FFFBFBFEEC09FB23" blockId="27.[123,1174,1070,1400]" box="[123,618,1147,1172]" pageId="27" pageNumber="164">
<taxonomicName id="4C18B9AFFFE0A144FFFBFBFEEC09FB23" authority="Reichenow, 1874" authorityName="Reichenow" authorityYear="1874" box="[123,618,1147,1172]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="27" pageNumber="164" phylum="Chordata" rank="species" species="cameronensis">
<emphasis id="B96C1E3EFFE0A144FFFBFBFEEC09FB23" bold="true" box="[123,618,1147,1172]" pageId="27" pageNumber="164">
<emphasis id="B96C1E3EFFE0A144FFFBFBFEEFC7FB23" bold="true" box="[123,420,1147,1172]" italics="true" pageId="27" pageNumber="164">Petropedetes cameronensis</emphasis>
Reichenow, 1874
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30291A7FFE0A144FF2BFB19EE92FB63" pageId="27" pageNumber="164" type="materials_examined">
<paragraph id="8BA7C22CFFE0A144FF2BFB19EE92FB63" blockId="27.[123,1174,1070,1400]" pageId="27" pageNumber="164">
<materialsCitation id="3B70C871FFE0A144FF2BFB19EE89FB62" collectionCode="LOCALITY" country="Cameroon" location="Cameroon" pageId="27" pageNumber="164" specimenCount="1" typeStatus="holotype">
<smallCapsWord id="8D4154F0FFE0A144FF2BFB19EE82FB03" baselines="1198,1199" box="[171,225,1180,1204]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="27" pageNumber="164">
<typeStatus id="54A37C8EFFE0A144FF2BFB19EE82FB03" box="[171,225,1180,1204]" pageId="27" pageNumber="164">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D4154F0FFE0A144FF71FB1AEF31FB04" baselines="1198" box="[241,338,1183,1203]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="27" pageNumber="164">
<collectionCode id="ED095AE9FFE0A144FF71FB1AEF31FB04" box="[241,338,1183,1203]" pageId="27" pageNumber="164">LOCALITY</collectionCode>
</smallCapsWord>
.— “Gebirgsbach bei Bimbia, in den Vorbergen des Camerun”, Limbé,
<collectingCountry id="F30F82BCFFE0A144FFFBFB39EE89FB62" box="[123,234,1212,1237]" name="Cameroon" pageId="27" pageNumber="164">Cameroon</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30291A7FFE0A144FF2BFB58EED9FAC0" pageId="27" pageNumber="164" type="distribution">
<paragraph id="8BA7C22CFFE0A144FF2BFB58EF2AFA81" blockId="27.[123,1174,1070,1400]" pageId="27" pageNumber="164">
<smallCapsWord id="8D4154F0FFE0A144FF2BFB58EF5FFB43" baselines="1263,1263" box="[171,316,1245,1269]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Distribution" pageId="27" pageNumber="164">DISTRIBUTION</smallCapsWord>
.— It ranges from southeastern
<collectingCountry id="F30F82BCFFE0A144FD1EFB58EC8DFB41" box="[670,750,1245,1270]" name="Nigeria" pageId="27" pageNumber="164">Nigeria</collectingCountry>
to southwestern
<collectingCountry id="F30F82BCFFE0A144FC30FB59EA7CFB42" box="[944,1055,1244,1269]" name="Cameroon" pageId="27" pageNumber="164">Cameroon</collectingCountry>
, including Bioko, the only place in
<collectingCountry id="F30F82BCFFE0A144FE15FB79EC38FAA1" box="[405,603,1276,1302]" name="Equatorial Guinea" pageId="27" pageNumber="164">Equatorial Guinea</collectingCountry>
where it has been recorded, at Musola (
<bibRefCitation id="EF89BFDDFFE0A144FBA6FB79EEDFFA81" author="BOULENGER, G. A." pageId="27" pageNumber="164" pagination="157 - 172" refId="ref29449" refString="BOULENGER, G. A. 1906 a. Report on the reptiles collected by the late L. Fea in West Africa. Annali del Museo Civico di Storia Naturale di Genova 3 (2): 157 - 172." type="journal article" year="1906">Boulenger 1906a</bibRefCitation>
) (
<figureCitation id="1323DEA9FFE0A144FF55FA9BEF59FA81" box="[213,314,1309,1334]" captionStart="MAPS" captionStartId="84.[155,174,1667,1687]" captionTargetBox="[170,1125,242,1634]" captionTargetId="figure-51@84.[139,1156,216,1654]" captionTargetPageId="84" captionText="MAPS 19AC. Distribution maps for Equatorial Guinean records of (A) Opisthothylax immaculatus; (B) Phlyctimantis cf. leonardi; (C) Petropedetes cameronensis." figureDoi="http://doi.org/10.5281/zenodo.11106103" httpUri="https://zenodo.org/record/11106103/files/figure.png" pageId="27" pageNumber="164">Map 19C</figureCitation>
).
</paragraph>
<paragraph id="8BA7C22CFFE0A144FF2BFABBEED9FAC0" blockId="27.[123,1174,1070,1400]" pageId="27" pageNumber="164">
<smallCapsWord id="8D4154F0FFE0A144FF2BFABBEF7FFAE1" baselines="1361,1361" box="[171,284,1342,1367]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Specimens" pageId="27" pageNumber="164">SPECIMENS</smallCapsWord>
<smallCapsWord id="8D4154F0FFE0A144FEA5FAC7EFF3FAE1" baselines="1361" box="[293,400,1346,1366]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="examined" pageId="27" pageNumber="164">EXAMINED</smallCapsWord>
.— No specimens were found at the collections examined by us in this study.
</paragraph>
</subSubSection>
</treatment>
</document>