250 lines
28 KiB
XML
250 lines
28 KiB
XML
<document id="1D490DDF4DDFC0F9BF6EE41C0912C83D" ID-DOI="10.1206/0003-0090(2002)273<0001:MAROAA>2.0.CO;2" ID-ISSN="0003-0090" ID-Zenodo-Dep="5397639" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630326589637" checkinUser="felipe" docAuthor="ASHER, ROBERT J., McKENNA, MALCOLM C., EMRY, ROBERT J., TABRUM, ALAN R. & KRON, DONALD G." docDate="2002" docId="001AB62EFFAFFF95FD256FF687EDF9A4" docLanguage="en" docName="B273.pdf" docOrigin="Bulletin of the American Museum of Natural History 2002 (273)" docSource="http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0090(2002)273%3C0001%3AMAROAA%3E2.0.CO%3B2" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.3:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="3" docTitle="Apternodus dasophylakas ASHER & McKENNA & EMRY & TABRUM & KRON 2002, new species" docType="treatment" docVersion="3" lastPageNumber="58" masterDocId="FC23CE56FF9AFFACFFFD6E458709FFE3" masterDocTitle="Morphology And Relationships Of Apternodus And Other Extinct, Zalambdodont, Placental Mammals" masterLastPageNumber="117" masterPageNumber="1" pageNumber="54" updateTime="1699228884684" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D29C46EDEBA6F317D4598979539C8946">Morphology And Relationships Of Apternodus And Other Extinct, Zalambdodont, Placental Mammals</mods:title>
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<mods:namePart id="87A7E052D659BB4D312A54141E84FEF7">ASHER, ROBERT J.</mods:namePart>
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<mods:namePart id="B867F0813203BD262CEF01DD4660EF50">McKENNA, MALCOLM C.</mods:namePart>
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<mods:namePart id="CD26C9259EBC9C271706AE762741D6E4">EMRY, ROBERT J.</mods:namePart>
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<mods:namePart id="F57E02FC20805C3D803F82A6EFA7A5E6">TABRUM, ALAN R.</mods:namePart>
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<mods:namePart id="AC5E7EAF57F93F24BCFFA8D7546FCA2E">KRON, DONALD G.</mods:namePart>
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<mods:title id="6506DAAF8C5C9ED52063068F25C80649">Bulletin of the American Museum of Natural History</mods:title>
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<mods:date id="5A1328EAEC0D409A068A44074B6BB933">2002</mods:date>
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<mods:number id="E10EDE0E063144ED705F3AE94525957D">2002-11-27</mods:number>
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<mods:number id="0ADB3F3307DEEE6D759A374B7DB98917">2002</mods:number>
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<mods:number id="AF6658573B29BF1712E362107C2D1CD8">273</mods:number>
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<mods:identifier id="D571DF7C26251DE7462831A98380C424" type="DOI">10.1206/0003-0090(2002)273<0001:MAROAA>2.0.CO;2</mods:identifier>
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<mods:identifier id="95EFE52B27D528A904B68CF83726044A" type="Zenodo-Dep">5397639</mods:identifier>
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<treatment id="001AB62EFFAFFF95FD256FF687EDF9A4" LSID="urn:lsid:plazi:treatment:001AB62EFFAFFF95FD256FF687EDF9A4" httpUri="http://treatment.plazi.org/id/001AB62EFFAFFF95FD256FF687EDF9A4" lastPageId="57" lastPageNumber="58" pageId="53" pageNumber="54">
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<subSubSection id="C0A954B3FFAFFF99FD256FF683A7FE29" box="[728,1198,435,459]" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="880C0738FFAFFF99FD256FF683A7FE29" blockId="53.[728,1198,435,459]" box="[728,1198,435,459]" pageId="53" pageNumber="54">
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<heading id="D344B054FFAFFF99FD256FF683A7FE29" box="[728,1198,435,459]" centered="true" fontSize="9" level="2" pageId="53" pageNumber="54" reason="2">
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<taxonomicName id="4FB37CBBFFAFFF99FD256FF68307FE28" authority="ASHER & McKENNA & EMRY & TABRUM & KRON, 2002" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" box="[728,1038,435,459]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="dasophylakas" status="sp. nov.">
