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<document id="FC3F6F47E3BCADB5A952AFAE1C3B07FF" ID-DOI="10.5281/zenodo.202142" ID-GBIF-Dataset="913bcfab-f6fd-4da9-968c-ef4fb6e71d89" ID-ISSN="1175-5326" ID-Zenodo-Dep="202142" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460337371841" checkinUser="plazi" docAuthor="Makarov, Slobodan E., Ćurčić, Božidar P. M., Tomić, Vladimir T., Rađa, Tonći, Rađa, Biljana, Ćurčić, Srećko B., Mitić, Bojan M. &amp; Lučić, Luka R." docDate="2011" docId="9862879FFFFBFFDCFF49FA8D89D9D11E" docLanguage="en" docName="zt02994p044.pdf" docOrigin="Zootaxa 2994" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Heterolatzeliidae Verhoeff 1897" docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="645BFFE7FFFAFFD9FFDEFF988D20D52A" masterDocTitle="Revision of the family Heterolatzeliidae (Diplopoda, Chordeumatida)" masterLastPageNumber="44" masterPageNumber="33" pageNumber="34" updateTime="1698260178417" updateUser="plazi">
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<mods:title id="582A0AF192E21C87A5BBED5422246C69">Revision of the family Heterolatzeliidae (Diplopoda, Chordeumatida)</mods:title>
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<mods:name id="CB241936ACD1F8A4DCC7D4C95289D94E" type="personal">
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<mods:namePart id="A4ACBE2DE99067059F848AF45694A526">Makarov, Slobodan E.</mods:namePart>
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<mods:namePart id="BDF01405B01A4D7D10F0F66F76C427D2">Ćurčić, Božidar P. M.</mods:namePart>
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<mods:namePart id="244719126F83DA83543BB811062961CB">Tomić, Vladimir T.</mods:namePart>
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<mods:namePart id="76AA19643A7D510EAE8BC8978ED1EACA">Rađa, Tonći</mods:namePart>
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<mods:namePart id="B90D70D50C5AD21761EDD4A8C4EDA2A0">Rađa, Biljana</mods:namePart>
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<mods:namePart id="CFC3EC5DE4EE7B3E444C21413759D7FA">Ćurčić, Srećko B.</mods:namePart>
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<mods:namePart id="AFCE3C5F21807E6B5B6A912A0D2BF823">Mitić, Bojan M.</mods:namePart>
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<mods:namePart id="C441F80092AEB26DFDDE4B66503747C8">Lučić, Luka R.</mods:namePart>
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<mods:date id="CF660AE10A592C52829CCF0D2919887C">2011</mods:date>
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<mods:number id="58039B5F4498F3953C1DDDC54D57509A">2994</mods:number>
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<treatment id="9862879FFFFBFFDCFF49FA8D89D9D11E" ID-DOI="http://doi.org/10.5281/zenodo.6186622" ID-GBIF-Taxon="119405460" ID-Zenodo-Dep="6186622" LSID="urn:lsid:plazi:treatment:9862879FFFFBFFDCFF49FA8D89D9D11E" httpUri="http://treatment.plazi.org/id/9862879FFFFBFFDCFF49FA8D89D9D11E" lastPageId="5" lastPageNumber="38" pageId="1" pageNumber="34">
<subSubSection id="58D16502FFFBFFD8FF49FA8D8FA7D005" box="[151,647,1301,1327]" pageId="1" pageNumber="34" type="nomenclature">
<paragraph id="10743689FFFBFFD8FF49FA8D8FA7D005" blockId="1.[151,647,1301,1327]" box="[151,647,1301,1327]" pageId="1" pageNumber="34">
<heading id="4B3C81E5FFFBFFD8FF49FA8D8FA7D005" bold="true" box="[151,647,1301,1327]" fontSize="11" level="1" pageId="1" pageNumber="34" reason="1">
<emphasis id="22BFEA9BFFFBFFD8FF49FA8D8FA7D005" bold="true" box="[151,647,1301,1327]" pageId="1" pageNumber="34">
Family
<taxonomicName id="D7CB4D0AFFFBFFD8FF28FA8D8FA7D005" ID-CoL="7NJM5" authority="Verhoeff, 1897" authorityName="Verhoeff" authorityYear="1897" box="[246,647,1301,1327]" class="Diplopoda" family="Heterolatzeliidae" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="34" phylum="Arthropoda" rank="family">
Heterolatzeliidae
<bibRefCitation id="745A4B78FFFBFFD8FE0CFA8D8FA7D005" author="Verhoeff" box="[466,647,1301,1327]" pageId="1" pageNumber="34" refString="Verhoeff, K. W. (1897) Ueber Diplopoden aus Bosnien, Herzegowina und Dalmatien. III Theil: Chordeumidae und Lysiopetalidae. Archiv fur Naturgeschichte, 63, 181 - 204." type="journal article" year="1897">Verhoeff, 1897</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="58D16502FFFBFFD8FF49FAC48E2ED0BD" pageId="1" pageNumber="34" type="materials_examined">
<paragraph id="10743689FFFBFFD8FF49FAC48FA0D05E" blockId="1.[151,1436,1372,1648]" box="[151,640,1372,1396]" pageId="1" pageNumber="34">
<emphasis id="22BFEA9BFFFBFFD8FF49FAC48C3CD05E" bold="true" box="[151,284,1372,1396]" pageId="1" pageNumber="34">
<typeStatus id="CF70882BFFFBFFD8FF49FAC48DF2D05E" box="[151,210,1372,1396]" pageId="1" pageNumber="34">Type</typeStatus>
genus
</emphasis>
:
<taxonomicName id="D7CB4D0AFFFBFFD8FEF2FAC48F5CD05E" authority="Verhoeff, 1897" authorityName="Verhoeff" authorityYear="1897" box="[300,636,1372,1396]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="22BFEA9BFFFBFFD8FEF2FAC48CEBD05E" box="[300,459,1372,1396]" italics="true" pageId="1" pageNumber="34">Heterolatzelia</emphasis>
<bibRefCitation id="745A4B78FFFBFFD8FE0CFAC48F5CD05E" author="Verhoeff" box="[466,636,1372,1396]" pageId="1" pageNumber="34" refString="Verhoeff, K. W. (1897) Ueber Diplopoden aus Bosnien, Herzegowina und Dalmatien. III Theil: Chordeumidae und Lysiopetalidae. Archiv fur Naturgeschichte, 63, 181 - 204." type="journal article" year="1897">Verhoeff, 1897</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="10743689FFFBFFD8FF1BFA188E2ED0BD" blockId="1.[151,1436,1372,1648]" box="[197,782,1407,1432]" pageId="1" pageNumber="34">
<emphasis id="22BFEA9BFFFBFFD8FF1BFA188C40D0B2" bold="true" box="[197,352,1407,1432]" pageId="1" pageNumber="34">
<typeStatus id="CF70882BFFFBFFD8FF1BFA188DDED0B2" box="[197,254,1408,1432]" pageId="1" pageNumber="34">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="D7CB4D0AFFFBFFD8FEB9FA188E29D0BD" authority="Verhoeff, 1897" authorityName="Verhoeff" authorityYear="1897" box="[359,777,1407,1432]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="nivalis">
<emphasis id="22BFEA9BFFFBFFD8FEB9FA188F78D0B2" box="[359,600,1408,1432]" italics="true" pageId="1" pageNumber="34">Heterolatzelia nivalis</emphasis>
<bibRefCitation id="745A4B78FFFBFFD8FD81FAE78E29D0BD" author="Verhoeff" box="[607,777,1407,1431]" pageId="1" pageNumber="34" refString="Verhoeff, K. W. (1897) Ueber Diplopoden aus Bosnien, Herzegowina und Dalmatien. III Theil: Chordeumidae und Lysiopetalidae. Archiv fur Naturgeschichte, 63, 181 - 204." type="journal article" year="1897">Verhoeff, 1897</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="58D16502FFFBFFD8FF1BFA3B8FE3D3F2" pageId="1" pageNumber="34" type="diagnosis">
<paragraph id="10743689FFFBFFD8FF1BFA3B8E4CD35A" blockId="1.[151,1436,1372,1648]" pageId="1" pageNumber="34">
<emphasis id="22BFEA9BFFFBFFD8FF1BFA3B8C1FD096" bold="true" box="[197,319,1443,1468]" pageId="1" pageNumber="34">Diagnosis.</emphasis>
A family in the superfamily Brannerioidea. Includes species with 29 pleurotegites + telson in adults. Lateral keels relatively small, rounded or dorsally flattened. Head with ocelli. Gnathochilarium with promentum. Leg-pairs 3–7 with tarsal papillae, without any additional processes. Leg-pairs 10 and 11 of adult males with coxal glands and without coxal processes. Anterior gonopods without flagella or cheirites, with straight or claw-like angiocoxites, simple colpocoxites, and comb-like posterior processes. Posterior gonopods with prominent wide ladle-like coxites, with or without apical setulose part.
