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<document id="F6E403DCE23D4631A07BA48234E06E75" ID-DOI="10.11646/zootaxa.3731.4.4" ID-GBIF-Dataset="26facfec-e615-4814-88c5-c966fcbc9efc" ID-ISSN="1175-5326" ID-Zenodo-Dep="248259" ID-ZooBank="2066C0D1-2ED6-41EB-AF3C-3DAB58EA1718" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460694621800" checkinUser="plazi" docAuthor="Krikken, Jan &amp; Li, Chun-Lin" docDate="2013" docId="03ABBA06FFBDFFDDA181FF20FD0FFD62" docLanguage="en" docName="zt03731p519.pdf" docOrigin="Zootaxa 3731 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bolbochromus (Bolbochromops) walshi Pouillaude 1914" docType="treatment" docVersion="9" lastPageNumber="517" masterDocId="FF92C27EFFABFFCBA116FFB7FFCBFFE1" masterDocTitle="Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae)" masterLastPageNumber="519" masterPageNumber="495" pageNumber="517" updateTime="1698353700889" updateUser="plazi">
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<mods:title id="EEC4A6110894133BC380C63C7008D3CB">Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae)</mods:title>
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<mods:namePart id="FB7B2FFBCF031C3F3FCF13E648A7AB2A">Krikken, Jan</mods:namePart>
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<mods:namePart id="D69E1DD3C18FEFDC6C38BC02E6E6EB8A">Li, Chun-Lin</mods:namePart>
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<mods:date id="5DC68D6D04756A05D0C1B499371A175B">2013</mods:date>
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<mods:number id="05331E062BDA16613B8EF9B55112CD1E">3731</mods:number>
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<treatment id="03ABBA06FFBDFFDDA181FF20FD0FFD62" ID-DOI="http://doi.org/10.5281/zenodo.5680783" ID-GBIF-Taxon="119581767" ID-Zenodo-Dep="5680783" LSID="urn:lsid:plazi:treatment:03ABBA06FFBDFFDDA181FF20FD0FFD62" httpUri="http://treatment.plazi.org/id/03ABBA06FFBDFFDDA181FF20FD0FFD62" lastPageNumber="517" pageId="22" pageNumber="517">
<subSubSection id="C318589BFFBDFFDDA181FF20FE74FDF7" pageId="22" pageNumber="517" type="nomenclature">
<paragraph id="8BBD0B10FFBDFFDDA181FF20FC85FF53" blockId="22.[151,846,151,211]" box="[151,846,151,178]" pageId="22" pageNumber="517">
<heading id="D0F5BC7CFFBDFFDDA181FF20FC85FF53" bold="true" box="[151,846,151,178]" fontSize="11" level="1" pageId="22" pageNumber="517" reason="1">
<taxonomicName id="4C027093FFBDFFDDA181FF20FC85FF53" authority="Pouillaude, 1914" authorityName="Pouillaude" authorityYear="1914" box="[151,846,151,178]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="517" phylum="Arthropoda" rank="species" species="walshi" subGenus="Bolbochromops">
<emphasis id="B976D702FFBDFFDDA181FF20FC85FF53" bold="true" box="[151,846,151,178]" pageId="22" pageNumber="517">
<emphasis id="B976D702FFBDFFDDA181FF20FE82FF50" bold="true" box="[151,329,151,177]" italics="true" pageId="22" pageNumber="517">Bolbochromus</emphasis>
(
<emphasis id="B976D702FFBDFFDDA04FFF20FDDCFF50" bold="true" box="[345,535,151,177]" italics="true" pageId="22" pageNumber="517">Bolbochromops</emphasis>
)
<emphasis id="B976D702FFBDFFDDA33FFF20FDBCFF50" bold="true" box="[553,631,151,177]" italics="true" pageId="22" pageNumber="517">walshi</emphasis>
Pouillaude, 1914
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBD0B10FFBDFFDDA181FF0DFE93FF32" blockId="22.[151,846,151,211]" box="[151,344,186,211]" pageId="22" pageNumber="517">
(
<figureCitation id="13391795FFBDFFDDA189FF0DFEECFF32" box="[159,295,186,211]" captionStart="FIGURES 17 24" captionStartId="10.[151,264,1925,1947]" captionTargetBox="[161,1426,197,1897]" captionTargetId="figure@10.[151,1436,193,1904]" captionTargetPageId="10" captionText="FIGURES 17 24. Habitus of male Bolbochromus (oblique and lateral views). 17 18, B. sulcicollis; 19 20, B. ludekingi; 21 22, B. walshi; 23 24, B. laetus or near. Scale lines 1 mm." httpUri="https://zenodo.org/record/248262/files/figure.png" pageId="22" pageNumber="517">Figs. 2122</figureCitation>
