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<document id="E18FEA9F5CFB4CD4E5F465B676EB3CCE" ID-CLB-Dataset="30873" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-Zenodo-Dep="1163422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. &amp; Yağmur, Ersen A." docDate="2018" docId="03B9B44C56600C3E514A8191FD214953" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola anadolu Zonstein &amp; Kunt &amp; Yağmur 2018, sp. nov." docType="treatment" docUuid="D5C7C4A0-1FAD-4D2E-A531-E2F59280D327" docUuidSource="ZooBank" docVersion="13" lastPageNumber="26" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="24" updateTime="1698518020641" updateUser="ExternalLinkService">
<mods:mods id="9DBF2E32DDD7FE38E17590DD61868ED7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5B4C345FB236B25454A7ECAE97DFB7BC">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
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<mods:namePart id="75171DA8E4920B2DAA1532978BF1C087">Zonstein, Sergei</mods:namePart>
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<mods:namePart id="5B3F1ED5F259798D373E38D661F6FECD">Kunt, Kadir B.</mods:namePart>
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<mods:namePart id="459BA4B97216E01069FB4348BFBF0CFD">Yağmur, Ersen A.</mods:namePart>
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<mods:title id="F4760C38C7F7C3C1F3D313455E3078B0">European Journal of Taxonomy</mods:title>
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<mods:date id="124FCFE00D1E7B990FCFCC3887B9019B">2018</mods:date>
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<mods:number id="BC3155AA9289B34EA6462F2C479F2AFA">2018-01-30</mods:number>
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<treatment id="03B9B44C56600C3E514A8191FD214953" ID-DOI="http://doi.org/10.5281/zenodo.5980294" ID-GBIF-Taxon="140561349" ID-Zenodo-Dep="5980294" LSID="urn:lsid:zoobank.org:act:D5C7C4A0-1FAD-4D2E-A531-E2F59280D327" httpUri="http://treatment.plazi.org/id/03B9B44C56600C3E514A8191FD214953" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection id="C30A56D156600C30514A8191FB104CC7" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BAF055A56600C30514A8191FC674C98" blockId="23.[361,1226,1154,1254]" box="[630,957,1154,1181]" pageId="23" pageNumber="24">
<heading id="D0E7B23656600C30514A8191FC674C98" box="[630,957,1154,1181]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<taxonomicName id="4C107ED956600C30514A8191FC814C98" ID-CoL="4RMTG" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[630,859,1154,1181]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="anadolu" status="sp. nov.">
<emphasis id="B964D94856600C30514A8191FC814C98" bold="true" box="[630,859,1154,1181]" italics="true" pageId="23" pageNumber="24">Raveniola anadolu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356600C30505F8190FC674C98" box="[867,957,1155,1181]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BAF055A56600C30525581B4FB104CC7" blockId="23.[361,1226,1154,1254]" box="[361,1226,1191,1218]" pageId="23" pageNumber="24">
urn:lsid:zoobank.org:act:D5C7C4A0-1FAD-4D2E-A531-E2
<accessionNumber id="944398B956600C30571481B4FB5D4CC7" box="[1064,1159,1191,1218]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F59280" pageId="23" pageNumber="24" type="EnaNcbi">F59280</accessionNumber>
D327
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156600C30517981D8FC344CE0" box="[581,1006,1227,1254]" pageId="23" pageNumber="24" type="description">
<paragraph id="8BAF055A56600C30517981D8FC344CE0" blockId="23.[361,1226,1154,1254]" box="[581,1006,1227,1254]" pageId="23" pageNumber="24">
<figureCitation id="132B19DF56600C30517981D8FD454CE3" box="[581,671,1227,1254]" captionStart="Figs43–51" captionStartId="57.[189,241,1838,1864]" captionTargetBox="[190,1398,265,1787]" captionTargetId="figure@57.[189,1398,265,1787]" captionText="Figs43–51. Raveniolaspp.,habitusof ♂♂(43,45–46,48,50–51)and ♀♀(44,47,49),dorsalviews.— 43–44. R. adjarica sp. nov. 45. R. dunini sp. nov. 46–47. R. hyrcanica Dunin, 1988. 48–49. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163438/files/figure.png" pageId="23" pageNumber="24">Figs 51</figureCitation>
,