</paragraph>
<paragraph id="10743689FFFBFFD8FF49F9268FE3D3F2" blockId="1.[151,707,1725,1752]" box="[151,707,1725,1752]" pageId="1" pageNumber="34">
<emphasis id="22BFEA9BFFFBFFD8FF49F9268F34D3F2" bold="true" box="[151,532,1726,1752]" pageId="1" pageNumber="34">
<emphasis id="22BFEA9BFFFBFFD8FF49F9268C6FD3F2" bold="true" box="[151,335,1726,1752]" italics="true" pageId="1" pageNumber="34">Massarilatzelia</emphasis>
Makarov &amp; Ra
</emphasis>
đ
<emphasis id="22BFEA9BFFFBFFD8FDFAF9268FE3D3F2" bold="true" box="[548,707,1726,1752]" pageId="1" pageNumber="34">a, new genus</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="58D16502FFFBFFD8FF49F89D8EF2D237" box="[151,978,1797,1822]" pageId="1" pageNumber="34" type="materials_examined">
<paragraph id="10743689FFFBFFD8FF49F89D8EF2D237" blockId="1.[151,1436,1797,2038]" box="[151,978,1797,1822]" pageId="1" pageNumber="34">
<emphasis id="22BFEA9BFFFBFFD8FF49F89D8C12D234" bold="true" box="[151,306,1797,1822]" pageId="1" pageNumber="34">
<typeStatus id="CF70882BFFFBFFD8FF49F89D8DF2D237" box="[151,210,1797,1821]" pageId="1" pageNumber="34">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="D7CB4D0AFFFBFFD8FEE7F89D8E60D237" authority="Makarov &amp; Rada" authorityName="Makarov &amp; Rada" box="[313,832,1797,1821]" class="Diplopoda" family="Heterolatzeliidae" genus="Massarilatzelia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="dugopoljica" status="sp. nov.">
<emphasis id="22BFEA9BFFFBFFD8FEE7F89D8F54D237" box="[313,628,1797,1821]" italics="true" pageId="1" pageNumber="34">Massarilatzelia dugopoljica</emphasis>
Makarov &amp; Rađa
</taxonomicName>
,
<taxonomicNameLabel id="398C57E0FFFBFFD8FC95F89E8EF2D237" box="[843,978,1797,1821]" pageId="1" pageNumber="34" rank="species">new species</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="58D16502FFFBFFD8FF19F8B2898BD26B" box="[199,1195,1833,1858]" pageId="1" pageNumber="34" type="etymology">
<paragraph id="10743689FFFBFFD8FF19F8B2898BD26B" blockId="1.[151,1436,1797,2038]" box="[199,1195,1833,1858]" pageId="1" pageNumber="34">
<emphasis id="22BFEA9BFFFBFFD8FF19F8B28C6ED268" bold="true" box="[199,334,1834,1858]" pageId="1" pageNumber="34">Etymology.</emphasis>
The new genus is named after Massarum, the Latin name for the Mt. Mosor.
</paragraph>
</subSubSection>
<subSubSection id="58D16502FFFBFFDBFF19F8D58CC0D415" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34" type="description">
<paragraph id="10743689FFFBFFD8FF19F8D58E66D2DF" blockId="1.[151,1436,1797,2038]" pageId="1" pageNumber="34">
<emphasis id="22BFEA9BFFFBFFD8FF19F8D58C79D24C" bold="true" box="[199,345,1869,1894]" pageId="1" pageNumber="34">Description.</emphasis>
Adults with 29 pleurotergites + telson. Body length 21.1–
<quantity id="D7339B6CFFFBFFD8FC3AF8D58969D24F" box="[996,1097,1869,1893]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="1" pageNumber="34" unit="mm" value="15.1">15.1 mm</quantity>
. Lateral keels small, with flattened dorsal side, disappearing on somites XXVIII and XXIX. Macrochaetae almost in a straight line; on all somites length and shape of macrochaetae are identical. Frontal side of head in adult male flattened, in females concave. Head with ocelli arranged in 4 or 5 rows. Antennomeres III–VI with one, four, one, and four long sensitive setae, respectively. Leg pairs III–VII with tarsal papillae.
</paragraph>
<paragraph id="10743689FFF8FFDBFF19FF008CC3D5DD" blockId="2.[151,1436,151,463]" pageId="2" pageNumber="35">
<emphasis id="22BFEA9BFFF8FFDBFF19FF008C63D59A" box="[199,323,152,176]" italics="true" pageId="2" pageNumber="35">Gonopods:</emphasis>
Anterior gonopods with massive angiocoxites, claw-like, curved caudally; reduced subquadrangular or ovoid colpocoxites and comb-like caudal processes. Posterior gonopods with prominent coxite and ovoid or subovoid setulose apical part.