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<emphasis id="BAC7DB2AFFAFFF99FD256FF68307FE28" bold="true" box="[728,1038,435,459]" italics="true" pageId="53" pageNumber="54">Apternodus dasophylakas</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A1F46651FFAFFF99FBE26FF683A7FE29" box="[1055,1198,435,458]" pageId="53" pageNumber="54" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C0A954B3FFAFFF99FD256FA384ECFCC0" pageId="53" pageNumber="54" type="materials_examined">
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<paragraph id="880C0738FFAFFF99FD256FA384ECFCC0" blockId="53.[699,1228,486,1741]" pageId="53" pageNumber="54">
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<typeStatus id="5708B99AFFAFFF99FD256FA38418FE1E" box="[728,785,486,509]" pageId="53" pageNumber="54">TYPE</typeStatus>
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AND ONLY SPECIMEN:
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<collectionCode id="EEA29FFDFFAFFF99FBC76FA38363FE1E" box="[1082,1130,486,509]" country="USA" name="University of Washington Fish Collection" pageId="53" pageNumber="54" type="Museum">UW</collectionCode>
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14072, crushed but otherwise complete skull (fig. 35) preserving left I1, CM3, right I1, broken C, P2, broken P3, P4M3, articulated left dentary with complete dentition, associated right dentary with i1i2, cm3 (fig. 36), associated tibia, atlas, axis, partial scapula, proximal humerus, distal ulna, ribs, and other fragmentary postcranial fragments (fig. 37). All elements are slightly distorted due to postmortem deformation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C0A954B3FFAFFF99FD256D6C8347FB38" pageId="53" pageNumber="54" type="etymology">
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<paragraph id="880C0738FFAFFF99FD256D6C8430FB60" blockId="53.[699,1228,486,1741]" pageId="53" pageNumber="54">
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ETYMOLOGY: This species is named in hon or of the faculty, students, and staff of the Department of Geology and Geophysics, University of
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<collectingRegion id="4A77C9DAFFAFFF99FC976DC484E9FC7B" box="[874,992,897,920]" country="United States of America" name="Wyoming" pageId="53" pageNumber="54">Wyoming</collectingRegion>
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, under whose auspices the
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<typeStatus id="5708B99AFFAFFF99FCC96DDB846FFC56" box="[820,870,926,949]" pageId="53" pageNumber="54">type</typeStatus>
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and only specimen was collected and is now curated. The Greek word transliterated as ‘‘
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<taxonomicName id="4FB37CBBFFAFFF99FC6C6D9C8326FC13" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" box="[913,1071,985,1008]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="dasophylakas">
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<emphasis id="BAC7DB2AFFAFFF99FC6C6D9C8326FC13" box="[913,1071,985,1008]" italics="true" pageId="53" pageNumber="54">dasophylakas</emphasis>
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</taxonomicName>
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’’ is an interpretation of the English word ‘‘cowboy’’, the UW mascot. Literally, the trivial name means ‘‘ranger’’ or ‘‘guardian of wild land and forest’’ (N. Soulunias and I. Trivilas, personal commun.).
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</paragraph>
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<paragraph id="880C0738FFAFFF99FD256ACC8347FB38" blockId="53.[699,1228,486,1741]" pageId="53" pageNumber="54">
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TEMPORAL AND GEOGRAPHIC DISTRIBUTION: Chadronian (late Eocene) of southeastern
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<collectingRegion id="4A77C9DAFFAFFF99FD466A818427FB38" box="[699,814,1220,1243]" country="United States of America" name="Wyoming" pageId="53" pageNumber="54">Wyoming</collectingRegion>
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(Harshman Quarry 21).