<figureCitation id="132B19DF56600C30519781D8FD114CE3" box="[683,715,1227,1254]" captionStart="Figs 74–78" captionStartId="61.[189,241,1818,1844]" captionTargetBox="[196,1397,266,1706]" captionTargetId="figure@61.[189,1401,265,1721]" captionText="Figs 74–78. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 74. R. biresekensis sp. nov. (dwarf conspecific male?). 75. R. anadolu sp. nov. (holotype). 76. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 77. R. turcica sp. nov. (holotype).78. R. micropa (Ausserer, 1871) (Uludağ). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163446/files/figure.png" pageId="23" pageNumber="24">75</figureCitation>
,
<figureCitation id="132B19DF56600C3051EB81D8FCDF4CE0" box="[727,773,1227,1253]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="23" pageNumber="24">108</figureCitation>
,
<figureCitation id="132B19DF56600C30502E81D8FC9B4CE0" box="[786,833,1227,1253]" captionStart="Figs 133–136" captionStartId="69.[189,241,1907,1933]" captionTargetBox="[255,1325,265,1873]" captionTargetId="figure@69.[227,1361,265,1874]" captionText="Figs 133–136. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 133. R. birecikensis sp. nov.(holotype).134. Same speciesas preceding (paratype, Birecik). 135. Samespecies as preceding (dwarf conspecific male?). 136. R. anadolu sp. nov.(holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163462/files/figure.png" pageId="23" pageNumber="24">136</figureCitation>
,
<figureCitation id="132B19DF56600C30507181D8FCA64CE3" box="[845,892,1227,1254]" captionStart="Figs 153–156" captionStartId="74.[189,241,1894,1920]" captionTargetBox="[189,1398,265,1868]" captionTargetId="figure@74.[189,1398,265,1868]" captionText="Figs 153–156. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 153. R. birecikensis sp. nov. (holotype). 154. Same species as preceding (paratype, Birecik, revolved bulbshowninventralposition).155.Samespeciesaspreceding(dwarf conspecific♂?). 156.R.anadolu sp. nov. (holotype). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1163472/files/figure.png" pageId="23" pageNumber="24">156</figureCitation>
,
<figureCitation id="132B19DF56600C3050BB81D8FC6F4CE0" box="[903,949,1227,1253]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="23" pageNumber="24">183</figureCitation>
,
<figureCitation id="132B19DF56600C3050FE81D8FC344CE0" box="[962,1006,1227,1253]" captionStart="Figure221" captionStartId="87.[189,272,842,868]" captionTargetBox="[189,1398,267,807]" captionTargetId="figure@87.[189,1398,265,810]" captionText="Figure221. Localitiesof Raveniolaspp.inTurkey:R.anadolu sp.nov.(),R.arthuriKunt&amp;Yağmur, 2010(),R.birecikensissp.nov.(),R.micropa(Ausserer,1871)(),R.nanasp.nov.(),R.sinani sp. nov.(), and R. turcica sp. nov. ().Abbreviation:AM =Armenia." httpUri="https://zenodo.org/record/1163498/files/figure.png" pageId="23" pageNumber="24">221</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156600C305381801EFC1F4DA5" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BAF055A56600C305381801EFEEC4D22" blockId="23.[189,310,1293,1319]" box="[189,310,1293,1319]" pageId="23" pageNumber="24">
<heading id="D0E7B23656600C305381801EFEEC4D22" bold="true" box="[189,310,1293,1319]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B964D94856600C305381801EFEEC4D22" bold="true" box="[189,310,1293,1319]" pageId="23" pageNumber="24">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56600C305381802EFC1F4DA5" blockId="23.[189,1399,1341,1440]" pageId="23" pageNumber="24">
In the structure of the palpal organ this species resembles
<taxonomicName id="4C107ED956600C305065802EFBF24D52" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[857,1064,1341,1367]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="turcica" status="sp. nov.">
<emphasis id="B964D94856600C305065802EFBF24D52" box="[857,1064,1341,1367]" italics="true" pageId="23" pageNumber="24">Raveniola turcica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356600C305713802EFB514D52" box="[1071,1163,1341,1367]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
, but differs from the latter by the presence of a small triangular embolic keel vestige and by the basal bend being confined to the median part of a shorter embolus (
<figureCitation id="132B19DF56600C3051478096FD3B4D9A" box="[635,737,1413,1439]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="23" pageNumber="24">Fig. 183</figureCitation>
; cf.