</paragraph>
<paragraph id="10743689FFF8FFDBFF19FE9C8CC0D415" blockId="2.[151,1436,151,463]" pageId="2" pageNumber="35">
<emphasis id="22BFEA9BFFF8FFDBFF19FE9C8C3AD436" box="[199,282,260,284]" italics="true" pageId="2" pageNumber="35">Vulvae:</emphasis>
Bursae slightly flattened dorsally, tuberculated and covered with setae only on lateral side. Operculum on both sides with setal field.
</paragraph>
</subSubSection>
<subSubSection id="58D16502FFF8FFDBFF19FED3897ED44E" box="[199,1118,331,356]" pageId="2" pageNumber="35" type="distribution">
<paragraph id="10743689FFF8FFDBFF19FED3897ED44E" blockId="2.[151,1436,151,463]" box="[199,1118,331,356]" pageId="2" pageNumber="35">
<emphasis id="22BFEA9BFFF8FFDBFF19FED38C40D44E" bold="true" box="[199,352,331,356]" pageId="2" pageNumber="35">Distribution.</emphasis>
Middle Dalmatia,
<collectingCountry id="68DC7619FFF8FFDBFDEDFED48FA9D44E" box="[563,649,332,356]" name="Croatia" pageId="2" pageNumber="35">Croatia</collectingCountry>
; troglophilic and possibly endemic taxon.
</paragraph>
</subSubSection>
<subSubSection id="58D16502FFF8FFDBFF19FEE88C42D774" pageId="2" pageNumber="35" type="discussion">
<paragraph id="10743689FFF8FFDBFF19FEE88C89D4E5" blockId="2.[151,1436,151,463]" pageId="2" pageNumber="35">
<emphasis id="22BFEA9BFFF8FFDBFF19FEE88C1BD4A2" bold="true" box="[199,315,368,392]" pageId="2" pageNumber="35">Remarks.</emphasis>
The new genus clearly differs from the representatives of the genus
<taxonomicName id="D7CB4D0AFFF8FFDBFBECFEE889F1D4A2" box="[1074,1233,368,392]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="22BFEA9BFFF8FFDBFBECFEE889F1D4A2" box="[1074,1233,368,392]" italics="true" pageId="2" pageNumber="35">Heterolatzelia</emphasis>
</taxonomicName>
in the presence of massive claw-like angiocoxites, considerably shorter colpocoxites, and in the distribution of the pleurotergal macrochaetae almost in line.
</paragraph>
<paragraph id="10743689FFF8FFDBFF49FDBC8C42D774" blockId="2.[151,893,547,606]" pageId="2" pageNumber="35">
<taxonomicName id="D7CB4D0AFFF8FFDBFF49FDBC8F9FD717" authority="Makarov and Ra" authorityName="Makarov and Ra" box="[151,703,547,574]" class="Diplopoda" family="Heterolatzeliidae" genus="Massarilatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="dugopoljica">
<emphasis id="22BFEA9BFFF8FFDBFF49FDBC8F9FD717" bold="true" box="[151,703,547,574]" pageId="2" pageNumber="35">
<emphasis id="22BFEA9BFFF8FFDBFF49FDBC8CC4D714" bold="true" box="[151,484,548,574]" italics="true" pageId="2" pageNumber="35">Massarilatzelia dugopoljica</emphasis>
Makarov and Ra
</emphasis>
</taxonomicName>
đ
<emphasis id="22BFEA9BFFF8FFDBFD0EFDBB8E5DD717" bold="true" box="[720,893,547,573]" pageId="2" pageNumber="35">a, new species</emphasis>
<figureCitation id="88F02A0CFFF8FFDBFF49FDDE8C31D774" box="[151,273,582,606]" captionStart-0="FIGURES 2 – 12" captionStart-1="FIGURES 13 – 19" captionStart-2="FIGURES 20 – 23" captionStartId-0="3.[151,264,1899,1922]" captionStartId-1="4.[151,264,1484,1507]" captionStartId-2="6.[151,264,1870,1893]" captionTargetBox-0="[307,1277,199,1857]" captionTargetBox-1="[169,1430,197,1441]" captionTargetBox-2="[153,1426,378,1869]" captionTargetId-0="figure@3.[299,1288,193,1878]" captionTargetId-1="figure@4.[151,1436,193,1463]" captionTargetId-2="figure@6.[151,1436,329,1929]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="FIGURES 2 – 12. Massarilatzelia dugopoljica n. gen., n. sp., holotype male from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 2, leg I, lateral view; 3, leg II, lateral view; 4, leg III, lateral view; 5, tarsus III, lateral view; 6, leg IV, lateral view; 7, leg V, lateral view; 8, leg VI, lateral view; 9, leg VII, lateral view; 10, tarsus VII, lateral view; 11, lag X, lateral view; 12, coxa XI, lateral view. Scale line = 1 mm." captionText-1="FIGURES 13 – 19. Massarilatzelia dugopoljica n. gen., n. sp., holotype male (Figs 13 – 16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view. a, angiocoxite; b, colpocoxite; c, comb-like process; d, coxite; e, apical lobe. Scale line = 0.2 mm; figure 18 without scale line." captionText-2="FIGURES 20 – 23. Massarilatzelia dugopoljica n. gen., n. sp., allotype female (Fig. 19), from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia, Heterolatzelia nivalis rupivaga, paratype male (Figs. 21 and 22), from Mt. Plasa, Bosnia and Herzegovina, and H. nivalis absoloni, syntype male (Fig. 23), from Ponor Pit, Gatačko polje, Trebinje region, Bosnia and Hercegowina: 20, vulva, dorsal view; 21, anterior gonopods, caudal view; 22, posterior gonopods, oral view; 23, posterior gonopods, oral view. Scale lines = 0.2 mm." httpUri-0="https://zenodo.org/record/202144/files/figure.png" httpUri-1="https://zenodo.org/record/202145/files/figure.png" httpUri-2="https://zenodo.org/record/202146/files/figure.png" pageId="2" pageNumber="35">Figs. 2–20</figureCitation>
; 26–27
</paragraph>
</subSubSection>
<subSubSection id="58D16502FFF8FFDBFF49FD168F96D638" pageId="2" pageNumber="35" type="materials_examined">
<paragraph id="10743689FFF8FFDBFF49FD168F96D638" blockId="2.[151,1437,654,1254]" pageId="2" pageNumber="35">
<emphasis id="22BFEA9BFFF8FFDBFF49FD168CA0D78D" bold="true" box="[151,384,654,679]" pageId="2" pageNumber="35">Material examined.</emphasis>
