treatments-xml/data/C9/F8/FF/C9F8FF178AC75B21A3CA9EEF8A6AFD13.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.889.39090" ID-GBIF-Dataset="f50f6008-500c-492f-84b8-295ba991ec7b" ID-PMC="PMC6872603" ID-Pensoft-Pub="1313-2970-889-81" ID-Pensoft-UUID="0C546DE0BA425539879C1421D8C25A63" ID-PubMed="31777439" ID-ZooBank="7E7A3D196D7040398C087B248A27EB33" ModsDocID="1313-2970-889-81" checkinTime="1574358895821" checkinUser="pensoft" docAuthor="Hendrich, Lars, Shaverdo, Helena, Hajek, Jiri &amp; Balke, Michael" docDate="2019" docId="C9F8FF178AC75B21A3CA9EEF8A6AFD13" docLanguage="en" docName="ZooKeys 889: 81-152" docOrigin="ZooKeys 889" docSource="http://dx.doi.org/10.3897/zookeys.889.39090" docTitle="Copelatus bakewelli J. Balfour-Browne 1939" docType="treatment" docVersion="4" id="0C546DE0BA425539879C1421D8C25A63" lastPageNumber="81" masterDocId="0C546DE0BA425539879C1421D8C25A63" masterDocTitle="Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="152" masterPageNumber="81" pageNumber="81" updateTime="1668168070667" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hendrich, Lars</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hajek, Jiri</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Balke, Michael</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2019</mods:date>
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<mods:number>889</mods:number>
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<mods:start>81</mods:start>
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<treatment ID-GBIF-Taxon="160511455" LSID="urn:lsid:plazi:treatment:C9F8FF178AC75B21A3CA9EEF8A6AFD13" httpUri="http://treatment.plazi.org/id/C9F8FF178AC75B21A3CA9EEF8A6AFD13" lastPageNumber="81" pageId="0" pageNumber="81">
<subSubSection pageId="0" pageNumber="81" type="nomenclature">
<paragraph pageId="0" pageNumber="81">
<taxonomicName LSID="C9F8FF17-8AC7-5B21-A3CA-9EEF8A6AFD13" authority="J. Balfour-Browne, 1939" authorityName="J. Balfour-Browne" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">Copelatus bakewelli J. Balfour-Browne, 1939</taxonomicName>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Figures 1</figureCitation>
, 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Copelatus bakewelli A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure11" httpUri="https://binary.pensoft.net/fig/356480" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure11">11</figureCitation>
, 
<figureCitation captionStart="Figures 20, 21" captionStartId="F20" captionText="Figures 20, 21. Distribution of 20 Copelatus bakewelli 21 C. daemeli." figureDoi="10.3897/zookeys.889.39090.figures20-21" httpUri="https://binary.pensoft.net/fig/356489" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figures20-21">20</figureCitation>
, 
<figureCitation captionStart="Figure 30" captionStartId="F28" captionText="Figure 30. Habitat of Copelatus bakewelli, rocky pool, with rotten leaves near Gunlom Waterfall, Northern Territory. Photographs by L. Hendrich." figureDoi="10.3897/zookeys.889.39090.figure30" httpUri="https://binary.pensoft.net/fig/356497" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure30">30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="reference_group">
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">Copelatus bakewelli</taxonomicName>
J. 
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" author="Balfour-Browne, J" journalOrPublisher="The Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="81" pagination="57 - 88" refId="B1" refString="Balfour-Browne, J, 1939. On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with descriptions of new species. The Transactions of the Royal Entomological Society of London 88: 57 - 88" title="On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with descriptions of new species." url="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" volume="88" year="1939">Balfour-Browne 1939</bibRefCitation>
: 86 (original description); 
<bibRefCitation DOI="https://doi.org/10.1071/AJZS057" author="Watts, CHS" journalOrPublisher="Australian Journal of Zoology Supplement Series" pageId="0" pageNumber="81" pagination="1 - 166" refId="B36" refString="Watts, CHS, 1978. A revision of the Australian Dytiscidae (Coleoptera). Australian Journal of Zoology Supplement Series 57: 1 - 166" title="A revision of the Australian Dytiscidae (Coleoptera)." url="https://doi.org/10.1071/AJZS057" volume="57" year="1978">Watts 1978</bibRefCitation>
: 127 (description); 
<bibRefCitation author="Watts, CHS" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="81" pagination="22 - 28" refId="B37" refString="Watts, CHS, 1985. A faunal assessment of Australian Hydradephaga. Proceedings of the Academy of Natural Sciences of Philadelphia 137: 22 - 28" title="A faunal assessment of Australian Hydradephaga." volume="137" year="1985">Watts 1985</bibRefCitation>
: 26 (general distribution); 
<bibRefCitation author="Watts, CHS" journalOrPublisher="Identification Ecology Guide" pageId="0" pageNumber="81" pagination="1 - 110" refId="B38" refString="Watts, CHS, 2002. Checklist and guides to the identification, to genus, of adults and larval Australian water beetles of the families Dytiscidae, Noteridae, Hygrobiidae, Haliplidae, Gyrinidae, Hydraenidae and the superfamily Hydrophiloidea (Insecta-Coleoptera). Cooperative Research Centre for Freshwater Ecology (Australia). Identification Ecology Guide 43: 1 - 110" title="Checklist and guides to the identification, to genus, of adults and larval Australian water beetles of the families Dytiscidae, Noteridae, Hygrobiidae, Haliplidae, Gyrinidae, Hydraenidae and the superfamily Hydrophiloidea (Insecta-Coleoptera). Cooperative Research Centre for Freshwater Ecology (Australia)." volume="43" year="2002">Watts 2002</bibRefCitation>
: 42 (checklist); 
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="81" publicationUrl="http://www.waterbeetles.eu" refId="B28" refString="Nilsson, AN, Hajek, J, 2019. .http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2019">
Nilsson and 
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
2019
</bibRefCitation>
: 62 (catalogue).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type locality">
<paragraph pageId="0" pageNumber="81">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="81">&quot;North Australia&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type material">
<paragraph pageId="0" pageNumber="81">Type material.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Holotype:</emphasis>
Female, 
<normalizedToken originalValue="“Holotype”">&quot;Holotype&quot;</normalizedToken>
[round label with red frame], 
<normalizedToken originalValue="“Na”">&quot;Na&quot;</normalizedToken>
[hw label], 
<normalizedToken originalValue="“6756”">&quot;6756&quot;</normalizedToken>
[printed label], &quot; 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">Copelatus bakewelli</taxonomicName>
B-B. Type&quot; [hw label] (NHMUK).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="materials_examined">
<paragraph pageId="0" pageNumber="81">Additional material studied (285 specimens).</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Western Australia:</emphasis>
2 exs., &quot;AUSTRALIA WA, Synnot Range 1/9/1969, 
<geoCoordinate degrees="16" direction="south" minutes="25" orientation="latitude" precision="925" value="-16.416666">16°25'S</geoCoordinate>
, 
<geoCoordinate degrees="125" direction="east" minutes="28" orientation="longitude" precision="925" value="125.46667">125°28'E</geoCoordinate>
, Giuliani D.D. leg.&quot; (WAM); 1 ex., &quot;AUSTRALIA/WA/Shire of Wyndham - East Kimberley, Gibb River Road, Drysdale River Crossing, 380 m, 14.6.1999, Hendrich leg. Loc. 9/109&quot; (CLH); 4 exs., &quot;AUSTRALIA/WA/ Shire of Wyndham-East Kimberley, Gibb River Road, King Edward River Crossing, 280 m, 15.6.1999, Hendrich leg./coll. Loc. 10/110&quot; (CLH); 77 exs., &quot;AUSTRALIA/WA/Shire of Wyndham-East Kimberley, Mitchell Plateau, Little Mertens Falls, 300 m, 15.6.1999, Hendrich leg./coll. Loc. 11/111b&quot; (CLH, ZSM); 8 exs., &quot;AUSTRALIA/WA/ East Kimberley, Mitchell Plateau, Little Mertens Falls, 300 m, 15.6.1999, Hendrich leg./coll. Loc. 11a/111a&quot; (NMW); 2 exs., &quot;AUSTRALIA/WA/Shire of Wyndham - East Kimberley, Mitchell Plateau, Port Warrender Road/Kalumburu Road, Lowya Creek, 290 m, 18.6.1999, Hendrich leg. Loc. 13/113&quot; (CLH); 1 ex., &quot;AUSTRALIA/WA/Shire of Wyndham - East Kimberley, Mitchell Plateau, Kalumburu Road, 25 km NW King Edward Homestead, N.N. Creek, 370 m, 18.6.1999, Hendrich leg. Loc. 114&quot; (CLH); 4 exs., &quot;AUSTRALIA/WA/Shire of Wyndham - East Kimberley, Gibb River Road, 10 km E Hann River, Snake Creek, 470 m, 19.6.1999, Hendrich leg. Loc. 16/116&quot; (CLH). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Northern Territory</emphasis>
: 1 ex., &quot;Australia: NT, Cullen River n. Cullen, at Stuart Hwy, 102 m, 23.VIII.2006, 14.02.052S 131.56.561E, L. &amp; E. Hendrich leg. (NT 13)&quot;, &quot;DNA Balke 2213&quot; [green printed label] (ZSM); 1 ex., &quot;Australia: NT, Kakadu Hwy, Harriet Creek at Hwy Cross., 153 m, 24.VIII.2006, 13.44.512S 131.54.012E, L. &amp; E. Hendrich leg. (NT 14)&quot;, &quot;DNA Balke 1614&quot; [green printed label] (ZSM); 2 exs., &quot;Australia: NT, Litchfield NP, Creek near Wanggi Falls, 191 m, 20.VIII.2006, 13.11.221S 130.43.327E, L. &amp; E. Hendrich leg. (NT 4)&quot;, &quot;DNA Balke 2190&quot;, &quot;DNA Balke 2191&quot; [green printed label] (ZSM); 2 exs., &quot;AUSTRALIA NT Kakadu NP Alligator R. Gungaree Rainforest Dec. 22/93 S&amp;J Peck&quot; (ZSM); 1 ex., &quot;NT, Horse Creek fossil site, Camfield Station [
<geoCoordinate degrees="17.1166" direction="south" orientation="latitude" precision="5" value="-17.1166">-17.1166</geoCoordinate>
<geoCoordinate degrees="131.5166" direction="east" orientation="longitude" precision="5" value="131.5166">131.5166</geoCoordinate>
] 2/8/1992&quot;, 
<normalizedToken originalValue="“1004435”">&quot;1004435&quot;</normalizedToken>
(MAGNT); 1 ex., &quot;NT, Top Humbert Yard, Humbert River [
<geoCoordinate degrees="16.4333" direction="south" orientation="latitude" precision="5" value="-16.4333">-16.4333</geoCoordinate>
, 
<geoCoordinate degrees="130.4666" direction="east" orientation="longitude" precision="5" value="130.4666">130.4666</geoCoordinate>
] 7/1986&quot;, 
<normalizedToken originalValue="“1004443”">&quot;1004443&quot;</normalizedToken>
(MAGNT); 2 exs., &quot;Bathurst Is. + Melville Is. N.T. XII. 1975 D. Curry&quot;, &quot;QDPC 0-172712&quot;, &quot;QDPC 0-172713&quot; (QDPC); 2 exs., &quot;Australia, NT, Litchfield NP, Florence Falls, 
<geoCoordinate degrees="12" direction="south" minutes="51" orientation="latitude" precision="15" seconds="15" value="-12.854167">12°51'15&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="45" orientation="longitude" precision="15" seconds="16" value="132.75444">132°45'16&quot;E</geoCoordinate>
, 63 m, at light, 12.V.2006, leg. Berger &amp; Dostal (5/06)&quot; (CGW); 8 exs., &quot;Australia, NT, 80 km W Roper Bar, 
<geoCoordinate degrees="14" direction="south" minutes="54" orientation="latitude" precision="15" seconds="19" value="-14.905277">14°54'19&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="133" direction="east" minutes="57" orientation="longitude" precision="15" seconds="28" value="133.95778">133°57'28&quot;E</geoCoordinate>
, 78 m, at light, 14.V.2006, leg. Berger &amp; Dostal (7/06)&quot; (CGW); 1 ex., &quot;Australia, N.T./Katherine Gorge, Butterfly Gorge Walk, 150 m, 4.7.1999, Hendrich leg. Loc. 33/133&quot; (CLH); 3 exs., &quot;Australia, N.T. Kakadu N.P., Mary River District, 3 km ESE Gunlom Camping Area. South Alligator River, 50 m, 2.11.1996, 
<geoCoordinate degrees="13" direction="south" minutes="27.276" orientation="latitude" precision="1" value="-13.4546">13°27.276'S</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="26.268" orientation="longitude" precision="1" value="132.4378">132°26.268'E</geoCoordinate>
L. Hendrich leg./Lok. 14&quot; (CLH); 3 exs., &quot;Australia, N.T./Kakadu N.P., Jim Jim District, Gungurul Lookout, 50 m, 1.11.1996, 
<geoCoordinate degrees="13" direction="south" minutes="59.359" orientation="latitude" precision="1" value="-13.989317">13°59.359'S</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="19.904" orientation="longitude" precision="1" value="132.33173">132°19.904'E</geoCoordinate>
L. Hendrich leg./Lok. 11&quot; (CLH); 19 exs., &quot;Australia, N.T./Kakadu N.P., Mary River District, Gunlom Camping Area, 50 m, 3.11.1996, 
<geoCoordinate degrees="13" direction="south" minutes="26.082" orientation="latitude" precision="1" value="-13.4347">13°26.082'S</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="24.929" orientation="longitude" precision="1" value="132.41548">132°24.929'E</geoCoordinate>
L. Hendrich leg./Lok. 15&quot; (CLH); 2 exs., &quot;Australia, N.T./Kakadu N.P., Jim Jim District, Barramundie Gorge, Maguk, 50 m, 31.10.1996, 
<geoCoordinate degrees="13" direction="south" minutes="18.823" orientation="latitude" precision="1" value="-13.313717">13°18.823'S</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="26.198" orientation="longitude" precision="1" value="132.43663">132°26.198'E</geoCoordinate>
. L. Hendrich leg./ Lok. 9&quot; (CLH); 12 exs., &quot;Australia, NT/Kakadu N.P., Mary River District, Gunlom Waterfall Creek, 150 m, 2.11.1996, 
<geoCoordinate degrees="13" direction="south" minutes="26.082" orientation="latitude" precision="1" value="-13.4347">13°26.082'S</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="24.929" orientation="longitude" precision="1" value="132.41548">132°24.929'E</geoCoordinate>
L. Hendrich leg./Lok. 13&quot; (CLH); 51 exs., &quot;Australia, N.T./Kakadu N.P., Jim Jim District, 1 km S Jim Jim Falls, 70 m, 26.10.1996, 
<geoCoordinate degrees="13" direction="south" minutes="16.718" orientation="latitude" precision="1" value="-13.278633">13°16.718'S</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="49.490" orientation="longitude" precision="1" value="132.82483">132°49.490'E</geoCoordinate>
L. Hendrich leg./Lok. 3&quot; (CLH); 39 exs., &quot;Australia, N.T./Kakadu N.P., Jim Jim District, Jim Jim Falls Camping Area, Jim Jim Creek, 60 m, 26. &amp; 27.10.1996 
<geoCoordinate degrees="13" direction="south" minutes="16.218" orientation="latitude" precision="1" value="-13.2703">13°16.218'S</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="49.276" orientation="longitude" precision="1" value="132.82127">132°49.276'E</geoCoordinate>
L. Hendrich leg./ Lok. 2.&quot; (CLH); 13 exs., &quot;Australia N.T./Kakadu N.P., Nourlangie District, Gubara, 50 m, 25.10.1996, 
<geoCoordinate degrees="12" direction="south" minutes="50.101" orientation="latitude" precision="1" value="-12.835016">12°50.101'S</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="52.501" orientation="longitude" precision="1" value="132.87502">132°52.501'E</geoCoordinate>
L. Hendrich leg./Lok. 1&quot; (CLH); 1 ex., &quot;Australia, NT, Pine Creek, Roper Gulf (S)Waterfall Ck., 60 mi. E of Pine Ck. [-13.81667, 132.7], 7.08.1964, Carne, P.B. leg. (ANIC); 1 ex., &quot;Australia, NT, Daly Basin, Katherine (T) Tindal, 
<geoCoordinate degrees="14.51667" direction="south" orientation="latitude" precision="1" value="-14.51667">-14.51667</geoCoordinate>
, 
<geoCoordinate degrees="132.3667" direction="east" orientation="longitude" precision="5" value="132.3667">132.3667</geoCoordinate>
, 1.12.1967, Vestjens, W.J.M. leg. (ANIC); 10 exs., &quot;Australia: NT, Litchfield NP, TJAYNERA FALLS, 
<geoCoordinate degrees="13" direction="south" minutes="15" orientation="latitude" precision="925" value="-13.25">13°15'S</geoCoordinate>
, 
<geoCoordinate degrees="130" direction="east" minutes="44" orientation="longitude" precision="925" value="130.73334">130°44'E</geoCoordinate>
, 63 m, S. 
<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
leg., 20 
<normalizedToken originalValue="–27.XI.2008”">-27.XI.2008&quot;</normalizedToken>
(NMPC). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Queensland</emphasis>
: 2 exs., &quot;QLD Cook, 2 km NNW Jowalbinna [-15.75, 144.25] 17/01/1994 Zborowski P. &amp; Edwards, E.D.&quot;, 
<normalizedToken originalValue="“25-023624-470”">&quot;25-023624-470&quot;</normalizedToken>
(ANIC); 1 ex., &quot;QLD Cook, 14 km ENE Heathlands [-11.68, 142.7], rainforest, 21/10/1993 Zborowski P. &amp; Rentz D.C.F.&quot;, 
<normalizedToken originalValue="“25-022999-353”">&quot;25-022999-353&quot;</normalizedToken>
(ANIC); 1 ex., &quot;QLD, Townsville [
<geoCoordinate degrees="19.4833" direction="south" orientation="latitude" precision="5" value="-19.4833">-19.4833</geoCoordinate>
, 
<geoCoordinate degrees="146.8167" direction="east" orientation="longitude" precision="5" value="146.8167">146.8167</geoCoordinate>
], 25.3.1996, CH Watts&quot;, 
<normalizedToken originalValue="“25-001942”">&quot;25-001942&quot;</normalizedToken>
(SAMA); 1 ex., &quot;QLD, Townsville [-19.2, 146.68], 6-11/2/1998, AJ Watts&quot;, 
<normalizedToken originalValue="“25-001940”">&quot;25-001940&quot;</normalizedToken>
(SAMA); 1 ex., &quot;Bamaga N. Qld. March 1984 J.W. Turner&quot;, &quot;QDPC 0-172711&quot; (QDPC); 3 exs., &quot;Bamaca [=Bamaga] xii.1983&quot;, &quot;Australian QLD J. Sedlacek&quot; (CLH); 1 ex., &quot;Australia, Queensland, Bald Hills Station 4 km N of Isabella Falls, 15°15' 145°00', 29. December 1984, mv lamp, G. &amp; A. Daniels&quot; (QM).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Description of male.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Body shape</emphasis>
.
</emphasis>
In dorsal view, oblong oval, broadest at midlength of elytra, moderately convex. Body outline continuous, without discontinuity between pronotum and elytra. Head relatively broad; anterior margin of clypeus not bordered. Pronotum broadest between posterior angles, lateral margins moderately curved. Base of elytra as broad as pronotal base; lateral margins of elytra moderately curved (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Fig. 1</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 1.</emphasis>
Habitus and colouration of 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus bakewelli</emphasis>
</taxonomicName>
, female. Total length 5.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Colouration:</emphasis>
Body black, clypeus anteriorly, sides of pronotum, base of elytra, and humeral angle of elytra paler, ferruginous; ventral side reddish, appendages testaceous (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Fig. 1</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Dorsal surface sculpture:</emphasis>
Whole surface shiny (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Fig. 1</figureCitation>
). Head uniformly microreticulated, reticulation composed of moderately deeply impressed isodiametric meshes. Punctation composed of very small punctures, sparsely spread on surface; rows of deep and coarse punctures present around inner margin of eyes and in small depression anterolaterally of eyes. Pronotum with some moderately strong, short striae, more dense and weak laterally; lateral beading of pronotum very thin and indistinct. Microreticulation and punctation similar to that of head; row of coarse setigerous punctures present along anterior margin, basal margin (except for basomedially), and laterally close to sides. Elytra with microreticulation similar to that of head and pronotum, but less impressed. Punctation consisting of very fine sparse punctures. Each elytron with 11 strongly impressed discal and one submarginal longitudinal striae, intervals subequal, alternate striae tending to be shorter apically; subbasal stria reaching from middle of elytron almost to end of stria 10.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Antennae and legs:</emphasis>
Antenna with antennomeres long and slender. Protibia straight, not modified. Pro- and mesotarsomeres 1-3 distinctly broadened, with adhesive discs on their ventral side; claws simple.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Ventral part:</emphasis>
Finely microreticulated, with intermixed, sparsely distributed, very small punctures. Prosternum obtusely keeled medially. Prosternal process shortly lanceolate, apex obtuse; distinctly bordered laterally. Lateral parts of metaventrite tongue-shaped, very slender. Metacoxal lines almost complete and absent only close to metaventrite. Metacoxae with some short and deep striae, and abdominal ventrites 1-3 with numerous longitudinal striae.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Male genitalia:</emphasis>
Median lobe consisting of a few sclerites, closely attached together; apex in lateral view thin and elongate (
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Copelatus bakewelli A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure11" httpUri="https://binary.pensoft.net/fig/356480" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure11">Fig. 11A, B</figureCitation>
). Shape of paramere narrowly triangular, with thin, elongate subdistal part and weak but evident long setae along dorsal margin (
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Copelatus bakewelli A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure11" httpUri="https://binary.pensoft.net/fig/356480" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure11">Fig. 11D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="female">
<paragraph pageId="0" pageNumber="81">Female.</paragraph>
<paragraph pageId="0" pageNumber="81">Similar to male in habitus. Pro- and mesotarsomeres not broadened, without adhesive setae. Pronotal striae finer and denser.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="81">TL = 5.1-5.25 mm; TL-H = 4.5-4.65 mm; MW = 2.45-2.50 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="variability">
<paragraph pageId="0" pageNumber="81">Variability.</paragraph>
<paragraph pageId="0" pageNumber="81">All specimens studied are rather uniform in shape and colour, and vary only little in body length.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="differential diagnosis">
<paragraph pageId="0" pageNumber="81">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus bakewelli</emphasis>
</taxonomicName>
is close to the form of 
<taxonomicName lsidName="C. daemeli" pageId="0" pageNumber="81" rank="species" species="daemeli">
<emphasis italics="true" pageId="0" pageNumber="81">C. daemeli</emphasis>
</taxonomicName>
with 11 elytral striae and 
<taxonomicName lsidName="C. irregularis" pageId="0" pageNumber="81" rank="species" species="irregularis">
<emphasis italics="true" pageId="0" pageNumber="81">C. irregularis</emphasis>
</taxonomicName>
but smaller and without testaceous basal and apical markings on the elytra. Pronotal striae are weak (absent in 
<taxonomicName lsidName="C. daemeli" pageId="0" pageNumber="81" rank="species" species="daemeli">
<emphasis italics="true" pageId="0" pageNumber="81">C. daemeli</emphasis>
</taxonomicName>
) and the male protibia is not bent as in 
<taxonomicName lsidName="C. irregularis" pageId="0" pageNumber="81" rank="species" species="irregularis">
<emphasis italics="true" pageId="0" pageNumber="81">C. irregularis</emphasis>
</taxonomicName>
. Furthermore, all three species can easily be separated by the form of their median lobes.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="distribution">
<paragraph pageId="0" pageNumber="81">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="81">
Endemic. The species occurs from the Kimberley region in north-western Australia and in the Arnhemland in the Northern Territory to northern Queensland. In that area, the species is mainly distributed in rainforest pockets of the stone country (
<figureCitation captionStart="Figures 20, 21" captionStartId="F20" captionText="Figures 20, 21. Distribution of 20 Copelatus bakewelli 21 C. daemeli." figureDoi="10.3897/zookeys.889.39090.figures20-21" httpUri="https://binary.pensoft.net/fig/356489" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figures20-21">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="habitat">
<paragraph pageId="0" pageNumber="81">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="81">
A lentic species, which occurs in isolated pools of different size in otherwise dry riverbeds of seasonal streams, creeks and rivers. The bottom is consisted of gravel, sand and a layer of rotten plant debris. Occasionally, it occurs in protected bays, at the edge of large (25-50 m width), slow flowing and shallow (up to 20 cm depth) rivers, shaded by old riverine Melaleuca trees. There, the adults can be found among floating roots and organic debris of the paperbark trees. In northern Australia, at Gunlom waterfall, a large series was collected in a small (ca. 50 cm², 10 cm depth), water filled and shaded pothole (
<figureCitation captionStart="Figure 30" captionStartId="F28" captionText="Figure 30. Habitat of Copelatus bakewelli, rocky pool, with rotten leaves near Gunlom Waterfall, Northern Territory. Photographs by L. Hendrich." figureDoi="10.3897/zookeys.889.39090.figure30" httpUri="https://binary.pensoft.net/fig/356497" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure30">Fig. 30</figureCitation>
), filled up with rotten leaves. In Kakadu and the Kimberley region, 
<taxonomicName lsidName="C. bakewelli" pageId="0" pageNumber="81" rank="species" species="bakewelli">
<emphasis italics="true" pageId="0" pageNumber="81">C. bakewelli</emphasis>
</taxonomicName>
is more or less restricted to the stone country and the monsoon rainforest pockets. The species is also attracted to light.
</paragraph>
</subSubSection>
</treatment>
</document>