<figureCitation id="132B19DF56600C30502F8096FC624DA5" box="[787,952,1413,1440]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="23" pageNumber="24">Figs 185–186</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156600C30538180D4FE2F4E30" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BAF055A56600C30538180D4FE9E4DE4" blockId="23.[189,324,1479,1505]" box="[189,324,1479,1505]" pageId="23" pageNumber="24">
<heading id="D0E7B23656600C30538180D4FE9E4DE4" bold="true" box="[189,324,1479,1505]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B964D94856600C30538180D4FE9E4DE4" bold="true" box="[189,324,1479,1505]" pageId="23" pageNumber="24">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56600C30538180E5FE2F4E30" blockId="23.[189,1399,1526,1589]" pageId="23" pageNumber="24">The specific epithet is the Turkish name for Anatolia; the type locality of this species lies in the East Anatolia region of Turkey.</paragraph>
</subSubSection>
<subSubSection id="C30A56D156600C305381834EFBD54F50" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BAF055A56600C305381834EFE704E72" blockId="23.[189,426,1629,1655]" box="[189,426,1629,1655]" pageId="23" pageNumber="24">
<heading id="D0E7B23656600C305381834EFE704E72" bold="true" box="[189,426,1629,1655]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B964D94856600C305381834EFE704E72" bold="true" box="[189,426,1629,1655]" pageId="23" pageNumber="24">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B780F0756600C3053D6839EFC6E4EEA" ID-GBIF-Occurrence="1806244357" collectingDate="2011-09-10" collectionCode="AUZM" collectorName="K. B. Kunt" country="Turkey" elevation="1690" latitude="38.400932" location="surroundings of Altınsaç Village" longLatPrecision="1" longitude="42.891083" municipality="Gevaş District" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Van Province" typeStatus="holotype">
<paragraph id="8BAF055A56600C3053D6839EFE864EA2" blockId="23.[189,1399,1677,1775]" box="[234,348,1677,1703]" pageId="23" pageNumber="24">
<typeStatus id="54ABBBF856600C3053D6839EFE864EA2" box="[234,348,1677,1703]" pageId="23" pageNumber="24" type="holotype">
<emphasis id="B964D94856600C3053D6839EFE864EA2" bold="true" box="[234,348,1677,1703]" pageId="23" pageNumber="24">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BAF055A56600C30538183A2FC6E4EEA" blockId="23.[189,1399,1677,1775]" pageId="23" pageNumber="24">
<collectingCountry id="F30745CA56600C30538183A2FEEC4ECE" box="[189,310,1713,1739]" name="Turkey" pageId="23" pageNumber="24">TURKEY</collectingCountry>
:
<specimenCount id="9D16CED356600C30527783A2FEB94ECE" box="[331,355,1713,1739]" pageId="23" pageNumber="24" type="male">♂</specimenCount>
,
<collectingRegion id="49D4CBB856600C30524683A2FDFE4ECF" box="[378,548,1712,1739]" country="Turkey" name="Van" pageId="23" pageNumber="24">Van Province</collectingRegion>
,
<collectingMunicipality id="6BCB9F2056600C30510783A2FD2B4ECF" box="[571,753,1712,1739]" pageId="23" pageNumber="24">Gevaş District</collectingMunicipality>
,
<location id="8ECF538156600C30503B83A3FB7C4ECF" LSID="urn:lsid:plazi:treatment:03B9B44C56600C3E514A8191FD214953:8ECF538156600C30503B83A3FB7C4ECF" box="[775,1190,1712,1738]" country="Turkey" latitude="38.400932" longLatPrecision="1" longitude="42.891083" municipality="Gevaş District" name="surroundings of Altınsaç Village" pageId="23" pageNumber="24" stateProvince="Van Province">surroundings of Altınsaç Village</location>
,
<geoCoordinate id="EE24639D56600C30578083BCFAA84ECE" box="[1212,1394,1711,1739]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="38.400932">38°24′3.35″ N</geoCoordinate>
,
<geoCoordinate id="EE24639D56600C30538183C0FEA94EEA" box="[189,371,1747,1775]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="42.891083">42°53′27.89″ E</geoCoordinate>
,
<elevation id="003DE26956600C3052BC83C7FE014EEA" box="[384,475,1748,1775]" metricMagnitude="3" metricUnit="m" metricValue="1.69" pageId="23" pageNumber="24" unit="m" value="1690.0">
<quantity id="4CE8A8BF56600C3052BC83C7FE014EEA" box="[384,475,1748,1775]" metricMagnitude="3" metricUnit="m" metricValue="1.69" pageId="23" pageNumber="24" unit="m" value="1690.0">1690 m</quantity>
</elevation>
,
<collectingDate id="EFEADA7256600C3052DB83C6FD594EEA" box="[487,643,1749,1775]" pageId="23" pageNumber="24" value="2011-09-10">
<date id="FFAE239A56600C3052DB83C6FD594EEA" box="[487,643,1749,1775]" pageId="23" pageNumber="24" value="2011-09-10">10 Sep. 2011</date>
</collectingDate>
,