<typeStatus id="CF70882BFFF8FFDBFE56FD168CD0D78C" box="[392,496,654,678]" pageId="2" pageNumber="35" type="holotype">Holotype</typeStatus>
male, allotype female,
<typeStatus id="CF70882BFFF8FFDBFD26FD178E78D78D" box="[760,856,655,679]" pageId="2" pageNumber="35" type="paratype">paratype</typeStatus>
male, and
<typeStatus id="CF70882BFFF8FFDBFC0AFD178914D78D" box="[980,1076,655,679]" pageId="2" pageNumber="35" type="paratype">paratype</typeStatus>
juvenile, from the Kraljeva Peċ Cave (= Baliċa Špilja) (
<figureCitation id="88F02A0CFFF8FFDBFE79FD2A8CCBD7E0" box="[423,491,690,714]" captionStart="FIGURE 1" captionStartId="2.[151,250,1933,1956]" captionTargetBox="[340,1247,1325,1912]" captionTargetId="figure@2.[340,1247,1308,1913]" captionTargetPageId="2" captionText="FIGURE 1. Entrance in Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia." httpUri="https://zenodo.org/record/202143/files/figure.png" pageId="2" pageNumber="35">Fig. 1</figureCitation>
), Baliċi, village Dugopolje, Mt. Mosor, near Split, Dalmatia,
<collectingCountry id="68DC7619FFF8FFDBFB71FD2A8825D7E0" box="[1199,1285,690,714]" name="Croatia" pageId="2" pageNumber="35">Croatia</collectingCountry>
; collected on
<date id="64751049FFF8FFDBFF49FD4E8C3ED7C4" box="[151,286,726,750]" pageId="2" pageNumber="35" value="1981-08-22">22.viii 1981</date>
by T. Rađa (
<collectionCode id="76DAAE4CFFF8FFDBFE71FD4E8F34D7C4" box="[431,532,726,750]" pageId="2" pageNumber="35">NHMSC</collectionCode>
616);
<typeStatus id="CF70882BFFF8FFDBFD83FD4F8F9DD7C5" box="[605,701,727,751]" pageId="2" pageNumber="35" type="paratype">paratype</typeStatus>
female, and 12
<typeStatus id="CF70882BFFF8FFDBFCADFD4F8EF3D7C5" box="[883,979,727,751]" pageId="2" pageNumber="35" type="paratype">paratype</typeStatus>
juveniles, same locality; collected on
<date id="64751049FFF8FFDBFA5FFD4E8DEFD638" pageId="2" pageNumber="35" value="1980-04">iv. 1980</date>
without collectors name (IZB
<date id="64751049FFF8FFDBFDF9FD628F8AD638" box="[551,682,762,786]" pageId="2" pageNumber="35" value="1210" valueMax="1223">1210–1223</date>
).
</paragraph>
</subSubSection>
<subSubSection id="58D16502FFF8FFDBFF19FC87883AD61C" box="[199,1306,798,823]" pageId="2" pageNumber="35" type="etymology">
<paragraph id="10743689FFF8FFDBFF19FC87883AD61C" blockId="2.[151,1437,654,1254]" box="[199,1306,798,823]" pageId="2" pageNumber="35">
<emphasis id="22BFEA9BFFF8FFDBFF19FC878C6ED61D" bold="true" box="[199,334,799,823]" pageId="2" pageNumber="35">Etymology.</emphasis>
The species name is given after the locality of the new form, the village of Dugopolje.
</paragraph>
</subSubSection>
<subSubSection id="58D16502FFF8FFDCFF19FCDA89D9D11E" lastPageId="5" lastPageNumber="38" pageId="2" pageNumber="35" type="description">
<paragraph id="10743689FFF8FFDBFF19FCDA8859D688" blockId="2.[151,1437,654,1254]" pageId="2" pageNumber="35">
<emphasis id="22BFEA9BFFF8FFDBFF19FCDA8C79D671" bold="true" box="[199,345,834,859]" pageId="2" pageNumber="35">Description.</emphasis>
Body with 29 pleurotegites + telson (adults). Dorsal side of metazonites brownish, prozonites and ventral side of metazonites yellowish (
<figureCitation id="88F02A0CFFF8FFDBFD99FCFE8FF1D654" box="[583,721,870,894]" captionStart-0="FIGURES 24 – 26" captionStart-1="FIGURE 27" captionStartId-0="7.[151,264,948,971]" captionStartId-1="7.[151,250,1630,1653]" captionTargetBox-0="[161,1426,223,927]" captionTargetBox-1="[340,1246,1052,1609]" captionTargetId-0="figure@7.[161,1426,223,927]" captionTargetId-1="figure@7.[340,1247,1052,1609]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="FIGURES 24 – 26. Heterolatzelia nivalis Verhoeff, 1897 (Fig. 24); Heterolatzelia durmitorensis Gulička, 1968 (Fig. 25); Massarilatzelia dugopoljica n. gen., n. sp., (Fig. 26); without scale line." captionText-1="FIGURE 27. Massarilatzelia dugopoljica, n. gen., n. sp., paratype female, from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia." httpUri-0="https://zenodo.org/record/202147/files/figure.png" httpUri-1="https://zenodo.org/record/202148/files/figure.png" pageId="2" pageNumber="35">Figs. 26–27</figureCitation>
). Body length
<quantity id="D7339B6CFFF8FFDBFCAAFCFE8EF8D654" box="[884,984,870,894]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="2" pageNumber="35" unit="mm" value="15.1">15.1 mm</quantity>
(
<typeStatus id="CF70882BFFF8FFDBFC38FCFE896AD654" box="[998,1098,870,894]" pageId="2" pageNumber="35" type="holotype">holotype</typeStatus>
male), and
<quantity id="D7339B6CFFF8FFDBFB10FCFE8812D654" box="[1230,1330,870,894]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="2" pageNumber="35" unit="mm" value="21.1">21.1 mm</quantity>
(allotype female). Vertical diameter of the largest pleurotergites
<quantity id="D7339B6CFFF8FFDBFD25FC128E7FD688" box="[763,863,906,930]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" pageId="2" pageNumber="35" unit="mm" value="0.97">0.97 mm</quantity>
(
<typeStatus id="CF70882BFFF8FFDBFCB3FC128EF1D688" box="[877,977,906,930]" pageId="2" pageNumber="35" type="holotype">holotype</typeStatus>
male), and
<quantity id="D7339B6CFFF8FFDBFB8BFC12898AD688" box="[1109,1194,906,930]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="35" unit="mm" value="1.3">1.3 mm</quantity>
(allotype female).