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</paragraph>
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</subSubSection>
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<subSubSection id="C0A954B3FFAFFF9BFD256AA4852BF92E" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="diagnosis">
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<paragraph id="880C0738FFAFFF9BFD256AA4852BF92E" blockId="53.[699,1228,486,1741]" lastBlockId="55.[123,652,1486,1741]" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
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DIAGNOSIS: Protocones on the upper molars are absent, a condition somewhat exacerbated by wear. The anterior dentition is premolariform, not bulbous. P3 has an elongate stylar margin, and diastemata are evident on either side of the upper canine, between I1 and I2, and probably also between P2 and P3; some distortion in the maxilla between P2P3 on both sides precludes certainty as to the size of the P2P3 diastema. Both P2 and p2 are smaller than adjacent teeth and are similar in proportion to those of
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<taxonomicName id="4FB37CBBFFAFFF99FD276804848AF9BB" authorityName="Schlaikjer" authorityYear="1934" box="[730,899,1601,1624]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="brevirostris">
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<emphasis id="BAC7DB2AFFAFFF99FD276804848AF9BB" box="[730,899,1601,1624]" italics="true" pageId="53" pageNumber="54">A. brevirostris</emphasis>
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</taxonomicName>
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. The mental foramen on the dentary is located just below p2, a position slightly anterior to that of other
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<taxonomicName id="4FB37CBBFFAFFF99FBB86839841FF953" class="Mammalia" family="Apternodontidae" genus="Apternodus" kingdom="Animalia" order="Soricomorpha" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="BAC7DB2AFFAFFF99FBB8683983C2F970" box="[1093,1227,1660,1683]" italics="true" pageId="53" pageNumber="54">Apternodus</emphasis>
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species.
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</taxonomicName>
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The p3 shows a slight buccal cingulid and a distinct posterior cusp. The ex ternal auditory meatus of UW 14072 is not marked by a torus, in contrast to that of
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<taxonomicName id="4FB37CBBFFADFF9BFD8F6BA987C9F9C3" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="major">
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<emphasis id="BAC7DB2AFFADFF9BFD8F6BA987C9F9C3" italics="true" pageId="55" pageNumber="56">A. major</emphasis>
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</taxonomicName>
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, but is flat as in
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<taxonomicName id="4FB37CBBFFADFF9BFE69684C8513F9C3" box="[404,538,1545,1568]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="iliffensis">
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<emphasis id="BAC7DB2AFFADFF9BFE69684C8513F9C3" box="[404,538,1545,1568]" italics="true" pageId="55" pageNumber="56">A. iliffensis</emphasis>
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</taxonomicName>
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. Another similarity to
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<taxonomicName id="4FB37CBBFFADFF9BFEED6863869DF9DE" box="[272,404,1574,1597]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="iliffensis">
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<emphasis id="BAC7DB2AFFADFF9BFEED6863869DF9DE" box="[272,404,1574,1597]" italics="true" pageId="55" pageNumber="56">A. iliffensis</emphasis>
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</taxonomicName>
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is the ventral margin of the lambdoid plate, which reaches just slightly ventral to the level of the jaw joint. The rostral tympanic process of the petrosal is flat. A piriform fenestra is evident, lateral to a small anterior carotid foramen.
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</paragraph>
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</subSubSection>
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<caption id="DCCC57B0FFACFF9AFF6B687585D3F983" ID-DOI="http://doi.org/10.5281/zenodo.5397738" ID-Zenodo-Dep="5397738" httpUri="https://zenodo.org/record/5397738/files/figure.png" pageId="54" pageNumber="55" startId="54.[150,188,1584,1605]" targetBox="[196,1156,187,1562]" targetPageId="54">
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<paragraph id="880C0738FFACFF9AFF6B687585D3F983" blockId="54.[124,1228,1584,1632]" pageId="54" pageNumber="55">
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Fig. 35. UW 14072, skull of
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<taxonomicName id="4FB37CBBFFACFF9AFE1D687585E6F9A6" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" box="[480,751,1584,1605]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="dasophylakas">
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<emphasis id="BAC7DB2AFFACFF9AFE1D687585E6F9A6" box="[480,751,1584,1605]" italics="true" pageId="54" pageNumber="55">Apternodus dasophylakas</emphasis>
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</taxonomicName>
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type specimen from Harshman Quarry, Wyoming in stereo ventral (top) and dorsal (bottom) views.