<collectorName id="26E5608C56600C3051B383C6FCD34EEA" box="[655,777,1749,1775]" pageId="23" pageNumber="24">K.B. Kunt</collectorName>
leg. (
<collectionCode id="ED019D9F56600C30507683C6FC7D4EEA" box="[842,935,1749,1775]" collectionName="AUZM" pageId="23" pageNumber="24">AUZM</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BAF055A56600C3053D68204FEBD4F34" blockId="23.[189,1039,1815,1877]" box="[234,359,1815,1841]" pageId="23" pageNumber="24">
<emphasis id="B964D94856600C3053D68204FEBD4F34" bold="true" box="[234,359,1815,1841]" pageId="23" pageNumber="24">Paratypes</emphasis>
</paragraph>
<paragraph id="8BAF055A56600C3053818228FBD54F50" blockId="23.[189,1039,1815,1877]" box="[189,1039,1851,1877]" pageId="23" pageNumber="24">
<materialsCitation id="3B780F0756600C3053818228FBD54F50" ID-GBIF-Occurrence="1806244396" box="[189,1039,1851,1877]" pageId="23" pageNumber="24">TURKEY: 4 ♂♂, collected together with the holotype (AUZM &amp; TAU).</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156600C3F5381826EFC4E4F50" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
<paragraph id="8BAF055A56600C305381826EFE944F92" blockId="23.[189,334,1917,1943]" box="[189,334,1917,1943]" pageId="23" pageNumber="24">
<heading id="D0E7B23656600C305381826EFE944F92" bold="true" box="[189,334,1917,1943]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B964D94856600C305381826EFE944F92" bold="true" box="[189,334,1917,1943]" pageId="23" pageNumber="24">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56600C3053D682BEFE714FCD" blockId="23.[189,439,1965,2027]" box="[234,427,1965,1992]" pageId="23" pageNumber="24">
<emphasis id="B964D94856600C3053D682BEFEF04FC2" bold="true" box="[234,298,1965,1991]" pageId="23" pageNumber="24">Male</emphasis>
(holotype)
</paragraph>
<paragraph id="8BAF055A56600C30538182C2FE6D4FEE" blockId="23.[189,439,1965,2027]" box="[189,439,2000,2027]" pageId="23" pageNumber="24">
HABITUS. See
<figureCitation id="132B19DF56600C30526182C2FE6D4FEE" box="[349,439,2000,2027]" captionStart="Figs43–51" captionStartId="57.[189,241,1838,1864]" captionTargetBox="[190,1398,265,1787]" captionTargetId="figure@57.[189,1398,265,1787]" captionText="Figs43–51. Raveniolaspp.,habitusof ♂♂(43,45–46,48,50–51)and ♀♀(44,47,49),dorsalviews.— 43–44. R. adjarica sp. nov. 45. R. dunini sp. nov. 46–47. R. hyrcanica Dunin, 1988. 48–49. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163438/files/figure.png" pageId="23" pageNumber="24">Fig. 51.</figureCitation>
</paragraph>
<paragraph id="8BAF055A566F0C3F5381841FFB7D4920" blockId="24.[189,1191,267,294]" box="[189,1191,267,294]" pageId="24" pageNumber="25">MEASUREMENTS. TBL 14.10, CL 4.93, CW 4.48, LL 0.36, LW 0.84, SL 2.56, SW 2.31.</paragraph>
<paragraph id="8BAF055A566F0C3F5381845DFB0049D1" blockId="24.[189,1399,333,468]" pageId="24" pageNumber="25">COLOUR. Carapace, chelicerae, palps and first pair of legs dorsally dark brownish rufous; eye tubercle somewhat darker with brownish black rings around eyes; sternum, labium, maxillae and legs II–IV light brownish rufous; abdomen dorsally brownish grey with numerous darker transverse fasciae and small partially fused lighter spots, ventral abdominal surface and spinnerets pale brownish grey.</paragraph>
<paragraph id="8BAF055A566F0C3F538184EFFE394AA0" blockId="24.[189,1399,507,677]" pageId="24" pageNumber="25">
PROSOMA. Carapace and chelicerae as shown in
<figureCitation id="132B19DF566F0C3F503E84E8FC864A13" box="[770,860,507,534]" captionStart="Figs 74–78" captionStartId="61.[189,241,1818,1844]" captionTargetBox="[196,1397,266,1706]" captionTargetId="figure@61.[189,1401,265,1721]" captionText="Figs 74–78. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 74. R. biresekensis sp. nov. (dwarf conspecific male?). 75. R. anadolu sp. nov. (holotype). 76. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 77. R. turcica sp. nov. (holotype).78. R. micropa (Ausserer, 1871) (Uludağ). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163446/files/figure.png" pageId="24" pageNumber="25">Fig. 75</figureCitation>
. Clypeus and eye group as in
<figureCitation id="132B19DF566F0C3F57EE84E8FAE04A10" box="[1234,1338,507,533]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="24" pageNumber="25">Fig. 108</figureCitation>
. Eye diameters and interdistances: AME 0.11(0.15), ALE 0.19, PLE 0.16, PME 0.16, AME–AME 0.11(0.08), ALE–AME 0.08(0.06), ALE–PLE 0.07, PLE–PME 0.03, PME–PME 0.36. Each cheliceral furrow with 9 promarginal teeth and 7–8 very small mesobasal denticles. Maxillae each with 20–22 cuspules arranged in triangle area.
</paragraph>
<paragraph id="8BAF055A566F0C3F538187DEFAFA4B2B" blockId="24.[189,1399,717,815]" pageId="24" pageNumber="25">
LEGS. Tibia and metatarsus I as shown in
<figureCitation id="132B19DF566F0C3F518287DEFCFC4AE2" box="[702,806,717,743]" captionStart="Figs 133–136" captionStartId="69.[189,241,1907,1933]" captionTargetBox="[255,1325,265,1873]" captionTargetId="figure@69.[227,1361,265,1874]" captionText="Figs 133–136. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 133. R. birecikensis sp. nov.(holotype).134. Same speciesas preceding (paratype, Birecik). 135. Samespecies as preceding (dwarf conspecific male?). 136. R. anadolu sp. nov.(holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163462/files/figure.png" pageId="24" pageNumber="25">Fig. 136</figureCitation>
. Scopula: distal on metatarsi I–II, entire on tarsi I, divided on tarsi II; absent on tarsi III–IV. Trichobothria: 2 rows of 8–10 each on tibiae, 10–12 on metatarsi, 10–12 on tarsi, 6–8 on cymbium. Paired tarsal claws with 9–13 teeth on each margin.
</paragraph>
<paragraph id="8BAF055A566F0C3F53818644FE7A4B74" blockId="24.[189,416,855,881]" box="[189,416,855,881]" pageId="24" pageNumber="25">LEG MEASUREMENTS.</paragraph>
<paragraph id="8BAF055A566F0C3F52568681FAE74C7E" pageId="24" pageNumber="25">
<table id="F910F7FA566FF3D853F48682FAE74C79" box="[200,1341,913,1148]" colsContinueFrom="22.[200,1341,1653,1889]" colsContinueIn="35.[200,1387,330,566]" gridcols="7" gridrows="6" pageId="24" pageNumber="25">
<tr id="35200718566FF3D853F48682FAE74BA9" box="[200,1341,913,940]" gridrow="0" pageId="24" pageNumber="25" rowspan-0="1">
<th id="76F16E64566FF3D852568682FE6D4BA9" box="[362,439,913,940]" gridcol="1" gridrow="0" pageId="24" pageNumber="25">Femur</th>
<th id="76F16E64566FF3D851228682FDB44BA9" box="[542,622,913,940]" gridcol="2" gridrow="0" pageId="24" pageNumber="25">Patella</th>
<th id="76F16E64566FF3D851E18682FCC04BA9" box="[733,794,913,940]" gridcol="3" gridrow="0" pageId="24" pageNumber="25">Tibia</th>
<th id="76F16E64566FF3D8504D8682FC284BA9" box="[881,1010,913,940]" gridcol="4" gridrow="0" pageId="24" pageNumber="25">Metatarsus</th>
<th id="76F16E64566FF3D8577C8682FB564BA9" box="[1088,1164,913,940]" gridcol="5" gridrow="0" pageId="24" pageNumber="25">Tarsus</th>
<th id="76F16E64566FF3D857C78682FAE74BA9" box="[1275,1341,913,940]" gridcol="6" gridrow="0" pageId="24" pageNumber="25">Total</th>
</tr>
<tr id="35200718566FF3D853F486A8FAE74BD3" box="[200,1341,955,982]" gridrow="1" pageId="24" pageNumber="25">
<th id="76F16E64566FF3D853F486A8FF264BD3" box="[200,252,955,982]" gridcol="0" gridrow="1" pageId="24" pageNumber="25">Palp</th>
<td id="76F16E64566FF3D8525686A8FE6D4BD3" box="[362,439,955,982]" gridcol="1" gridrow="1" pageId="24" pageNumber="25">2.55</td>
<td id="76F16E64566FF3D8512286A8FDB44BD3" box="[542,622,955,982]" gridcol="2" gridrow="1" pageId="24" pageNumber="25">1.50</td>
<td id="76F16E64566FF3D851E186A8FCC04BD3" box="[733,794,955,982]" gridcol="3" gridrow="1" pageId="24" pageNumber="25">2.16</td>
<td id="76F16E64566FF3D8504D86A8FC284BD3" box="[881,1010,955,982]" gridcol="4" gridrow="1" pageId="24" pageNumber="25">–</td>
<td id="76F16E64566FF3D8577C86A8FB564BD3" box="[1088,1164,955,982]" gridcol="5" gridrow="1" pageId="24" pageNumber="25">0.90</td>