</paragraph>
<paragraph id="10743689FFF8FFDBFF19FC368908D124" blockId="2.[151,1437,654,1254]" pageId="2" pageNumber="35">Head: With 4–5 rows of 20–22 ocellae, occipital axial suture clear and densely setose. Labrum with 10–12 labral and 4 supralabral setae. Gnathochilarium: promentum triangular; stipites each with 5–6 long apical and medial setae, and 14–15 basal microsetae. Lingual plates with one row of 5 setae.</paragraph>
<paragraph id="10743689FFF8FFDBFF19FB828FD5D1CC" blockId="2.[151,1437,654,1254]" pageId="2" pageNumber="35">
Antennal length
<quantity id="D7339B6CFFF8FFDBFE5CFB828CF8D118" box="[386,472,1050,1074]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="35" unit="mm" value="2.3">2.3 mm</quantity>
(
<typeStatus id="CF70882BFFF8FFDBFE38FB828F6AD118" box="[486,586,1050,1074]" pageId="2" pageNumber="35" type="holotype">holotype</typeStatus>
male), and
<quantity id="D7339B6CFFF8FFDBFD11FB828E05D118" box="[719,805,1050,1074]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="35" unit="mm" value="2.5">2.5 mm</quantity>
(allotype female). Length of antennomeres I–VII (
<typeStatus id="CF70882BFFF8FFDBFA81FB828DE7D17D" pageId="2" pageNumber="35" type="holotype">holotype</typeStatus>
male and allotype female):
<quantity id="D7339B6CFFF8FFDBFDD5FBA68F8FD17C" box="[523,687,1086,1110]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="2" pageNumber="35" unit="mm" value="0.16" valueMax="0.17" valueMin="0.15">0.15–0.17 mm</quantity>
(I);
<quantity id="D7339B6CFFF8FFDBFD3BFBA68EA9D17C" box="[741,905,1086,1110]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" metricValueMax="3.5" metricValueMin="2.7" pageId="2" pageNumber="35" unit="mm" value="0.31" valueMax="0.35" valueMin="0.27">0.27–0.35 mm</quantity>
(II);
<quantity id="D7339B6CFFF8FFDBFC17FBA6894DD17C" box="[969,1133,1086,1110]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.6" metricValueMin="5.8999999999999995" pageId="2" pageNumber="35" unit="mm" value="0.625" valueMax="0.66" valueMin="0.59">0.59–0.66 mm</quantity>
(III);
<quantity id="D7339B6CFFF8FFDBFB68FBA6887AD17C" box="[1206,1370,1086,1110]" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="3.2" metricValueMin="1.7" pageId="2" pageNumber="35" unit="mm" value="0.245" valueMax="0.32" valueMin="0.17">0.17–0.32 mm</quantity>
(IV);
<quantity id="D7339B6CFFF8FFDBFF49FBFA8C1CD150" box="[151,316,1122,1146]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" metricValueMax="6.2" metricValueMin="5.4" pageId="2" pageNumber="35" unit="mm" value="0.58" valueMax="0.62" valueMin="0.54">0.54–0.62 mm</quantity>
(V);
<quantity id="D7339B6CFFF8FFDBFEA2FBFA8F01D150" box="[380,545,1122,1146]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="2" pageNumber="35" unit="mm" value="0.25" valueMax="0.26" valueMin="0.24">0.24–0.26 mm</quantity>
(VI); and
<quantity id="D7339B6CFFF8FFDBFD43FBFA8E62D150" box="[669,834,1122,1146]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="2" pageNumber="35" unit="mm" value="0.21" valueMax="0.22" valueMin="0.2">0.20–0.22 mm</quantity>
(VII). Length/breadth ratios of antennomeres I–VII: 1.2–1.36 (I); 2.45–2.9 (II); 5.9–6.0 (III); 1.7–3.2 (IV); 3.8–5.63 (V); 1,3–2.36 (VI); and 1.3–2 (VII). Antennomeres III, IV, and VI with one, four, and four long sensitive setae, respectively. Antennomeres V with one long subapical sensitive seta, and apical ring of slightly shorter setae.
</paragraph>
<caption id="44B46601FFF8FFDBFF49F8158D97D2ED" httpUri="https://zenodo.org/record/202143/files/figure.png" pageId="2" pageNumber="35" targetBox="[340,1247,1325,1912]" targetPageId="2">
<paragraph id="10743689FFF8FFDBFF49F8158D97D2ED" blockId="2.[151,1436,1933,1991]" pageId="2" pageNumber="35">
<emphasis id="22BFEA9BFFF8FFDBFF49F8158C32D289" bold="true" box="[151,274,1933,1956]" pageId="2" pageNumber="35">FIGURE 1.</emphasis>
Entrance in Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia.
</paragraph>
</caption>
<caption id="44B46601FFF9FFDAFF49F8F3891FD2F4" httpUri="https://zenodo.org/record/202144/files/figure.png" pageId="3" pageNumber="36" targetBox="[307,1277,199,1857]" targetPageId="3">
<paragraph id="10743689FFF9FFDAFF49F8F3891FD2F4" blockId="3.[151,1436,1899,2014]" pageId="3" pageNumber="36">
<emphasis id="22BFEA9BFFF9FFDAFF49F8F38C6AD2AB" bold="true" box="[151,330,1899,1922]" pageId="3" pageNumber="36">FIGURES 2–12.</emphasis>
<taxonomicName id="D7CB4D0AFFF9FFDAFE8CF8F48F52D2A8" box="[338,626,1900,1922]" class="Diplopoda" family="Heterolatzeliidae" genus="Massarilatzelia" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="dugopoljica" status="gen. nov.">
<emphasis id="22BFEA9BFFF9FFDAFE8CF8F48F52D2A8" box="[338,626,1900,1922]" italics="true" pageId="3" pageNumber="36">Massarilatzelia dugopoljica</emphasis>
</taxonomicName>
<emphasis id="22BFEA9BFFF9FFDAFDA5F8F48E30D2AB" bold="true" box="[635,784,1900,1921]" pageId="3" pageNumber="36">
<taxonomicNameLabel id="398C57E0FFF9FFDAFDA5F8F48FE6D2AB" box="[635,710,1900,1921]" pageId="3" pageNumber="36" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
, holotype male from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 2, leg I, lateral view; 3, leg II, lateral view; 4, leg III, lateral view; 5, tarsus III, lateral view; 6, leg IV, lateral view; 7, leg V, lateral view; 8, leg VI, lateral view; 9, leg VII, lateral view; 10, tarsus VII, lateral view; 11, lag X, lateral view; 12, coxa XI, lateral view. Scale line = 1 mm.
</paragraph>
</caption>
<caption id="44B46601FFFEFFDDFF49FA548869D377" httpUri="https://zenodo.org/record/202145/files/figure.png" pageId="4" pageNumber="37" targetBox="[169,1430,197,1441]" targetPageId="4">
<paragraph id="10743689FFFEFFDDFF49FA548869D377" blockId="4.[151,1436,1484,1629]" pageId="4" pageNumber="37">
<emphasis id="22BFEA9BFFFEFFDDFF49FA548C73D0CB" bold="true" box="[151,339,1484,1507]" pageId="4" pageNumber="37">FIGURES 13–19.</emphasis>
<taxonomicName id="D7CB4D0AFFFEFFDDFE87FA548F55D0C8" box="[345,629,1484,1506]" class="Diplopoda" family="Heterolatzeliidae" genus="Massarilatzelia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="dugopoljica" status="gen. nov.">
<emphasis id="22BFEA9BFFFEFFDDFE87FA548F55D0C8" box="[345,629,1484,1506]" italics="true" pageId="4" pageNumber="37">Massarilatzelia dugopoljica</emphasis>
</taxonomicName>
<emphasis id="22BFEA9BFFFEFFDDFDA2FA558E28D0C8" bold="true" box="[636,776,1485,1506]" pageId="4" pageNumber="37">
<taxonomicNameLabel id="398C57E0FFFEFFDDFDA2FA558FE4D0C8" box="[636,708,1485,1506]" pageId="4" pageNumber="37" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
, holotype male (Figs 13–16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view.