, 43)
</paragraph>
<paragraph id="8BBD0B10FFBDFFDDA181FEB5FE74FDF7" blockId="22.[151,1436,258,643]" pageId="22" pageNumber="517">
<emphasis id="B976D702FFBDFFDDA181FEB5FE8BFEFA" bold="true" box="[151,320,258,283]" pageId="22" pageNumber="517">Identification.</emphasis>
Robust species, with long median horn on both clypeal apex and postfrons; base of clypeal horn broad (approximately 0.3 of clypeal width). Colour pattern basically constant, generally black; pronotal sides usually black, densely, strongly punctate; broad median area of pronotum usually plain (purplish-)red. Pronotum usually with pair of slight but distinct antediscal protrusions. Black colour of pronotum and elytra may be dark brown, and entirely black individuals were seen from east Java. Confusion with other Java species possible, certainly where unicolourous individuals occur (as just mentioned, then confusion with
<taxonomicName id="4C027093FFBDFFDDA58EFE00FB3DFE2F" box="[1176,1270,439,462]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="517" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B976D702FFBDFFDDA58EFE00FB3DFE2F" box="[1176,1270,439,462]" italics="true" pageId="22" pageNumber="517">B. niger</emphasis>
</taxonomicName>
likely, which, aforementioned features aside, has very different genitalia). Clypeogenal angles without tubercle. Body length
<taxonomicName id="4C027093FFBDFFDDA492FE6CFF2BFDF7" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="517" phylum="Arthropoda" rank="species" species="walshi">
<emphasis id="B976D702FFBDFFDDA492FE6CFF2BFDF7" italics="true" pageId="22" pageNumber="517">B. walshi</emphasis>
</taxonomicName>
usually
<quantity id="4CFAA6F5FFBDFFDDA028FE48FE70FDF7" box="[318,443,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="22" pageNumber="517" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">1013 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C318589BFFBDFFDDA1D1FD95FD0FFD62" pageId="22" pageNumber="517" type="discussion">
<paragraph id="8BBD0B10FFBDFFDDA1D1FD95FD0FFD62" blockId="22.[151,1436,258,643]" pageId="22" pageNumber="517">
<emphasis id="B976D702FFBDFFDDA1D1FD95FE8FFDDA" bold="true" box="[199,324,546,571]" pageId="22" pageNumber="517">Comment.</emphasis>
Bornean specimens similar to
<taxonomicName id="4C027093FFBDFFDDA3BFFD94FCD8FDDA" box="[681,787,547,571]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="517" phylum="Arthropoda" rank="species" species="walshi">
<emphasis id="B976D702FFBDFFDDA3BFFD94FCD8FDDA" box="[681,787,547,571]" italics="true" pageId="22" pageNumber="517">B. walshi</emphasis>
</taxonomicName>
with old labels were seen, reputedly coming from
<collectingCountry id="F3154B80FFBDFFDDA477FD95FF27FDBF" name="Malaysia" pageId="22" pageNumber="517">north Borneo</collectingCountry>
, including Mt Kinabalu (SMTD ex Felsche, probably from a dealer)—these may have been mislabelled.
<emphasis id="B976D702FFBDFFDDA1D1FDDDFEABFD62" bold="true" box="[199,352,618,643]" pageId="22" pageNumber="517">Distribution.</emphasis>
<collectingCountry id="F3154B80FFBDFFDDA071FDDDFE13FD62" box="[359,472,618,643]" name="Indonesia" pageId="22" pageNumber="517">Indonesia</collectingCountry>
: Java;
<collectingCountry id="F3154B80FFBDFFDDA335FDDDFD71FD62" box="[547,698,618,643]" name="Malaysia" pageId="22" pageNumber="517">north Borneo</collectingCountry>
?
</paragraph>
</subSubSection>
</treatment>
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