<td id="76F16E64566FF3D857C786A8FAE74BD3" box="[1275,1341,955,982]" gridcol="6" gridrow="1" pageId="24" pageNumber="25">7.11</td>
</tr>
<tr id="35200718566FF3D853F486F7FAE74BFA" box="[200,1341,996,1023]" gridrow="2" pageId="24" pageNumber="25">
<th id="76F16E64566FF3D853F486F7FF264BFA" box="[200,252,996,1023]" gridcol="0" gridrow="2" pageId="24" pageNumber="25">I</th>
<td id="76F16E64566FF3D8525686F7FE6D4BFA" box="[362,439,996,1023]" gridcol="1" gridrow="2" pageId="24" pageNumber="25">4.51</td>
<td id="76F16E64566FF3D8512286F7FDB44BFA" box="[542,622,996,1023]" gridcol="2" gridrow="2" pageId="24" pageNumber="25">2.41</td>
<td id="76F16E64566FF3D851E186F7FCC04BFA" box="[733,794,996,1023]" gridcol="3" gridrow="2" pageId="24" pageNumber="25">3.44</td>
<td id="76F16E64566FF3D8504D86F7FC284BFA" box="[881,1010,996,1023]" gridcol="4" gridrow="2" pageId="24" pageNumber="25">3.44</td>
<td id="76F16E64566FF3D8577C86F7FB564BFA" box="[1088,1164,996,1023]" gridcol="5" gridrow="2" pageId="24" pageNumber="25">1.97</td>
<td id="76F16E64566FF3D857C786F7FAE74BFA" box="[1275,1341,996,1023]" gridcol="6" gridrow="2" pageId="24" pageNumber="25">15.77</td>
</tr>
<tr id="35200718566FF3D853F4811DFAE74C2C" box="[200,1341,1038,1065]" gridrow="3" pageId="24" pageNumber="25">
<th id="76F16E64566FF3D853F4811DFF264C2C" box="[200,252,1038,1065]" gridcol="0" gridrow="3" pageId="24" pageNumber="25">II</th>
<td id="76F16E64566FF3D85256811DFE6D4C2C" box="[362,439,1038,1065]" gridcol="1" gridrow="3" pageId="24" pageNumber="25">4.09</td>
<td id="76F16E64566FF3D85122811DFDB44C2C" box="[542,622,1038,1065]" gridcol="2" gridrow="3" pageId="24" pageNumber="25">2.08</td>
<td id="76F16E64566FF3D851E1811DFCC04C2C" box="[733,794,1038,1065]" gridcol="3" gridrow="3" pageId="24" pageNumber="25">2.92</td>
<td id="76F16E64566FF3D8504D811DFC284C2C" box="[881,1010,1038,1065]" gridcol="4" gridrow="3" pageId="24" pageNumber="25">2.86</td>
<td id="76F16E64566FF3D8577C811DFB564C2C" box="[1088,1164,1038,1065]" gridcol="5" gridrow="3" pageId="24" pageNumber="25">1.69</td>
<td id="76F16E64566FF3D857C7811DFAE74C2C" box="[1275,1341,1038,1065]" gridcol="6" gridrow="3" pageId="24" pageNumber="25">13.64</td>
</tr>
<tr id="35200718566FF3D853F4812BFAE74C56" box="[200,1341,1080,1107]" gridrow="4" pageId="24" pageNumber="25">
<th id="76F16E64566FF3D853F4812BFF264C56" box="[200,252,1080,1107]" gridcol="0" gridrow="4" pageId="24" pageNumber="25">III</th>
<td id="76F16E64566FF3D85256812BFE6D4C56" box="[362,439,1080,1107]" gridcol="1" gridrow="4" pageId="24" pageNumber="25">3.61</td>
<td id="76F16E64566FF3D85122812BFDB44C56" box="[542,622,1080,1107]" gridcol="2" gridrow="4" pageId="24" pageNumber="25">1.83</td>
<td id="76F16E64566FF3D851E1812BFCC04C56" box="[733,794,1080,1107]" gridcol="3" gridrow="4" pageId="24" pageNumber="25">2.35</td>
<td id="76F16E64566FF3D8504D812BFC284C56" box="[881,1010,1080,1107]" gridcol="4" gridrow="4" pageId="24" pageNumber="25">3.05</td>
<td id="76F16E64566FF3D8577C812BFB564C56" box="[1088,1164,1080,1107]" gridcol="5" gridrow="4" pageId="24" pageNumber="25">1.81</td>
<td id="76F16E64566FF3D857C7812BFAE74C56" box="[1275,1341,1080,1107]" gridcol="6" gridrow="4" pageId="24" pageNumber="25">12.65</td>
</tr>
<tr id="35200718566FF3D853F48172FAE74C79" box="[200,1341,1121,1148]" gridrow="5" pageId="24" pageNumber="25">