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</paragraph>
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</caption>
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<caption id="DCCC57B0FFADFF9BFF686B35848CFA43" ID-DOI="http://doi.org/10.5281/zenodo.5397742" ID-Zenodo-Dep="5397742" httpUri="https://zenodo.org/record/5397742/files/figure.png" pageId="55" pageNumber="56" startId="55.[149,187,1392,1413]" targetBox="[123,1228,187,1371]" targetPageId="55">
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<paragraph id="880C0738FFADFF9BFF686B3583C2FA66" blockId="55.[123,1227,1392,1440]" box="[149,1227,1392,1413]" pageId="55" pageNumber="56">
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Fig. 36. UW 14072, mandible of
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<taxonomicName id="4FB37CBBFFADFF9BFDED6B358429FA66" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" box="[528,800,1392,1413]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="dasophylakas">
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<emphasis id="BAC7DB2AFFADFF9BFDED6B358429FA66" box="[528,800,1392,1413]" italics="true" pageId="55" pageNumber="56">Apternodus dasophylakas</emphasis>
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</taxonomicName>
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type specimen from Harshman Quarry,
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</paragraph>
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<paragraph id="880C0738FFADFF9BFF866BCE848CFA43" blockId="55.[123,1227,1392,1440]" box="[123,901,1419,1440]" pageId="55" pageNumber="56">Wyoming in occlusal (top), lateral (middle), and lingual (bottom) views.</paragraph>
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</caption>
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<subSubSection id="C0A954B3FFADFF95FD256B8B856DFA22" lastPageId="57" lastPageNumber="58" pageId="55" pageNumber="56" type="discussion">
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<paragraph id="880C0738FFADFF95FD256B8B8577FC98" blockId="55.[699,1227,1486,1741]" lastBlockId="57.[123,652,193,1419]" lastPageId="57" lastPageNumber="58" pageId="55" pageNumber="56">
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REMARKS: UW 14072 is a large version of
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<taxonomicName id="4FB37CBBFFADFF9BFD466BA98437F9E0" box="[699,830,1516,1539]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="iliffensis">
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<emphasis id="BAC7DB2AFFADFF9BFD466BA98437F9E0" box="[699,830,1516,1539]" italics="true" pageId="55" pageNumber="56">A. iliffensis</emphasis>
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</taxonomicName>
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, sharing with it aspects of the ear and lambdoid region as well as reduced upper molar protocones and an elongate P3 (polymorphic in
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<taxonomicName id="4FB37CBBFFADFF9BFC826806830BF9B9" box="[895,1026,1603,1626]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="iliffensis">
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<emphasis id="BAC7DB2AFFADFF9BFC826806830BF9B9" box="[895,1026,1603,1626]" italics="true" pageId="55" pageNumber="56">A. iliffensis</emphasis>
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</taxonomicName>
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). UW 14072 differs qualitatively from individuals of
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<taxonomicName id="4FB37CBBFFADFF9BFB83682585F4F977" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="iliffensis">
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<emphasis id="BAC7DB2AFFADFF9BFB83682585F4F977" italics="true" pageId="55" pageNumber="56">A. iliffensis</emphasis>
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</taxonomicName>
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in having an M3 similar in buccolingual width to the M2, an elongate rostrum with diastemata on either side of the canine, and a transversely wide basicranium. It differs from other
|
||
<taxonomicName id="4FB37CBBFFA3FF95FECE6E9A851CFF16" box="[307,533,222,246]" class="Mammalia" family="Apternodontidae" genus="Apternodus" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="BAC7DB2AFFA3FF95FECE6E9A86B0FF15" box="[307,441,223,246]" italics="true" pageId="57" pageNumber="58">Apternodus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in having a mental foramen ventral to p2; the position of the mental foramen(ina) is unknown in
|
||
<taxonomicName id="4FB37CBBFFA3FF95FD8F6F5C87D7FEAD" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="iliffensis">
|
||
<emphasis id="BAC7DB2AFFA3FF95FD8F6F5C87D7FEAD" italics="true" pageId="57" pageNumber="58">A. iliffensis</emphasis>
|
||
</taxonomicName>
|
||
. Cranial proportions are similar to those of
|
||
<taxonomicName id="4FB37CBBFFA3FF95FF196F118663FE88" authorityName="Schlaikjer" authorityYear="1933" box="[228,362,340,363]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="gregoryi">
|
||
<emphasis id="BAC7DB2AFFA3FF95FF196F118663FE88" box="[228,362,340,363]" italics="true" pageId="57" pageNumber="58">A. gregoryi</emphasis>
|
||
</taxonomicName>
|
||
, with the exception that
|
||
<taxonomicName id="4FB37CBBFFA3FF95FF866F348632FE6B" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" box="[123,315,369,392]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="dasophylakas">
