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<mods:title id="9ECE5C7A14FA16E2E426D1F44A2672D9">A Revision Of Species In The Subgenus Nidirana Dubois, 1992, With Special Attention To The Identity Of Specimens Allocated To Rana Adenopleura Boulenger, 1909, And Rana Chapaensis (Bourret, 1937) (Amphibia: Anura: Ranidae) From Thailand And Laos</mods:title>
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<taxonomicName id="7A98E1CDFF83FFDE278BF871FA95E2CE" ID-CoL="4RDG5" authority="Boulenger, 1909" authorityName="BOULENGER" authorityYear="1909" box="[931,1328,1939,1964]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="302" phylum="Chordata" rank="species" species="adenopleura">
<emphasis id="8FEC465CFF83FFDE278BF871FA95E2CE" bold="true" box="[931,1328,1939,1964]" pageId="11" pageNumber="302">
<emphasis id="8FEC465CFF83FFDE278BF871FBD4E2C9" bold="true" box="[931,1137,1940,1964]" italics="true" pageId="11" pageNumber="302">Rana adenopleura</emphasis>
<bibRefCitation id="D909E7BFFF83FFDE2051F876FA95E2CE" author="Boulenger, G" box="[1145,1328,1939,1963]" pageId="11" pageNumber="302" pagination="492 - 495" refId="ref17936" refString="Boulenger, G. A., 1909. Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa. Annals and Magazine of Natural History, series 8: 492 - 495." type="book chapter" year="1909">Boulenger, 1909</bibRefCitation>
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<treatmentCitation id="3C39BC5FFF83FFDE2702F837FB11E282" author="Boulenger, G" box="[810,1204,2002,2023]" page="492" pageId="11" pageNumber="302" year="1909">
<taxonomicName id="7A98E1CDFF83FFDE2702F837FB11E282" ID-CoL="4RDG5" authority="Boulenger, 1909: 492" authorityName="BOULENGER" authorityPageNumber="492" authorityYear="1909" box="[810,1204,2002,2023]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="302" phylum="Chordata" rank="species" species="adenopleura">
<emphasis id="8FEC465CFF83FFDE2702F837FC7EE282" box="[810,987,2002,2023]" italics="true" pageId="11" pageNumber="302">Rana adenopleura</emphasis>
<materialsCitation id="0DF09013FF83FFDE27C9F837FB11E282" box="[993,1204,2002,2023]" country="Taiwan" county="Wucheng" elevation="1219" location="Fuhacho Village" municipality="Maobu" pageId="11" pageNumber="302" specimenCount="1" stateProvince="Nantou" typeStatus="syntype">
<bibRefCitation id="D909E7BFFF83FFDE27C9F837FB11E282" author="Boulenger, G" box="[993,1204,2002,2023]" pageId="11" pageNumber="302" pagination="492 - 495" refId="ref17936" refString="Boulenger, G. A., 1909. Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa. Annals and Magazine of Natural History, series 8: 492 - 495." type="book chapter" year="1909">Boulenger, 1909: 492</bibRefCitation>
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. [
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: BM (“several specimens”). Type-locality:
<location id="B847CC95FF83FFDE205BF80AFABEED61" LSID="urn:lsid:plazi:treatment:35312B58FF83FFD9278BF871FE55E0E7:B847CC95FF83FFDE205BF80AFABEED61" box="[1139,1307,2031,2052]" country="Taiwan" county="Wucheng" municipality="Maobu" name="Fuhacho Village" pageId="11" pageNumber="302" stateProvince="Nantou">Fuhacho Village</location>
(=
<collectingMunicipality id="5D430034FF83FFDE216EF80AFA29ED61" box="[1350,1420,2031,2052]" pageId="11" pageNumber="302">Maobu</collectingMunicipality>
or
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,
<collectingRegion id="7F5C54ACFF83FFDE2795F7E9FBFFED44" box="[957,1114,2060,2081]" country="Taiwan" name="Nantou" pageId="11" pageNumber="302">Nantou County</collectingRegion>
), altitude about
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<elevation id="36B57D7DFF83FFDE2129F7E9FAC2ED44" box="[1281,1383,2060,2081]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="11" pageNumber="302" unit="ft" value="4000.0">4,000 feet</elevation>
</quantity>
(ca. 1
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<elevation id="36B57D7DFF83FFDE2767F7CCFC35ED5B" box="[847,912,2089,2110]" metricMagnitude="2" metricUnit="m" metricValue="2.19" pageId="11" pageNumber="302" unit="m" value="219.0">219 m</elevation>
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),
<collectingCountry id="C58FDADEFF83FFDE278CF7CCFC54ED5B" box="[932,1009,2089,2110]" name="Taiwan" pageId="11" pageNumber="302">Taiwan</collectingCountry>
.]
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<emphasis id="8FEC465CFF84FFD924AFFF58FEC6E5B7" bold="true" box="[135,355,189,210]" italics="true" pageId="12" pageNumber="303">Material examined. </emphasis>
<materialsCitation id="0DF09013FF84FFD92544FF58FE7FE420" accessionNumber="A 11618, CAS 19693, A 68242" country="Taiwan" county="Fushan Nature Reserve" location="Yang Ming Mountain" municipality="Horisha" pageId="12" pageNumber="303" specimenCount="1" stateProvince="Pingtung">
<collectingCountry id="C58FDADEFF84FFD92544FF58FE1CE5B7" box="[364,441,189,210]" name="Taiwan" pageId="12" pageNumber="303">Taiwan</collectingCountry>
:
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(NMNS 1512-1, 1512-3 to 11, 1512-14 to 23),
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(AMNH A 11618- 623),
<collectingRegion id="7F5C54ACFF84FFD92494FF12FEB7E469" box="[188,274,247,268]" country="Taiwan" name="Pingtung" pageId="12" pageNumber="303">Pingtung</collectingRegion>
(MCZ A-125372), Polisia Forosa (CAS 19693696), Tai-Chung Hsien (Huan-Shan) (AMNH
<accessionNumber id="A2CB07ADFF84FFD9262DFEF6FDFFE44D" box="[517,602,275,296]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A68242" pageId="12" pageNumber="303" type="EnaNcbi">A 68242</accessionNumber>
243),
<location id="B847CC95FF84FFD926B1FEF6FF41E420" LSID="urn:lsid:plazi:treatment:35312B58FF83FFD9278BF871FE55E0E7:B847CC95FF84FFD926B1FEF6FF41E420" country="Taiwan" county="Fushan Nature Reserve" municipality="Horisha" name="Yang Ming Mountain" pageId="12" pageNumber="303" stateProvince="Pingtung">Yang Ming Mountain</location>
(CAS-SUA 20485487)
</materialsCitation>
.
</paragraph>
<paragraph id="BD279A4EFF84FFD924AFFE8EFE0BE667" blockId="12.[135,774,363,770]" pageId="12" pageNumber="303">
<emphasis id="8FEC465CFF84FFD924AFFE8EFDF0E4E7" box="[135,597,363,387]" italics="true" pageId="12" pageNumber="303">
<emphasis id="8FEC465CFF84FFD924AFFE8EFED7E4E6" bold="true" box="[135,370,363,387]" italics="true" pageId="12" pageNumber="303">Taxonomic notes. </emphasis>
<taxonomicName id="7A98E1CDFF84FFD92556FE8EFDF0E4E7" authorityName="BOULENGER" authorityYear="1909" box="[382,597,363,386]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="adenopleura">Rana adenopleura</taxonomicName>
</emphasis>
was described by
<bibRefCitation id="D909E7BFFF84FFD92485FE6EFEDDE4C7" author="Boulenger, G" box="[173,376,395,418]" pageId="12" pageNumber="303" pagination="492 - 495" refId="ref17936" refString="Boulenger, G. A., 1909. Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa. Annals and Magazine of Natural History, series 8: 492 - 495." type="book chapter" year="1909">Boulenger (1909)</bibRefCitation>
from
<specimenCount id="AB9E51C7FF84FFD925EDFE6EFDD1E4C7" box="[453,628,395,418]" pageId="12" pageNumber="303" type="generic">four specimens</specimenCount>
collected at “Fuhacho”,
<collectingCountry id="C58FDADEFF84FFD92539FE4EFECDE4A7" box="[273,360,427,450]" name="Taiwan" pageId="12" pageNumber="303">Taiwan</collectingCountry>
. Several authors regarded the frogs from continental
<collectingCountry id="C58FDADEFF84FFD92560FE2EFE2CE487" box="[328,393,459,482]" name="China" pageId="12" pageNumber="303">China</collectingCountry>
belonging to
<taxonomicName id="7A98E1CDFF84FFD92601FE2EFD71E487" authorityName="BOULENGER" authorityYear="1909" box="[553,724,459,482]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="adenopleura">
<emphasis id="8FEC465CFF84FFD92601FE2EFD71E487" box="[553,724,459,482]" italics="true" pageId="12" pageNumber="303">R. adenopleura</emphasis>
</taxonomicName>
and regarded the congeneric species from Fukien namely
<taxonomicName id="7A98E1CDFF84FFD926C6FE0EFF4CE747" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="caldwelli">
<emphasis id="8FEC465CFF84FFD926C6FE0EFF4CE747" italics="true" pageId="12" pageNumber="303">R. caldwelli</emphasis>
</taxonomicName>
as a synonym (
<bibRefCitation id="D909E7BFFF84FFD925BAFDEEFDAAE747" author="Pope, C" box="[402,527,523,546]" pageId="12" pageNumber="303" pagination="397 - 611" refId="ref20440" refString="Pope, C. H., 1931. Notes on amphibians from Fukien, Hainan, and other parts of China. Bulletin of American Museum of Natural History, 61: 397 - 611." type="journal article" year="1931">Pope, 1931</bibRefCitation>
;
<bibRefCitation id="D909E7BFFF84FFD92633FDEEFD22E747" author="Liu, C" box="[539,647,523,546]" pageId="12" pageNumber="303" pagination="1 - 400" refId="ref19586" refString="Liu, C. - C., 1950. Amphibians of western China. Fieldiana: Zoololgy Memoirs, 2: 1 - 400." type="journal article" year="1950">Liu, 1950</bibRefCitation>
;
<bibRefCitation id="D909E7BFFF84FFD926BAFDEEFF1AE727" author="Kuramoto, M" pageId="12" pageNumber="303" pagination="150 - 158" refId="ref19376" refString="Kuramoto, M., 1985. A new frog (genus Rana) from the Yaeyama group of the Ryukyu Ialands. Herpetologica, 41 (2): 150 - 158." type="journal article" year="1985">Kuramoto, 1985</bibRefCitation>
;
<bibRefCitation id="D909E7BFFF84FFD924E4FDCEFEE9E727" author="Chou, W" box="[204,332,555,578]" pageId="12" pageNumber="303" pagination="389 - 400" refId="ref18476" refString="Chou, W. - H., 1999. A new frog of the genus Rana (Anura: Ranidae) from China. Herpetologica, 55 (3): 389 - 400." type="journal article" year="1999">Chou, 1999</bibRefCitation>
;
<bibRefCitation id="D909E7BFFF84FFD92571FDCEFDBBE727" author="Orlov, N &amp; Murphy, N &amp; Ananjeva, S" box="[345,542,555,578]" pageId="12" pageNumber="303" pagination="81 - 104" refId="ref20356" refString="Orlov, N. L., R. W. Murphy, N. B. Ananjeva, S. A. Ryabov &amp; T. C. Ho, 2002. Herpetofauna of Vietnam, a checklist. Part I. Amphibia. Russian Journal of Herpetology, 9 (2): 81 - 104." type="journal article" year="2002">Orlov et al., 2002</bibRefCitation>
). Based on published evidences,
<bibRefCitation id="D909E7BFFF84FFD924D7FDAEFE3EE707" author="Dubois, A" box="[255,411,587,610]" pageId="12" pageNumber="303" pagination="305 - 352" refId="ref18802" refString="Dubois, A., 1992. Notes sur la classification des Ranidae (Amphibiens Anoures). Bulletin Mensuel de la Societe Linneenne de Lyon, 61 (10): 305 - 352." type="journal article" year="1992">Dubois (1992)</bibRefCitation>
proposed to keep the name of
<taxonomicName id="7A98E1CDFF84FFD926C6FDAEFEABE7E7" authorityName="BOULENGER" authorityYear="1909" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="adenopleura">
<emphasis id="8FEC465CFF84FFD926C6FDAEFEABE7E7" italics="true" pageId="12" pageNumber="303">R. adenopleura</emphasis>
</taxonomicName>
for
<collectingCountry id="C58FDADEFF84FFD92513FD8EFE2EE7E7" box="[315,395,619,642]" name="Taiwan" pageId="12" pageNumber="303">Taiwan</collectingCountry>
population and restricted the name of
<taxonomicName id="7A98E1CDFF84FFD9248EFD6EFE8DE7C7" box="[166,296,651,674]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="caldwelli">
<emphasis id="8FEC465CFF84FFD9248EFD6EFE8DE7C7" box="[166,296,651,674]" italics="true" pageId="12" pageNumber="303">R. caldwelli</emphasis>
</taxonomicName>
for the Fujian population. As given above, the morphological and morphometrical comparisons confirm the differences between these two species as similar as the proposal of
<bibRefCitation id="D909E7BFFF84FFD92522FD0EFE0FE667" author="Dubois, A" box="[266,426,747,770]" pageId="12" pageNumber="303" pagination="305 - 352" refId="ref18802" refString="Dubois, A., 1992. Notes sur la classification des Ranidae (Amphibiens Anoures). Bulletin Mensuel de la Societe Linneenne de Lyon, 61 (10): 305 - 352." type="journal article" year="1992">Dubois (1992)</bibRefCitation>
.
</paragraph>
<paragraph id="BD279A4EFF84FFD924AFFCCEFDF4E027" blockId="12.[135,774,811,1346]" pageId="12" pageNumber="303">
<emphasis id="8FEC465CFF84FFD924AFFCCEFDFFE627" box="[135,602,811,835]" italics="true" pageId="12" pageNumber="303">
<emphasis id="8FEC465CFF84FFD924AFFCCEFE2AE626" bold="true" box="[135,399,811,835]" italics="true" pageId="12" pageNumber="303">Diagnostic characters. </emphasis>
<taxonomicName id="7A98E1CDFF84FFD925BDFCCEFDFFE627" authorityName="BOULENGER" authorityYear="1909" box="[405,602,811,834]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="adenopleura">Rana adenopleura</taxonomicName>
</emphasis>
is characterized by the following combination of characters: (1) body elongated; (2) SVL of adult males 48.5 ± 2.7 (
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), adult females 49.9 ± 1.9 (
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); (3) nostril directed laterally; (4) marginal grooves on fingers present or absent; (5) marginal grooves on toes present; (6) mid-dorsal stripe present or absent; (7) external vocal sac present; (8) nuptial pad present on thumb and first finger, forming a single pad; (9) suprabrachial gland present and large; (10) nest construction behavior absent; (11) spinules present on entire dorsum of head and body (in adult males); (12) spinules on ventrum present in adult males; (13) tibio-tarsal articulation reach snout tip or between eye to snout; (14) finger tips dilated; (15) toe tips dilated; (16) relative length of fingers II&lt;I&lt;IV&lt;III; (17) relative length of toes I&lt;II&lt;III=V&lt;IV; (18) pigmented eggs; (19) larval keratodont formula 1:1+1/1+1:2; (20) Calling: 24 loud “gai-gai-…” notes.
</paragraph>
</subSubSection>
<subSubSection id="F582C9C5FF84FFD924AFFA8EFE55E0E7" box="[135,496,1387,1411]" pageId="12" pageNumber="303" type="distribution">
<paragraph id="BD279A4EFF84FFD924AFFA8EFE55E0E7" blockId="12.[135,496,1387,1411]" box="[135,496,1387,1411]" pageId="12" pageNumber="303">
<emphasis id="8FEC465CFF84FFD924AFFA8EFE93E0E6" bold="true" box="[135,310,1387,1411]" pageId="12" pageNumber="303">
<emphasis id="8FEC465CFF84FFD924AFFA8EFEB7E0E6" bold="true" box="[135,274,1387,1411]" italics="true" pageId="12" pageNumber="303">Distribution.</emphasis>
</emphasis>
<collectingCountry id="C58FDADEFF84FFD92517FA8EFEDAE0E7" box="[319,383,1387,1410]" name="China" pageId="12" pageNumber="303">China</collectingCountry>
(
<collectingRegion id="7F5C54ACFF84FFD925A7FA8EFE49E0E7" box="[399,492,1387,1410]" country="China" name="Taiwan)" pageId="12" pageNumber="303">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>