<emphasis id="22BFEA9BFFFEFFDDFA9CF9B1886ED314" box="[1346,1358,1577,1598]" italics="true" pageId="4" pageNumber="37">a</emphasis>
, angiocoxite;
<emphasis id="22BFEA9BFFFEFFDDFF3CF9DF8DCFD377" box="[226,239,1607,1629]" italics="true" pageId="4" pageNumber="37">b</emphasis>
, colpocoxite;
<emphasis id="22BFEA9BFFFEFFDDFEA1F9D08CAAD377" box="[383,394,1608,1629]" italics="true" pageId="4" pageNumber="37">c</emphasis>
, comb-like process;
<emphasis id="22BFEA9BFFFEFFDDFD83F9DF8F4AD377" box="[605,618,1607,1629]" italics="true" pageId="4" pageNumber="37">d</emphasis>
, coxite;
<emphasis id="22BFEA9BFFFEFFDDFD1FF9D08FECD377" box="[705,716,1608,1629]" italics="true" pageId="4" pageNumber="37">e</emphasis>
, apical lobe. Scale line = 0.2 mm; figure 18 without scale line.
</paragraph>
</caption>
<paragraph id="10743689FFFEFFDDFF19F9118E2AD38B" blockId="4.[151,1436,1673,2027]" box="[199,778,1673,1697]" pageId="4" pageNumber="37">Collum narrower than head, with six macrochaetae.</paragraph>
<paragraph id="10743689FFFEFFDDFF19F9358CBBD2C1" blockId="4.[151,1436,1673,2027]" pageId="4" pageNumber="37">
Pleurotergites: Lateral keels of metazonites relatively small on anterior and median somites (
<figureCitation id="88F02A0CFFFEFFDDFB32F9358861D3EF" box="[1260,1345,1709,1733]" captionStart="FIGURES 13 – 19" captionStartId="4.[151,264,1484,1507]" captionTargetBox="[169,1430,197,1441]" captionTargetId="figure@4.[151,1436,193,1463]" captionTargetPageId="4" captionText="FIGURES 13 – 19. Massarilatzelia dugopoljica n. gen., n. sp., holotype male (Figs 13 – 16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view. a, angiocoxite; b, colpocoxite; c, comb-like process; d, coxite; e, apical lobe. Scale line = 0.2 mm; figure 18 without scale line." httpUri="https://zenodo.org/record/202145/files/figure.png" pageId="4" pageNumber="37">Fig. 19</figureCitation>
), disappearing on somites XXVIII and XXIX. In width: collum &lt;II &lt;III &lt;IV &lt;V &lt;VI &lt;VII &lt;VIII &lt;IX = X–XII &lt;XIII =XIV–XVIII&gt; XIX&gt; XX&gt; XXI = XXII–XXIV&gt; XXV&gt; XXVI&gt; XXVII&gt; XXVIII&gt; XXIX. Macrochaetae short; maximal length of macrochaetae on mid-body somites
<quantity id="D7339B6CFFFEFFDDFC9FF8818E87D21B" box="[833,935,1817,1841]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="37" unit="mm" value="0.14">0.14 mm</quantity>
. Macrochaetal index CIX (15),
<emphasis id="22BFEA9BFFFEFFDDFAD9F8838803D218" box="[1287,1315,1819,1842]" italics="true" pageId="4" pageNumber="37">i.e</emphasis>
., (distance between exterior and median macrochaetae/distance between interior and median macrochaetae) = 1.61 (female) – 1.33 (male). Median index MIX (15),
<emphasis id="22BFEA9BFFFEFFDDFDBAF8FB8FA0D250" box="[612,640,1891,1914]" italics="true" pageId="4" pageNumber="37">i.e</emphasis>
., (distance between interior macrochaetae and axial suture/distance between interior and median macrochaetae) = 0.67 (female) – 0.86 (male). Macrochaetal angle between arm created by median and exterior macrochaetae and arm formed by median and interior macrochaetae, MA (15)
<emphasis id="22BFEA9BFFFEFFDDFA8DF8358842D2EF" bold="true" box="[1363,1378,1965,1989]" pageId="4" pageNumber="37">≈</emphasis>
173o (female) – 176o (male).
</paragraph>
<paragraph id="10743689FFFFFFDCFF19FF0F8FEBD5FE" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="38">Epiproct truncated caudally, with two typical spinnerets; each paraproct with three long marginal setae; hypoproct subquadrangular, with two long apical setae.</paragraph>
<paragraph id="10743689FFFFFFDCFF19FF788CC0D436" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="38">
Legs elongated, 1.23–1.61 times as long as mid-body height. Leg-pairs 1 and 2 slightly reduced, with tarsal brushes ventrally (
<figureCitation id="88F02A0CFFFFFFDCFEB8FE9C8CF3D436" box="[358,467,260,284]" captionStart="FIGURES 2 – 12" captionStartId="3.[151,264,1899,1922]" captionTargetBox="[307,1277,199,1857]" captionTargetId="figure@3.[299,1288,193,1878]" captionTargetPageId="3" captionText="FIGURES 2 – 12. Massarilatzelia dugopoljica n. gen., n. sp., holotype male from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 2, leg I, lateral view; 3, leg II, lateral view; 4, leg III, lateral view; 5, tarsus III, lateral view; 6, leg IV, lateral view; 7, leg V, lateral view; 8, leg VI, lateral view; 9, leg VII, lateral view; 10, tarsus VII, lateral view; 11, lag X, lateral view; 12, coxa XI, lateral view. Scale line = 1 mm." httpUri="https://zenodo.org/record/202144/files/figure.png" pageId="5" pageNumber="38">Figs. 2–3</figureCitation>
).
</paragraph>
<paragraph id="10743689FFFFFFDCFF19FEBF8EB9D716" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="38">
Male sexual characters. Frontal side of head in adult male flattened. Leg-pairs 3–7 not greatly enlarged and modified as in many chordeumatids. In the new species these legs are supplied with tarsal papillae and with all podomeres are more or less slightly bulging. Tarsus of leg 3 (
<figureCitation id="88F02A0CFFFFFFDCFC9DFEF78E8DD4AD" box="[835,941,367,391]" captionStart="FIGURES 2 – 12" captionStartId="3.[151,264,1899,1922]" captionTargetBox="[307,1277,199,1857]" captionTargetId="figure@3.[299,1288,193,1878]" captionTargetPageId="3" captionText="FIGURES 2 – 12. Massarilatzelia dugopoljica n. gen., n. sp., holotype male from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 2, leg I, lateral view; 3, leg II, lateral view; 4, leg III, lateral view; 5, tarsus III, lateral view; 6, leg IV, lateral view; 7, leg V, lateral view; 8, leg VI, lateral view; 9, leg VII, lateral view; 10, tarsus VII, lateral view; 11, lag X, lateral view; 12, coxa XI, lateral view. Scale line = 1 mm." httpUri="https://zenodo.org/record/202144/files/figure.png" pageId="5" pageNumber="38">Figs. 4–5</figureCitation>
) with 3–4 rows of claw-like accessory setae and two rows of papillae. Legs 4–6 (
<figureCitation id="88F02A0CFFFFFFDCFDF0FE0C8FB9D486" box="[558,665,404,428]" captionStart="FIGURES 2 – 12" captionStartId="3.[151,264,1899,1922]" captionTargetBox="[307,1277,199,1857]" captionTargetId="figure@3.[299,1288,193,1878]" captionTargetPageId="3" captionText="FIGURES 2 – 12. Massarilatzelia dugopoljica n. gen., n. sp., holotype male from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 2, leg I, lateral view; 3, leg II, lateral view; 4, leg III, lateral view; 5, tarsus III, lateral view; 6, leg IV, lateral view; 7, leg V, lateral view; 8, leg VI, lateral view; 9, leg VII, lateral view; 10, tarsus VII, lateral view; 11, lag X, lateral view; 12, coxa XI, lateral view. Scale line = 1 mm." httpUri="https://zenodo.org/record/202144/files/figure.png" pageId="5" pageNumber="38">Figs. 6–8</figureCitation>
) with ventral side of coxae, basoventral and dorsal side of prefemora denticulate, and with rows of tarsal papillae. Leg 7 with slightly denticulate dorso-subapical margins of prefemora, and with one row of claw-like setae and row of papillae (
<figureCitation id="88F02A0CFFFFFFDCFCFCFE448EBED4DE" box="[802,926,476,500]" captionStart="FIGURES 2 – 12" captionStartId="3.[151,264,1899,1922]" captionTargetBox="[307,1277,199,1857]" captionTargetId="figure@3.[299,1288,193,1878]" captionTargetPageId="3" captionText="FIGURES 2 – 12. Massarilatzelia dugopoljica n. gen., n. sp., holotype male from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 2, leg I, lateral view; 3, leg II, lateral view; 4, leg III, lateral view; 5, tarsus III, lateral view; 6, leg IV, lateral view; 7, leg V, lateral view; 8, leg VI, lateral view; 9, leg VII, lateral view; 10, tarsus VII, lateral view; 11, lag X, lateral view; 12, coxa XI, lateral view. Scale line = 1 mm." httpUri="https://zenodo.org/record/202144/files/figure.png" pageId="5" pageNumber="38">Figs. 9–10</figureCitation>
). Leg 10 and 11 with both dorsal and ventral margins of prefemora denticulate, as well as all surfaces of coxa (leg 11) or only ventral side (leg 10) (
<figureCitation id="88F02A0CFFFFFFDCFAD9FE6788AED73D" box="[1287,1422,511,535]" captionStart="FIGURES 2 – 12" captionStartId="3.[151,264,1899,1922]" captionTargetBox="[307,1277,199,1857]" captionTargetId="figure@3.[299,1288,193,1878]" captionTargetPageId="3" captionText="FIGURES 2 – 12. Massarilatzelia dugopoljica n. gen., n. sp., holotype male from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 2, leg I, lateral view; 3, leg II, lateral view; 4, leg III, lateral view; 5, tarsus III, lateral view; 6, leg IV, lateral view; 7, leg V, lateral view; 8, leg VI, lateral view; 9, leg VII, lateral view; 10, tarsus VII, lateral view; 11, lag X, lateral view; 12, coxa XI, lateral view. Scale line = 1 mm." httpUri="https://zenodo.org/record/202144/files/figure.png" pageId="5" pageNumber="38">Figs. 11–12</figureCitation>
); both leg pairs with coxal glands, and without any lobes or processes.
</paragraph>
<paragraph id="10743689FFFFFFDCFF19FDDF8921D655" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="38">
Anterior gonopods. The anterior gonopods with a single wide protective subquadrangular part, covering almost half the cephalic side on both gonopods; on both lateral sides this element is expanded into two narrower points (
<figureCitation id="88F02A0CFFFFFFDCFF35FD178C1FD78D" box="[235,319,655,679]" captionStart="FIGURES 13 – 19" captionStartId="4.[151,264,1484,1507]" captionTargetBox="[169,1430,197,1441]" captionTargetId="figure@4.[151,1436,193,1463]" captionTargetPageId="4" captionText="FIGURES 13 – 19. Massarilatzelia dugopoljica n. gen., n. sp., holotype male (Figs 13 – 16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view. a, angiocoxite; b, colpocoxite; c, comb-like process; d, coxite; e, apical lobe. Scale line = 0.2 mm; figure 18 without scale line." httpUri="https://zenodo.org/record/202145/files/figure.png" pageId="5" pageNumber="38">Fig. 13</figureCitation>
). Both sides of gonopods clearly separated and consist of three elements. Massive angiocoxites are dominant, with wide basal part, and distal claw-like process curved caudally (
<figureCitation id="88F02A0CFFFFFFDCFBD5FD2C899FD7E6" box="[1035,1215,692,716]" captionStart="FIGURES 13 – 19" captionStartId="4.[151,264,1484,1507]" captionTargetBox="[169,1430,197,1441]" captionTargetId="figure@4.[151,1436,193,1463]" captionTargetPageId="4" captionText="FIGURES 13 – 19. Massarilatzelia dugopoljica n. gen., n. sp., holotype male (Figs 13 – 16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view. a, angiocoxite; b, colpocoxite; c, comb-like process; d, coxite; e, apical lobe. Scale line = 0.2 mm; figure 18 without scale line." httpUri="https://zenodo.org/record/202145/files/figure.png" pageId="5" pageNumber="38">Figs. 13–14, 18</figureCitation>
). In
<typeStatus id="CF70882BFFFFFFDCFB2AFD2C8876D7E6" box="[1268,1366,692,716]" pageId="5" pageNumber="38" type="holotype">holotype</typeStatus>
male, apical angiocoxites carry three small pointed processes and one sub-basal denticulated lamella (
<figureCitation id="88F02A0CFFFFFFDCFB3DFD4F884FD7C5" box="[1251,1391,727,751]" captionStart="FIGURES 13 – 19" captionStartId="4.[151,264,1484,1507]" captionTargetBox="[169,1430,197,1441]" captionTargetId="figure@4.[151,1436,193,1463]" captionTargetPageId="4" captionText="FIGURES 13 – 19. Massarilatzelia dugopoljica n. gen., n. sp., holotype male (Figs 13 – 16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view. a, angiocoxite; b, colpocoxite; c, comb-like process; d, coxite; e, apical lobe. Scale line = 0.2 mm; figure 18 without scale line." httpUri="https://zenodo.org/record/202145/files/figure.png" pageId="5" pageNumber="38">Figs. 13–14</figureCitation>
). In
<typeStatus id="CF70882BFFFFFFDCFF49FD658DD7D63F" box="[151,247,765,789]" pageId="5" pageNumber="38" type="paratype">paratype</typeStatus>
male this part consists of one pointed claw-like process and a shorter narrower one; subapically with two or three small lobes (
<figureCitation id="88F02A0CFFFFFFDCFE59FC878CFCD61D" box="[391,476,799,823]" captionStart="FIGURES 13 – 19" captionStartId="4.[151,264,1484,1507]" captionTargetBox="[169,1430,197,1441]" captionTargetId="figure@4.[151,1436,193,1463]" captionTargetPageId="4" captionText="FIGURES 13 – 19. Massarilatzelia dugopoljica n. gen., n. sp., holotype male (Figs 13 – 16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view. a, angiocoxite; b, colpocoxite; c, comb-like process; d, coxite; e, apical lobe. Scale line = 0.2 mm; figure 18 without scale line." httpUri="https://zenodo.org/record/202145/files/figure.png" pageId="5" pageNumber="38">Fig. 18</figureCitation>
). Second element are colpocoxites, subquadrangular in
<typeStatus id="CF70882BFFFFFFDCFB8BFC878997D61D" box="[1109,1207,799,823]" pageId="5" pageNumber="38" type="holotype">holotype</typeStatus>
male (
<figureCitation id="88F02A0CFFFFFFDCFADBFC8788AFD61D" box="[1285,1423,799,823]" captionStart="FIGURES 13 – 19" captionStartId="4.[151,264,1484,1507]" captionTargetBox="[169,1430,197,1441]" captionTargetId="figure@4.[151,1436,193,1463]" captionTargetPageId="4" captionText="FIGURES 13 – 19. Massarilatzelia dugopoljica n. gen., n. sp., holotype male (Figs 13 – 16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view. a, angiocoxite; b, colpocoxite; c, comb-like process; d, coxite; e, apical lobe. Scale line = 0.2 mm; figure 18 without scale line." httpUri="https://zenodo.org/record/202145/files/figure.png" pageId="5" pageNumber="38">Figs. 13–14</figureCitation>
), but ovoid with denticulated mesal side in
<typeStatus id="CF70882BFFFFFFDCFDA1FCDD8FC0D677" box="[639,736,837,861]" pageId="5" pageNumber="38" type="paratype">paratype</typeStatus>
male (
<figureCitation id="88F02A0CFFFFFFDCFCEDFCDC8EA9D676" box="[819,905,836,860]" captionStart="FIGURES 13 – 19" captionStartId="4.[151,264,1484,1507]" captionTargetBox="[169,1430,197,1441]" captionTargetId="figure@4.[151,1436,193,1463]" captionTargetPageId="4" captionText="FIGURES 13 – 19. Massarilatzelia dugopoljica n. gen., n. sp., holotype male (Figs 13 – 16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view. a, angiocoxite; b, colpocoxite; c, comb-like process; d, coxite; e, apical lobe. Scale line = 0.2 mm; figure 18 without scale line." httpUri="https://zenodo.org/record/202145/files/figure.png" pageId="5" pageNumber="38">Fig. 18</figureCitation>
). Third element is a comb-like process, fused basally with the angiocoxites (possibly also derived from coxa) (
<figureCitation id="88F02A0CFFFFFFDCFCB5FCFF8ED4D655" box="[875,1012,871,895]" captionStart="FIGURES 13 – 19" captionStartId="4.[151,264,1484,1507]" captionTargetBox="[169,1430,197,1441]" captionTargetId="figure@4.[151,1436,193,1463]" captionTargetPageId="4" captionText="FIGURES 13 – 19. Massarilatzelia dugopoljica n. gen., n. sp., holotype male (Figs 13 – 16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view. a, angiocoxite; b, colpocoxite; c, comb-like process; d, coxite; e, apical lobe. Scale line = 0.2 mm; figure 18 without scale line." httpUri="https://zenodo.org/record/202145/files/figure.png" pageId="5" pageNumber="38">Figs. 14, 18</figureCitation>
).
</paragraph>
<paragraph id="10743689FFFFFFDCFF19FC148FE3D6C6" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="38">
Posterior gonopods. The posterior gonopods consist of coxite (
<figureCitation id="88F02A0CFFFFFFDCFC57FC14892FD68E" box="[905,1039,908,932]" captionStart="FIGURES 13 – 19" captionStartId="4.[151,264,1484,1507]" captionTargetBox="[169,1430,197,1441]" captionTargetId="figure@4.[151,1436,193,1463]" captionTargetPageId="4" captionText="FIGURES 13 – 19. Massarilatzelia dugopoljica n. gen., n. sp., holotype male (Figs 13 – 16, 19) and paratype male (Figs. 17 and 18) from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia: 13, anterior gonopods, oral view; 14, anterior gonopods, caudal view; 15, posterior gonopods, caudal view; 16, posterior gonopods, oral view; 17, posterior gonopods, oral view; 18, anterior gonopods, caudal view; 19, pleurotergites XV, dorsal view. a, angiocoxite; b, colpocoxite; c, comb-like process; d, coxite; e, apical lobe. Scale line = 0.2 mm; figure 18 without scale line." httpUri="https://zenodo.org/record/202145/files/figure.png" pageId="5" pageNumber="38">Figs. 15–17</figureCitation>
) and ovoid or subovoid apical process. The later is fused with coxite on cephalic side, covered with numerous setae on dorsal side and functioning as protective part for elements of anterior gonopods.
</paragraph>
<paragraph id="10743689FFFFFFDCFF19FC6F89D9D11E" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="38">
Female sexual characters. Mesal side of bursae without setae; lateral side covered with tubercles and with subapical field with moderate long setae (
<figureCitation id="88F02A0CFFFFFFDCFD9DFB848FB7D11E" box="[579,663,1052,1076]" captionStart="FIGURES 20 – 23" captionStartId="6.[151,264,1870,1893]" captionTargetBox="[153,1426,378,1869]" captionTargetId="figure@6.[151,1436,329,1929]" captionTargetPageId="6" captionText="FIGURES 20 – 23. Massarilatzelia dugopoljica n. gen., n. sp., allotype female (Fig. 19), from the Kraljeva Peć Cave (= Balića Špilja), Balići, village Dugopolje, Mt. Mosor, near Split, Dalmatia, Croatia, Heterolatzelia nivalis rupivaga, paratype male (Figs. 21 and 22), from Mt. Plasa, Bosnia and Herzegovina, and H. nivalis absoloni, syntype male (Fig. 23), from Ponor Pit, Gatačko polje, Trebinje region, Bosnia and Hercegowina: 20, vulva, dorsal view; 21, anterior gonopods, caudal view; 22, posterior gonopods, oral view; 23, posterior gonopods, oral view. Scale lines = 0.2 mm." httpUri="https://zenodo.org/record/202146/files/figure.png" pageId="5" pageNumber="38">Fig. 20</figureCitation>
). Operculum with two groups of 6-7 long apical setae.
</paragraph>
</subSubSection>
</treatment>
</document>
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