|
||
<emphasis id="BAC7DB2AFFA3FF95FF866F348632FE6B" box="[123,315,369,392]" italics="true" pageId="57" pageNumber="58">A. dasophylakas</emphasis>
|
||
</taxonomicName>
|
||
has a very shallow external auditory meatus and a lambdoid plate that does not extend far ventral to the jaw joint. The orbitotemporal region of UW 14072 is crushed, leaving a very distorted lacrimal foramen on the left side, and none on the right. It is therefore difficult to tell if the posterior border of the lacrimal foramen is flush with the anterior orbit as in
|
||
<taxonomicName id="4FB37CBBFFA3FF95FE6B6C198582FD90" authorityName="Schlaikjer" authorityYear="1933" box="[406,651,604,627]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="gregoryi">
|
||
<emphasis id="BAC7DB2AFFA3FF95FE6B6C198582FD90" box="[406,651,604,627]" italics="true" pageId="57" pageNumber="58">Apternodus gregoryi</emphasis>
|
||
</taxonomicName>
|
||
(figs. 21, 22) and
|
||
<taxonomicName id="4FB37CBBFFA3FF95FEB26C3C86BFFD73" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" box="[335,438,633,656]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="BAC7DB2AFFA3FF95FEB26C3C86BFFD73" box="[335,438,633,656]" italics="true" pageId="57" pageNumber="58">A. major</emphasis>
|
||
</taxonomicName>
|
||
(fig. 32). The left craniomandibular joint, with the jaw still in articulation, preserves what might be part of an ossified articular disk separating the condyle from the glenoid fossa (fig. 35). However, due to the postmortem distortion of the specimen, it is difficult to discount the possibility that this region contains multiple fragments of the mandibular condyle itself.
|
||
</paragraph>
|
||
<caption id="DCCC57B0FFA2FF94FF6B68768393F981" ID-DOI="http://doi.org/10.5281/zenodo.5397744" ID-Zenodo-Dep="5397744" httpUri="https://zenodo.org/record/5397744/files/figure.png" pageId="56" pageNumber="57" startId="56.[150,188,1587,1608]" targetBox="[164,1188,187,1566]" targetPageId="56">
|
||
<paragraph id="880C0738FFA2FF94FF6B68768393F981" blockId="56.[124,1228,1587,1635]" pageId="56" pageNumber="57">
|
||
Fig. 37. UW 14072, scapula, tibia, fragmentary vertebral column including the axis (top and middle) and atlas (bottom) of
|
||
<taxonomicName id="4FB37CBBFFA2FF94FE9A680B857FF980" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" box="[359,630,1614,1635]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="dasophylakas">
|
||
<emphasis id="BAC7DB2AFFA2FF94FE9A680B857FF980" box="[359,630,1614,1635]" italics="true" pageId="56" pageNumber="57">Apternodus dasophylakas</emphasis>
|
||
</taxonomicName>
|
||
type specimen from Harshman Quarry, Wyoming.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="880C0738FFA3FF95FF656DC487B2FA68" blockId="57.[123,652,193,1419]" pageId="57" pageNumber="58">
|
||
Postcranially, what little is known of
|
||
<taxonomicName id="4FB37CBBFFA3FF95FD8F6DC48610FC55" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="dasophylakas">
|
||
<emphasis id="BAC7DB2AFFA3FF95FD8F6DC48610FC55" italics="true" pageId="57" pageNumber="58">A. dasophylakas</emphasis>
|
||
</taxonomicName>
|
||
(fig. 37) does not appear to be specialized for a nonterrestrial locomotor repertoire. The only exception to this observation in any
|
||
<taxonomicName id="4FB37CBBFFA3FF95FEDA6DB286A4FBED" authorityName="Matthew" authorityYear="1903" box="[295,429,1015,1038]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="genus">
|
||
<emphasis id="BAC7DB2AFFA3FF95FEDA6DB286A4FBED" box="[295,429,1015,1038]" italics="true" pageId="57" pageNumber="58">Apternodus</emphasis>
|
||
</taxonomicName>
|
||
specimen may be the structure of the proximal femur in
|
||
<taxonomicName id="4FB37CBBFFA3FF95FDC76A5187B1FBAB" authorityName="Schlaikjer" authorityYear="1933" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="gregoryi">
|
||
<emphasis id="BAC7DB2AFFA3FF95FDC76A5187B1FBAB" italics="true" pageId="57" pageNumber="58">A. gregoryi</emphasis>
|
||
</taxonomicName>
|
||
(UW 13508; see above). The distal tibia of
|
||
<taxonomicName id="4FB37CBBFFA3FF95FF426A0A868BFB85" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" box="[191,386,1103,1126]" class="Mammalia" family="Apternodontidae" genus="Apternodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="dasophylakas">
|
||
<emphasis id="BAC7DB2AFFA3FF95FF426A0A868BFB85" box="[191,386,1103,1126]" italics="true" pageId="57" pageNumber="58">A. dasophylakas</emphasis>
|
||
</taxonomicName>
|
||
is not fused with the fibula. The scapula has a triangular shape, narrow laterally at the glenoid fossa and broadening dorsomedially toward the vertebral column. The second cervical vertebra shows a ventrally projecting keel on the posteroventral aspect of its centrum (similar to the axis of shrews and
|
||
<taxonomicName id="4FB37CBBFFA3FF95FE776B598512FAD0" authorityName="Blainville" authorityYear="1838" box="[394,539,1308,1331]" class="Mammalia" family="Erinaceidae" genus="Echinosorex" kingdom="Animalia" order="Erinaceomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="genus">
|
||
<emphasis id="BAC7DB2AFFA3FF95FE776B598512FAD0" box="[394,539,1308,1331]" italics="true" pageId="57" pageNumber="58">Echinosorex</emphasis>
|
||
</taxonomicName>
|
||
); and the first cervical vertebra shows two widely separated facets for the occipital condyles of the skull.