<th id="76F16E64566FF3D853F48172FF264C79" box="[200,252,1121,1148]" gridcol="0" gridrow="5" pageId="24" pageNumber="25">IV</th>
<td id="76F16E64566FF3D852568172FE6D4C79" box="[362,439,1121,1148]" gridcol="1" gridrow="5" pageId="24" pageNumber="25">4.36</td>
<td id="76F16E64566FF3D851228172FDB44C79" box="[542,622,1121,1148]" gridcol="2" gridrow="5" pageId="24" pageNumber="25">2.09</td>
<td id="76F16E64566FF3D851E18172FCC04C79" box="[733,794,1121,1148]" gridcol="3" gridrow="5" pageId="24" pageNumber="25">3.19</td>
<td id="76F16E64566FF3D8504D8172FC284C79" box="[881,1010,1121,1148]" gridcol="4" gridrow="5" pageId="24" pageNumber="25">4.28</td>
<td id="76F16E64566FF3D8577C8172FB564C79" box="[1088,1164,1121,1148]" gridcol="5" gridrow="5" pageId="24" pageNumber="25">2.15</td>
<td id="76F16E64566FF3D857C78172FAE74C79" box="[1275,1341,1121,1148]" gridcol="6" gridrow="5" pageId="24" pageNumber="25">16.07</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF055A566F0C3F5381818EFDD84D8B" blockId="24.[189,1399,1181,1422]" pageId="24" pageNumber="25">SPINATION. Patellae I and II aspinose. Palp: femur d1–1–1–1, pd0–0–1, rd0–0–1; patella p1–1; tibia d1–1–1, p1–1–1–1, r1–1, pv1–1–1, v1–2–1; cymbium d5. Leg I: femur d1–1–1–1, pd0–1(0)–1; tibia p1–1–0, rv0–1–m–m, v1–0–1–0; metatarsus v0–0–2. Leg II: femur d1–1–1–1, pv1(0)–1–1; tibia p1– 1–1, v2–2–2; metatarsus v2–2–2. Leg III: femur d1–1–1–1, pd1–1–1, rd1–1–1; patella p1, r1; tibia d1–1(0)–0, p0–1–1, r1–1–1, v2–2–3; metatarsus d1–1–2, p0–1–1, r1–1–1, v2–2–3. Leg IV: femur d1– 1–1–1, pd0–0–1, rd0–0–1; patella r1; tibia d1–0–0, p1–1–1, r1–1–1–1, v2–2–3; metatarsus d1–1–2, p1–1–1, r1–1–1–1, v2–2–3.</paragraph>
<paragraph id="8BAF055A566F0C3F538180A4FCDE4DF1" blockId="24.[189,1399,1462,1524]" pageId="24" pageNumber="25">
PALP. Tibia, cymbium and palpal organ as shown in
<figureCitation id="132B19DF566F0C3F506880A5FC644DD4" box="[852,958,1462,1489]" captionStart="Figs 153–156" captionStartId="74.[189,241,1894,1920]" captionTargetBox="[189,1398,265,1868]" captionTargetId="figure@74.[189,1398,265,1868]" captionText="Figs 153–156. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 153. R. birecikensis sp. nov. (holotype). 154. Same species as preceding (paratype, Birecik, revolved bulbshowninventralposition).155.Samespeciesaspreceding(dwarf conspecific♂?). 156.R.anadolu sp. nov. (holotype). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1163472/files/figure.png" pageId="24" pageNumber="25">Fig. 156</figureCitation>
. Embolus with characteristic small, triangular vestige of dorsomedian keel (
<figureCitation id="132B19DF566F0C3F51AC80C9FD2D4DF1" box="[656,759,1498,1524]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="24" pageNumber="25">Fig. 183</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A566F0C3F5381830EFB794E5F" blockId="24.[189,1398,1564,1627]" pageId="24" pageNumber="25">SPINNERETS. PMS: length 0.46; diameter 0.18. PLS: maximal diameter 0.39; length of basal, medial and apical segments 0.71, 0.52, 0.51; total length 1.74; apical segment shortly digitiform.</paragraph>
<paragraph id="8BAF055A566F0C3F53D68390FE9F4E98" blockId="24.[189,325,1667,1728]" box="[234,325,1667,1693]" pageId="24" pageNumber="25">
<emphasis id="B964D948566F0C3F53D68390FE9F4E98" bold="true" box="[234,325,1667,1693]" pageId="24" pageNumber="25">Female</emphasis>
</paragraph>
<paragraph id="8BAF055A566F0C3F538183B5FEE34EC5" blockId="24.[189,325,1667,1728]" box="[189,313,1702,1728]" pageId="24" pageNumber="25">Unknown.</paragraph>
<paragraph id="8BAF055A566F0C3F538183FBFEE94F07" blockId="24.[189,307,1768,1794]" box="[189,307,1768,1794]" pageId="24" pageNumber="25">
<heading id="D0E7B236566F0C3F538183FBFEE94F07" bold="true" box="[189,307,1768,1794]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B964D948566F0C3F538183FBFEE94F07" bold="true" box="[189,307,1768,1794]" pageId="24" pageNumber="25">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A566F0C3F5381820BFC4E4F50" blockId="24.[189,1398,1815,1878]" pageId="24" pageNumber="25">Carapace length in males varies from 4.22 to 5.04. The size and shape of the embolic keel vestige in all paratype males do not differ from those in the holotypic male.</paragraph>
</subSubSection>
<subSubSection id="C30A56D1566F0C3F5381826DFCED4FEE" pageId="24" pageNumber="25" type="biology_ecology">
<paragraph id="8BAF055A566F0C3F5381826DFEFA4F9D" blockId="24.[189,288,1918,1944]" box="[189,288,1918,1944]" pageId="24" pageNumber="25">
<heading id="D0E7B236566F0C3F5381826DFEFA4F9D" bold="true" box="[189,288,1918,1944]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B964D948566F0C3F5381826DFEFA4F9D" bold="true" box="[189,288,1918,1944]" pageId="24" pageNumber="25">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A566F0C3F538182BEFCED4FEE" blockId="24.[189,1398,1965,2027]" pageId="24" pageNumber="25">
The type series, which includes only males, was collected with pitfall traps in an open rocky biotope sparsely covered with low
<taxonomicName id="4C107ED9566F0C3F52CB82C1FDB14FEE" box="[503,619,2002,2027]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B964D948566F0C3F52CB82C1FDB14FEE" box="[503,619,2002,2027]" italics="true" pageId="24" pageNumber="25">Juniperus</emphasis>
</taxonomicName>
trees and shrubs.
</paragraph>
</subSubSection>
<subSubSection id="C30A56D1566E0C3E53818418FD214953" box="[189,763,267,342]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BAF055A566E0C3E53818418FE8C4920" blockId="25.[189,342,267,293]" box="[189,342,267,293]" pageId="25" pageNumber="26">
<heading id="D0E7B236566E0C3E53818418FE8C4920" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B964D948566E0C3E53818418FE8C4920" bold="true" box="[189,342,267,293]" pageId="25" pageNumber="26">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A566E0C3E5381842FFD214953" blockId="25.[189,763,315,342]" box="[189,763,315,342]" pageId="25" pageNumber="26">
Known only from the type locality. See
<figureCitation id="132B19DF566E0C3E51AE8428FD2D4953" box="[658,759,315,342]" captionStart="Figure221" captionStartId="87.[189,272,842,868]" captionTargetBox="[189,1398,267,807]" captionTargetId="figure@87.[189,1398,265,810]" captionText="Figure221. Localitiesof Raveniolaspp.inTurkey:R.anadolu sp.nov.(),R.arthuriKunt&amp;Yağmur, 2010(),R.birecikensissp.nov.(),R.micropa(Ausserer,1871)(),R.nanasp.nov.(),R.sinani sp. nov.(), and R. turcica sp. nov. ().Abbreviation:AM =Armenia." httpUri="https://zenodo.org/record/1163498/files/figure.png" pageId="25" pageNumber="26">Fig. 221</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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