|
||
</paragraph>
|
||
<paragraph id="880C0738FFA3FF95FF666BEB856DFA22" blockId="57.[155,612,1454,1473]" box="[155,612,1454,1473]" pageId="57" pageNumber="58">
|
||
FAMILY
|
||
<emphasis id="BAC7DB2AFFA3FF95FF046BEB86C1FA22" bold="true" box="[249,456,1454,1473]" pageId="57" pageNumber="58">
|
||
<taxonomicName id="4FB37CBBFFA3FF95FF046BEB86CCFA22" authority="ASHER & McKENNA & EMRY & TABRUM & KRON, 2002" authorityName="ASHER & McKENNA & EMRY & TABRUM & KRON" authorityYear="2002" box="[249,453,1454,1473]" class="Insecta" family="Oligoryctidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="family" status="fam. nov.">OLIGORYCTIDAE</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A1F46651FFA3FF95FE2D6BEB856DFA22" box="[464,612,1454,1473]" pageId="57" pageNumber="58" rank="family">NEW FAMILY</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0A954B3FFA3FF95FF656B9D87BCF9EF" pageId="57" pageNumber="58" type="description">
|
||
<paragraph id="880C0738FFA3FF95FF656B9D87BCF9EF" blockId="57.[123,651,1496,1607]" pageId="57" pageNumber="58">
|
||
INCLUDED GENERA:
|
||
<taxonomicName id="4FB37CBBFFA3FF95FE6E6B9D87BCF9EF" authority="Hough, 1956" authorityName="Hough" authorityYear="1956" class="Mammalia" family="Apternodontidae" genus="Oligoryctes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="genus">
|
||
<emphasis id="BAC7DB2AFFA3FF95FE6E6B9D852AFA0C" box="[403,547,1496,1519]" italics="true" pageId="57" pageNumber="58">Oligoryctes</emphasis>
|
||
<bibRefCitation id="EC227AC9FFA3FF95FDCD6B9D87BCF9EF" author="Hough, M. J." pageId="57" pageNumber="58" pagination="531 - 541" refId="ref55684" refString="Hough, M. J. 1956. A new insectivore from the Oligocene of the Wind River Basin, Wyoming, with notes on the taxonomy of the Oligocene Tenrecoidea. Journal of Paleontology 30: 531 - 541." type="journal article" year="1956">Hough, 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0A954B3FFA3FF95FF65685787EDF9A4" pageId="57" pageNumber="58" type="distribution">
|
||
<paragraph id="880C0738FFA3FF95FF65685787EDF9A4" blockId="57.[123,651,1496,1607]" pageId="57" pageNumber="58">
|
||
DISTRIBUTION AND DIAGNOSIS: As for
|
||
<taxonomicName id="4FB37CBBFFA3FF95FDA1685687D4F9A4" authorityName="Hough" authorityYear="1956" class="Mammalia" family="Apternodontidae" genus="Oligoryctes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="58" phylum="Chordata" rank="genus">
|
||
<emphasis id="BAC7DB2AFFA3FF95FDA1685687D4F9A4" italics="true" pageId="57" pageNumber="58">Oligoryctes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |