353 lines
34 KiB
XML
353 lines
34 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1168.99302" ID-Pensoft-Pub="1313-2970-1168-267" ID-Pensoft-UUID="01F93FFC6397552587D46E5748153639" ID-ZooBank="28659D393708403584BFA95AC91C59FF" ModsDocID="1313-2970-1168-267" checkinTime="1688439528747" checkinUser="pensoft" docAuthor="Hayashi, Masakazu, Iwata, Tomofumi & Yoshitomi, Hiroyuki" docDate="2023" docId="F1B9F79A3CDF538C97C78C311848E4BC" docLanguage="en" docName="ZooKeys 1168: 267-294" docOrigin="ZooKeys 1168" docPubDate="2023-07-03" docSource="http://dx.doi.org/10.3897/zookeys.1168.99302" docTitle="Haliplus (Liaphlus) kotoshonis Kano & Kamiya 1931" docType="treatment" docVersion="1" id="01F93FFC6397552587D46E5748153639" lastPageNumber="267" masterDocId="01F93FFC6397552587D46E5748153639" masterDocTitle="Revision of the family Haliplidae (Insecta, Coleoptera) in Japan" masterLastPageNumber="294" masterPageNumber="267" pageNumber="267" updateTime="1688439528747" updateUser="pensoft">
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<mods:title id="B3EC0BAF3F7A4F1952A39F3C8A5E977A">Revision of the family Haliplidae (Insecta, Coleoptera) in Japan</mods:title>
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<mods:name id="8BFB215DEEC99BFB30EC6FEA17DBB9E0" type="personal">
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<mods:namePart id="54FACDAF534EA9E1EC35A967FE3C826C">Hayashi, Masakazu</mods:namePart>
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<mods:nameIdentifier id="67DB0FD3883F2E8AFD0B8F7D04DE05D2" type="ORCID">https://orcid.org/0000-0003-3708-9005</mods:nameIdentifier>
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<mods:affiliation id="C0848795E3EBA27CD27B6AA70AE01308">Hoshizaki Green Foundation, Sono, Izumo, 691 - 0076, Japan</mods:affiliation>
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<mods:nameIdentifier id="BAEE941B5F506C35300D6EEF5F060AB7" type="email">hgf-haya@green-f.or.jp</mods:nameIdentifier>
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<mods:roleTerm id="1E7D49467D8921C01214DB78D1401B1D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3822F96FF6E474089824157E44AF1B4E">Iwata, Tomofumi</mods:namePart>
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<mods:nameIdentifier id="12D737BE31A8A63D69764FD2EB8467D9" type="ORCID">https://orcid.org/0000-0002-7738-3199</mods:nameIdentifier>
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<mods:affiliation id="36F9D55646EF6B22EA68DC2B34375DA7">Toyama Science Museum, 1 - 8 - 31 Nishinakano-machi, Toyama, 939 - 8084, Japan</mods:affiliation>
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<mods:name id="DCFA95B8C41103CBA895324DA863504D" type="personal">
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<mods:namePart id="5AF0DF38274BAEBD52C636124F9BA92B">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:nameIdentifier id="D2A002EC9BA0A48D8CFA3F3681AA2E4A" type="ORCID">https://orcid.org/0000-0001-8416-9249</mods:nameIdentifier>
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<mods:affiliation id="ADC831E4BA993B65D3C7CDC5C812A061">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566, Japan</mods:affiliation>
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<mods:nameIdentifier id="7395712402FA2A30B3B31F6F956E9F2F" type="email">hymushi@agr.ehime-u.ac.jp</mods:nameIdentifier>
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<mods:typeOfResource id="E70CCF1687DC8642B5B5F5C05CF2106B">text</mods:typeOfResource>
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<mods:title id="EC8FC3F873A9D3E25A0BA4241B897B5D">ZooKeys</mods:title>
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<mods:part id="F79806EE7C907577E11861229D08B2CA">
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<mods:date id="F88850E1AA67F8AC4D360E9C99D97D90">2023</mods:date>
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<mods:number id="9FB90010D76ED0240B4B2DC9C875DFB3">2023-07-03</mods:number>
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<mods:detail id="90DCEC3490B2B925BEAE3809DBEC0BEF" type="volume">
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<mods:number id="0C13F8C13216C770CD2F2D8AA535D37B">1168</mods:number>
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<mods:url id="F293CFB76C8BBFB2C6E5BFCB91CB55C1">http://dx.doi.org/10.3897/zookeys.1168.99302</mods:url>
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<mods:classification id="C3173A101E602D22937D89D92BF53EEB">journal article</mods:classification>
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<mods:identifier id="F4EA5B7E84FA152DCADA2500AA0ECA4E" type="DOI">http://dx.doi.org/10.3897/zookeys.1168.99302</mods:identifier>
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<mods:identifier id="ECA85BFB2A24F60B73F0177C3E212915" type="Pensoft-Pub">1313-2970-1168-267</mods:identifier>
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<treatment id="F1B9F79A3CDF538C97C78C311848E4BC" LSID="urn:lsid:plazi:treatment:F1B9F79A3CDF538C97C78C311848E4BC" httpUri="http://treatment.plazi.org/id/F1B9F79A3CDF538C97C78C311848E4BC" lastPageNumber="267" pageId="0" pageNumber="267">
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<subSubSection id="A88B5088FE467021E034323D67F4C2E4" pageId="0" pageNumber="267" type="nomenclature">
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<paragraph id="62743079F7BB3B2F457CD02DA4B940AB" pageId="0" pageNumber="267">
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<taxonomicName id="A589721814A8C84A81633A369F2642C0" LSID="F1B9F79A-3CDF-538C-97C7-8C311848E4BC" authority="Kano & Kamiya, 1931" authorityName="Kano & Kamiya" authorityYear="1931" class="Insecta" family="Haliplidae" genus="Haliplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haliplus (Liaphlus) kotoshonis" order="Coleoptera" pageId="0" pageNumber="267" phylum="Arthropoda" rank="species" species="kotoshonis" subGenus="Liaphlus">Haliplus (Liaphlus) kotoshonis Kano & Kamiya, 1931</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="09E0DB1D77342D2A01CD00A91B54BEB4" pageId="0" pageNumber="267" type="description">
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<paragraph id="BD51FF3308F8A959913EB7295DA6C5A6" pageId="0" pageNumber="267">
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<figureCitation id="56F232F5E4B99197881F9CACEDA91433" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Haliplus kotoshonis A habitus B head C prosternal process D penis E left paramere F right paramere. Scale bars: 1.0 mm (A); 0.5 mm (B); 0.25 mm (C); 0.1 mm (D-F)." figureDoi="10.3897/zookeys.1168.99302.figure14" httpUri="https://binary.pensoft.net/fig/872274" pageId="0" pageNumber="267">Figs 14</figureCitation>
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<figureCitation id="432C4A9EB1D37B8F4C07ADD2535CB602" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution maps based on specimens examined A Peltodytes intermedius B Peltodytes sinensis C Haliplus japonicus D Haliplus kamiyai (circle), Haliplus morii (star), Haliplus regimbarti (square) E Haliplus simplex (black circle), Haliplus angustifrons (white circle) F Haliplus basinotatus (black circle), Haliplus kotoshonis (white circle)." figureDoi="10.3897/zookeys.1168.99302.figure18" httpUri="https://binary.pensoft.net/fig/872278" pageId="0" pageNumber="267">, 18F Japanese name: Koto-kogashira-mizumushi</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="60B959CC9EEB212A832219AE76B64DFF" pageId="0" pageNumber="267" type="reference_group">
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<paragraph id="DEE4FCF71566530533C6671744EA031E" pageId="0" pageNumber="267">
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<taxonomicName id="87FA0AFEB0E1188C313D503E0A67148E" authorityName="Kano & Kamiya" authorityYear="1931" class="Insecta" family="Haliplidae" genus="Haliplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haliplus kotoshonis" order="Coleoptera" pageId="0" pageNumber="267" phylum="Arthropoda" rank="species" species="kotoshonis">Haliplus kotoshonis</taxonomicName>
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Kano & Kamiya, 1931: 2.
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<bibRefCitation id="5A9A643E65D1A2CF7FFE92B4439B980D" author="Sato, M" editor="Ueno, S" journalOrPublisher="II). Hoikusha, Osaka" pageId="0" pageNumber="267" pagination="180 - 181" refId="B35" refString="Sato, M, 1985. Haliplidae. In: Ueno, S, Kurosawa, Y, Sato, M, Eds., The Coleoptera of Japan in Color (Vol. II). Hoikusha, Osaka: 180 - 181" title="Haliplidae." volumeTitle="The Coleoptera of Japan in Color (Vol." year="1985">
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<normalizedToken id="DB76072A0B5EF6227F45EBAD85A397B8" originalValue="Satô">Sato</normalizedToken>
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1985
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</bibRefCitation>
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: 181, pls 33-37;
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<bibRefCitation id="9BC98FE031B96075579FA78FD4F04745" author="Nakane, T" journalOrPublisher="Kita-Kyushu no Konchu" pageId="0" pageNumber="267" pagination="61 - 67" refId="B29" refString="Nakane, T, 1985. A revisional study on Japanese species of the genus Haliplus Latreille (Coleoptera, Haliplidae). Kita-Kyushu no Konchu 32: 61 - 67" title="A revisional study on Japanese species of the genus Haliplus Latreille (Coleoptera, Haliplidae)." volume="32" year="1985">Nakane 1985</bibRefCitation>
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: 63;
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<bibRefCitation id="669DC06C8449BF283EA9E6FEEB0BCDEB" author="Nakane, T" journalOrPublisher="Insects & Nature" pageId="0" pageNumber="267" pagination="26 - 30" refId="B30" refString="Nakane, T, 1987. The beetles of Japan (new series) 80. Insects & Nature 22 (11): 26 - 30" title="The beetles of Japan (new series) 80." volume="22" year="1987">1987</bibRefCitation>
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: 30;
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<bibRefCitation id="0D4784EFA093DF0E035EC70E0B4625FD" author="Nakajima, J" journalOrPublisher="Bun-ichi Sogo Shuppan, Tokyo" pageId="0" pageNumber="267" refId="B25" refString="Nakajima, J, Hayashi, M, Ishida, K, Kitano, T, Yoshitomi, H, 2020. Aquatic Coleoptera and Hemiptera of Japan. Bun-ichi Sogo Shuppan, Tokyo" title="Aquatic Coleoptera and Hemiptera of Japan." year="2020">Nakajima et al. 2020</bibRefCitation>
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: 21.
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</paragraph>
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<paragraph id="E73463F2BCFC591BB82EDCB4F6C044A8" pageId="0" pageNumber="267">
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<taxonomicName id="9905408CE58800ADB318F5B8D407CD68" authorityName="Kano & Kamiya" authorityYear="1931" class="Insecta" family="Haliplidae" genus="Haliplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haliplus (Liaphlus) kotoshonis" order="Coleoptera" pageId="0" pageNumber="267" phylum="Arthropoda" rank="species" species="kotoshonis" subGenus="Liaphlus">Haliplus (Liaphlus) kotoshonis</taxonomicName>
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:
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<bibRefCitation id="F1F6A9112CC3A2951AD1A9E459B27C53" author="Sato, M" journalOrPublisher="News" pageId="0" pageNumber="267" pagination="1 - 4" refId="B34" refString="Sato, M, 1984. Taxonomic notes on the aquatic Coleoptera of Japan, I. Coleopterists'. News 65: 1 - 4" title="Taxonomic notes on the aquatic Coleoptera of Japan, I. Coleopterists'." volume="65" year="1984">
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<normalizedToken id="3A017E04175A87B94C26DD4F55AC371A" originalValue="Satô">Sato</normalizedToken>
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1984
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</bibRefCitation>
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: 2.
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</paragraph>
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<paragraph id="8C2631F77C266AD0606FF14BD5A838CD" pageId="0" pageNumber="267">
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<taxonomicName id="6FAFAD929AB08FDFD3802C01375D66DE" authorityName="J.Balfour-Browne" authorityYear="1947" class="Insecta" family="Haliplidae" genus="Haliplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haliplus diruptus" order="Coleoptera" pageId="0" pageNumber="267" phylum="Arthropoda" rank="species" species="diruptus">Haliplus diruptus</taxonomicName>
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J. Balfour-Browne, 1946: 436. Vondel 1995: 120, figs 71-79. Syn. nov.
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</paragraph>
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<paragraph id="815FB601B2E8AE242847BBE2E0D7E69E" pageId="0" pageNumber="267">
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<taxonomicName id="8186AECD93702F7ACDF4EC2171B6585D" authorityName="Vondel" authorityYear="1991" class="Insecta" family="Haliplidae" genus="Haliplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haliplus davidi" order="Coleoptera" pageId="0" pageNumber="267" phylum="Arthropoda" rank="species" species="davidi">Haliplus davidi</taxonomicName>
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Vondel, 1991: 92. [synonymized with
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<taxonomicName id="606C96E1A1F0659F09B13B950A358427" authorityName="J.Balfour-Browne" authorityYear="1947" class="Insecta" family="Haliplidae" genus="Haliplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haliplus diruptus" order="Coleoptera" pageId="0" pageNumber="267" phylum="Arthropoda" rank="species" species="diruptus">Haliplus diruptus</taxonomicName>
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by
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<bibRefCitation id="9E45542A30FEE6DA970B0B66448FF03C" author="van Vondel, BJ" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="267" pagination="31 - 35" refId="B52" refString="van Vondel, BJ, 2017. Five new synonymies in Haliplus subgen. Liaphlus Guignot, based on the variability of the left paramere (Coleoptera: Haliplidae). Koleopterologische Rundschau 87: 31 - 35" title="Five new synonymies in Haliplus subgen. Liaphlus Guignot, based on the variability of the left paramere (Coleoptera: Haliplidae)." volume="87" year="2017">van Vondel (2017)</bibRefCitation>
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]
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</paragraph>
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</subSubSection>
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<subSubSection id="70AA59ECD174A9889CABB4B1D1B1ECB5" pageId="0" pageNumber="267" type="materials_examined">
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<paragraph id="A5347F2BCC9ECBB3E6015E383D9FE94E" pageId="0" pageNumber="267">Material examined.</paragraph>
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<paragraph id="9DE2C14B133B53BE55A89AB3B6602C6B" pageId="0" pageNumber="267">
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<materialsCitation id="A14BB41D23DC78BCD0075CCCA26E109B" collectingDate="1960-07-02" collectorName="M. Sato" county="Tokara" location="Kagoshima Prefecture" municipality="Takara-jima" specimenCount="1">
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<specimenCount id="915656D0D0C18E9A42BAC5C4F9DA53FA" type="generic">1 ex.</specimenCount>
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,
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<location id="3350F9748F65B6751D950F16EF2984D6" LSID="urn:lsid:plazi:treatment:F1B9F79A3CDF538C97C78C311848E4BC:3350F9748F65B6751D950F16EF2984D6" county="Tokara" municipality="Takara-jima" name="Kagoshima Prefecture">Kagoshima Prefecture</location>
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:
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<collectingMunicipality id="54B3FD5299292EF36EACB814149C9AD8">Takara-jima</collectingMunicipality>
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,
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<collectingCounty id="15BDC3CF7055CBC972EDC8536333CECF">Tokara</collectingCounty>
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,
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<collectingDate id="3D1234E920BDAA5D396361053D22F172" value="1960-07-02">2.VII.1960</collectingDate>
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,
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<collectorName id="F124F489F765B25283A8AEDF2F075F20">
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M.
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<normalizedToken id="476A77E62424AF37F71380F9F4BA37B3" originalValue="Satô">Sato</normalizedToken>
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</collectorName>
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leg. (EUMJ)
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</materialsCitation>
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;
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<materialsCitation id="4E7628171703FB59F586A012868BF878" collectingDate="1961-07-20" collectorName="Y. Hama" county="Tokara" location="Kagoshima Prefecture" municipality="Takara-jima" specimenCount="2">
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<specimenCount id="95CC0A6E6699120789D7ABB171EBC77A" type="generic">2 exs.</specimenCount>
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,
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<location id="55CC347648CBE9EDD9CC0E3403749B75" LSID="urn:lsid:plazi:treatment:F1B9F79A3CDF538C97C78C311848E4BC:55CC347648CBE9EDD9CC0E3403749B75" county="Tokara" municipality="Takara-jima" name="Kagoshima Prefecture">Kagoshima Prefecture</location>
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:
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<collectingMunicipality id="6AA386BA1C923D48928B46DE9CF300D5">Takara-jima</collectingMunicipality>
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,
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<collectingCounty id="EACB8ED3001ECDD358E34E5D70C1043F">Tokara</collectingCounty>
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,
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<collectingDate id="2E8AC9494C377772521CC4FB4FE18166" value="1961-07-20">20.VII.1961</collectingDate>
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,
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<collectorName id="4EE756758CA6CEF9C32BE4782EC82C72">Y. Hama</collectorName>
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leg. (EUMJ)
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</materialsCitation>
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;
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<materialsCitation id="A18575DA2890BCC985F870FF7D3795B7" collectingDate="1962-06-02" collectorName="M. Sato" county="Tokara" location="Kagoshima Prefecture" municipality="Takara-jima" specimenCount="7">
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<specimenCount id="ADE09A072CDB4D644CD79A993632EC16" type="generic">7 exs.</specimenCount>
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,
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<location id="BDADD894A7C97B23A7850AEA760166A5" LSID="urn:lsid:plazi:treatment:F1B9F79A3CDF538C97C78C311848E4BC:BDADD894A7C97B23A7850AEA760166A5" county="Tokara" municipality="Takara-jima" name="Kagoshima Prefecture">Kagoshima Prefecture</location>
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:
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<collectingMunicipality id="01ABE197371A41C331C0EF705754E87C">Takara-jima</collectingMunicipality>
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,
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<collectingCounty id="6C92E332406FCEFF1DD9C451A1FF973E">Tokara</collectingCounty>
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,
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<collectingDate id="28A45E4B1F3F8DA959DBA85C9AD04199" value="1962-06-02">2.VI.1962</collectingDate>
|
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,
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<collectorName id="4026BCADD921D7E6A9D19332386B1D20">
|
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M.
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<normalizedToken id="4EFD4758F2C5D268E79B97EF76573A69" originalValue="Satô">Sato</normalizedToken>
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</collectorName>
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leg. (EUMJ)
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</materialsCitation>
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;
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<materialsCitation id="88F0295BDB391C38E4340C8542D650F6" collectingDate="1975-09-04" collectorName="T. Takahashi" county="Ishigaki-jima" location="Okinawa Prefecture" municipality="Maesato" specimenCount="2">
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<specimenCount id="E461FBFAE9F6B04A2D5D03D38C1D144B" type="generic">2 exs.</specimenCount>
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,
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<location id="27346416323E28EA19F160F6F1E12E23" LSID="urn:lsid:plazi:treatment:F1B9F79A3CDF538C97C78C311848E4BC:27346416323E28EA19F160F6F1E12E23" county="Ishigaki-jima" municipality="Maesato" name="Okinawa Prefecture">Okinawa Prefecture</location>
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:
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<collectingMunicipality id="C8FD388C4D0351C40D8323B767A888AB">Maesato</collectingMunicipality>
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,
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<collectingCounty id="04CCAFAFA37ED71B99D4411D40E18315">Ishigaki-jima</collectingCounty>
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,
|
|
<collectingDate id="9787716E4C76F2FEABF41C36B4EE81A9" value="1975-09-04">4.IX.1975</collectingDate>
|
|
,
|
|
<collectorName id="7C63BB710D9942B6DF73DCB81551BC9F">T. Takahashi</collectorName>
|
|
leg. (EUMJ)
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="F0A7E6DC19313B8C486F3C16ACDD2931" collectingDate="1990-04-01" collectorName="Y. Uchida" county="Yonaguni-jima" location="Okinawa Prefecture" municipality="Mt. Urabu-dake" specimenCount="1">
|
|
<specimenCount id="6740AD740B716C812E0806285D3EEBC9" type="generic">1 ex.</specimenCount>
|
|
,
|
|
<location id="CC7C6B1D94EE2E38D3D8769BFBC269DF" LSID="urn:lsid:plazi:treatment:F1B9F79A3CDF538C97C78C311848E4BC:CC7C6B1D94EE2E38D3D8769BFBC269DF" county="Yonaguni-jima" municipality="Mt. Urabu-dake" name="Okinawa Prefecture">Okinawa Prefecture</location>
|
|
:
|
|
<collectingMunicipality id="2CC5C5ECA75798BBA27548270AD846E0">Mt. Urabu-dake</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="E778BEF9A60C0D7AF41EB367BEF3EFA7">Yonaguni-jima</collectingCounty>
|
|
,
|
|
<collectingDate id="134BA945A64A01C918300BC47174C941" value="1990-04-01">1.IV.1990</collectingDate>
|
|
,
|
|
<collectorName id="8BBC138BC0BBEA271F751631348AB0DF">Y. Uchida</collectorName>
|
|
leg. (EUMJ)
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="69E04BBBD7543D51D0D87E0B299D5585" collectingDate="2001-06-30" collectorName="T. Nakamura" county="Iriomote-jima" location="Okinawa Prefecture" municipality="Shirahama" specimenCount="1">
|
|
<specimenCount id="A9C3C48A3E6CC39627166738FD6D732D" type="generic">1 ex.</specimenCount>
|
|
,
|
|
<location id="DE3511CA6F074B018F2A2E4EB7FBB61E" LSID="urn:lsid:plazi:treatment:F1B9F79A3CDF538C97C78C311848E4BC:DE3511CA6F074B018F2A2E4EB7FBB61E" county="Iriomote-jima" municipality="Shirahama" name="Okinawa Prefecture">Okinawa Prefecture</location>
|
|
:
|
|
<collectingMunicipality id="B4268895E7D1CA489870DA65DF8101B1">Shirahama</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="A94E993D28EBC1840BF91118AA42E465">Iriomote-jima</collectingCounty>
|
|
,
|
|
<collectingDate id="349DE32AC261D314E3D5E9555511792E" value="2001-06-30">30.VI.2001</collectingDate>
|
|
,
|
|
<collectorName id="9B2252C9E709A7607486940A73F66B6E">T. Nakamura</collectorName>
|
|
leg. (TPM)
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="7250ECEF2AAB837D8608C5DBBEB11F99" collectingDate="1995-06-26" collectorName="T. Takara" county="Ryukyu" location="Okinawa Prefecture" municipality="Okinawa-jima" specimenCount="3">
|
|
<specimenCount id="2DC4EC3757EA71DD8543F89F19D9FECE" type="generic">3 exs.</specimenCount>
|
|
,
|
|
<location id="96880E410D1CC81A4BE2D0B91FECF4A7" LSID="urn:lsid:plazi:treatment:F1B9F79A3CDF538C97C78C311848E4BC:96880E410D1CC81A4BE2D0B91FECF4A7" county="Ryukyu" municipality="Okinawa-jima" name="Okinawa Prefecture">Okinawa Prefecture</location>
|
|
:
|
|
<collectingMunicipality id="1D3ED3EBF7FDF62699BA1D27F2D351C8">Okinawa-jima</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="926C0159D5A88B18C6EA1A3EC374BE59">Ryukyu</collectingCounty>
|
|
,
|
|
<collectingDate id="5632903A23862F9ECB3CE074DC7DF9D9" value="1995-06-26">26.VI.1995</collectingDate>
|
|
,
|
|
<collectorName id="000DC7AFEFF8139418184841BCD1B439">T. Takara</collectorName>
|
|
leg. (RUMF)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
<caption id="FB947F9C41EC5C2FCA4D433C991651CC" doi="10.3897/zookeys.1168.99302.figure14" httpUri="https://binary.pensoft.net/fig/872274" pageId="0" pageNumber="267" start="Figure 14" startId="F14">
|
|
<paragraph id="7E21922BE1F59CFA3A85EBADDC0F1BE2" pageId="0" pageNumber="267">
|
|
<emphasis id="9B4A413CA879E6DC762D74174E589CD5" bold="true" pageId="0" pageNumber="267">Figure 14.</emphasis>
|
|
<taxonomicName id="8A487637B32F9F4B127163D6FD41DFAC" authorityName="Kano & Kamiya" authorityYear="1931" class="Insecta" family="Characeae" genus="Haliplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haliplus kotoshonis" order="Coleoptera" pageId="0" pageNumber="267" phylum="Arthropoda" rank="species" species="kotoshonis">
|
|
<emphasis id="A2FB9F891229B620EA7A25872D6A5899" italics="true" pageId="0" pageNumber="267">Haliplus kotoshonis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="C6BA7D895C03F2B1D315EABFD8DB716F" bold="true" pageId="0" pageNumber="267">A</emphasis>
|
|
habitus
|
|
<emphasis id="8CD9AC2882E777B04BC41BA80EFD6337" bold="true" pageId="0" pageNumber="267">B</emphasis>
|
|
head
|
|
<emphasis id="9F8C96AA745C43D9CBEAE9637853CF22" bold="true" pageId="0" pageNumber="267">C</emphasis>
|
|
prosternal process
|
|
<emphasis id="6CDB671F81ADBD99D1185962A621FA48" bold="true" pageId="0" pageNumber="267">D</emphasis>
|
|
penis
|
|
<emphasis id="05A444027E40DA929288D9209CC94710" bold="true" pageId="0" pageNumber="267">E</emphasis>
|
|
left paramere
|
|
<emphasis id="0AF22BFA4B16D7885A084061C7CEB943" bold="true" pageId="0" pageNumber="267">F</emphasis>
|
|
right paramere. Scale bars: 1.0 mm (
|
|
<emphasis id="EE1DF578FD553126DDC92833E4F56794" bold="true" pageId="0" pageNumber="267">A</emphasis>
|
|
);
|
|
<quantity id="FD5AB61EE8E5A9F30A2540BDC92CE0C2" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
|
|
(
|
|
<emphasis id="F21CE4BE60993C403EC067C8B16C48F0" bold="true" pageId="0" pageNumber="267">B</emphasis>
|
|
);
|
|
<quantity id="BE7AFD7BBA84A3513C9BDEF6CEE9EAC0" metricMagnitude="-4" metricUnit="m" metricValue="2.5" unit="mm" value="0.25">0.25 mm</quantity>
|
|
(
|
|
<emphasis id="F8C6520E2C1F356D6F5FC39EDE209902" bold="true" pageId="0" pageNumber="267">C</emphasis>
|
|
);
|
|
<quantity id="DEAB35BA12C32AE4BBED5E3920AD4D94" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
|
|
(
|
|
<emphasis id="0ADF89165FF50108AEAF3EA3928F44AC" bold="true" pageId="0" pageNumber="267">D-F</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="B64361DCE5C31DF9743908308FD66C68" pageId="0" pageNumber="267" type="description">
|
|
<paragraph id="D18112BCBEE77EE3A12CB512D7BC0FD8" pageId="0" pageNumber="267">Measurements</paragraph>
|
|
<paragraph id="F0AC6235BFE7168B0BB04D0795FA62E9" pageId="0" pageNumber="267">
|
|
<emphasis id="DB50AEA1177DE5938167B95AEC0E225F" bold="true" pageId="0" pageNumber="267">
|
|
(
|
|
<emphasis id="9FB4390AA5BC40E72E724C822A417AA7" italics="true" pageId="0" pageNumber="267">n</emphasis>
|
|
= 10).
|
|
</emphasis>
|
|
TL 3.39-4.94 (4.47) mm; HW 0.74-0.87 (0.80) mm; CED 0.31-0.38 (0.33) mm; PL 0.65-0.79 (0.73) mm; PW 1.29-1.53 (1.42) mm; EL 2.30-2.62 (2.42) mm; EW 1.76-2.01 (1.88) mm; BT 1.39-1.70 (1.50) mm; HW/CED 2.30-2.56 (2.38); PW/PL 1.87-2.04 (1.95); EL/EW 1.23-1.32 (1.30).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="73EABFC47612D971EE946DFAF2C5B946" pageId="0" pageNumber="267" type="biology_ecology">
|
|
<paragraph id="7152BC78852AC6F26E35F0ABBE0B0E43" pageId="0" pageNumber="267">Biology.</paragraph>
|
|
<paragraph id="5B8788C059EB791829663DA739B1185B" pageId="0" pageNumber="267">The above specimens were collected from small ponds.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="0FEE4392DF1FFB2C4731480C05A8D3F0" pageId="0" pageNumber="267" type="immature stages">
|
|
<paragraph id="EF499109597A67EC07C432456585C9EE" pageId="0" pageNumber="267">Immature stages.</paragraph>
|
|
<paragraph id="8F1335584C06C47AFC56A80795FC091E" pageId="0" pageNumber="267">Unknown.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="448D1AE59B42BA6887C533CC991289A7" pageId="0" pageNumber="267" type="discussion">
|
|
<paragraph id="A7A78AEA455CD246374F4193758517B2" pageId="0" pageNumber="267">Discussion.</paragraph>
|
|
<paragraph id="004A3F109464E14B5AF1142C185FC769" pageId="0" pageNumber="267">
|
|
This species was described based on one specimen from
|
|
<normalizedToken id="44F017BB3A0A6EC67137A9CF92CE8A2C" originalValue="“Kotosho”">"Kotosho"</normalizedToken>
|
|
(Orchid Island), Taiwan (
|
|
<bibRefCitation id="ACA079E4C56B55EEE540AD7241CD41D6" author="Kano, T" journalOrPublisher="Transactions of the Kansai Entomological Society" pageId="0" pageNumber="267" pagination="1 - 4" refId="B18" refString="Kano, T, Kamiya, K, 1931. Two new species of Haliplidae from Japan. Transactions of the Kansai Entomological Society 2: 1 - 4" title="Two new species of Haliplidae from Japan." volume="2" year="1931">Kano and Kamiya 1931</bibRefCitation>
|
|
). The type specimen is likely to have been lost (
|
|
<bibRefCitation id="F362831ABDCCF09202C98D1628A8EF4D" author="Mita, T" journalOrPublisher="Journal of Agricultural Science, Tokyo University of Agriculture" pageId="0" pageNumber="267" pagination="50 - 56" refId="B22" refString="Mita, T, Kojima, H, Okajima, S, 2015. Name-bearing types of Coleoptera (Insecta) in the Museum of Agriculture, Tokyo University of Agriculture. Journal of Agricultural Science, Tokyo University of Agriculture 60: 50 - 56" title="Name-bearing types of Coleoptera (Insecta) in the Museum of Agriculture, Tokyo University of Agriculture." volume="60" year="2015">Mita et al. 2015</bibRefCitation>
|
|
), but the black marks on the dorsum are unique and the original description and figures are recognizable.
|
|
<bibRefCitation id="6145BBFA8E2F3E90095134E5CA3A2162" author="Nakane, T" journalOrPublisher="Kita-Kyushu no Konchu" pageId="0" pageNumber="267" pagination="61 - 67" refId="B29" refString="Nakane, T, 1985. A revisional study on Japanese species of the genus Haliplus Latreille (Coleoptera, Haliplidae). Kita-Kyushu no Konchu 32: 61 - 67" title="A revisional study on Japanese species of the genus Haliplus Latreille (Coleoptera, Haliplidae)." volume="32" year="1985">Nakane (1985</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="C50CFF783100DF4C578994C72E42E3AF" author="Nakane, T" journalOrPublisher="Insects & Nature" pageId="0" pageNumber="267" pagination="26 - 30" refId="B30" refString="Nakane, T, 1987. The beetles of Japan (new series) 80. Insects & Nature 22 (11): 26 - 30" title="The beetles of Japan (new series) 80." volume="22" year="1987">1987</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="19ED28BF2856341F7DF6DBC2413E07CD" author="Nakane, T" journalOrPublisher="Insects & Nature" pageId="0" pageNumber="267" pagination="22 - 23" refId="B31" refString="Nakane, T, 1990. The beetles of Japan (new series) 91. Insects & Nature 25 (10): 22 - 23" title="The beetles of Japan (new series) 91." volume="25" year="1990">1990</bibRefCitation>
|
|
) has repeatedly pointed out that
|
|
<taxonomicName id="D04996A5EB69CED9DDFC710B64F78BBD" authorityName="J.Balfour-Browne" authorityYear="1947" class="Insecta" family="Characeae" genus="Haliplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haliplus diruptus" order="Coleoptera" pageId="0" pageNumber="267" phylum="Arthropoda" rank="species" species="diruptus">
|
|
<emphasis id="34306A465D9B9D3C923AAEC249645652" italics="true" pageId="0" pageNumber="267">Haliplus diruptus</emphasis>
|
|
</taxonomicName>
|
|
J. Balfour-Browne, 1946 should be a junior synonym of the species, but a formal treatment has not been done. Judging from our investigation of the specimens and the description of
|
|
<bibRefCitation id="8E7D617F050FA3FE5CB619EE595562B5" author="van Vondel, BJ" journalOrPublisher="In: Jaech MA, Ji L (Eds) Water Beetles of China" pageId="0" pageNumber="267" pagination="111 - 154" refId="B48" refString="van Vondel, BJ, 1995. Haliplidae: Review of the Haliplidae of China. In: Jaech MA, Ji L (Eds) Water Beetles of China 1: 111 - 154" title="Haliplidae: Review of the Haliplidae of China." volume="1" year="1995">van Vondel (1995)</bibRefCitation>
|
|
,
|
|
<taxonomicName id="70B2B52D5B9920447EB376508165063D" family="Haliplidae" kingdom="Animalia" lsidName="H. diruptus" order="Coleoptera" pageId="0" pageNumber="267" rank="species" species="diruptus">
|
|
<emphasis id="01C5EA538DAE210421E3936A649958FB" italics="true" pageId="0" pageNumber="267">H. diruptus</emphasis>
|
|
</taxonomicName>
|
|
is treated as a junior synonym of this species in this paper. In addition,
|
|
<taxonomicName id="9948B55E5590F1590EA0B982A56A9B9F" authorityName="Vondel" authorityYear="1991" class="Insecta" family="Characeae" genus="Haliplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haliplus davidi" order="Coleoptera" pageId="0" pageNumber="267" phylum="Arthropoda" rank="species" species="davidi">
|
|
<emphasis id="7DCF9F7BB0372F99F3FD5FF56B90400A" italics="true" pageId="0" pageNumber="267">Haliplus davidi</emphasis>
|
|
</taxonomicName>
|
|
Vondel, 1991 was recorded from Japan (
|
|
<bibRefCitation id="E9F04E2CAD9774683D36F64FFEBA1ECD" author="van Vondel, BJ" journalOrPublisher="In: Jaech MA, Ji L (Eds) Water Beetles of China" pageId="0" pageNumber="267" pagination="131 - 136" refId="B49" refString="van Vondel, BJ, 1998. Haliplidae: Additional notes on the Haliplidae of China and neighboring countries (Coleoptera). In: Jaech MA, Ji L (Eds) Water Beetles of China 2: 131 - 136" title="Haliplidae: Additional notes on the Haliplidae of China and neighboring countries (Coleoptera)." volume="2" year="1998">van Vondel 1998</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="B56BEC48B62D2600B14F43C2734927F7" author="van Vondel, BJ" editor="Loebl, I" journalOrPublisher="1). Apollo Books, Vester Skerninge" pageId="0" pageNumber="267" pagination="30 - 33" refId="B50" refString="van Vondel, BJ, 2003a. Haliplidae. In: Loebl, I, Smetana, S, Eds., Catalogue of Palaearctic Coleoptera (Vol. 1). Apollo Books, Vester Skerninge: 30 - 33" title="Haliplidae." volumeTitle="Catalogue of Palaearctic Coleoptera (Vol." year="2003 a">2003a</bibRefCitation>
|
|
), but was later treated as a junior synonym of
|
|
<taxonomicName id="D80F7FC152107C5222A933BF310272ED" family="Haliplidae" kingdom="Animalia" lsidName="H. diruptus" order="Coleoptera" pageId="0" pageNumber="267" rank="species" species="diruptus">
|
|
<emphasis id="352D35CB8DEA95A4C70BA9815A006852" italics="true" pageId="0" pageNumber="267">H. diruptus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation id="85380A0E2BBEF232C0F8D6FA750303F9" author="van Vondel, BJ" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="267" pagination="31 - 35" refId="B52" refString="van Vondel, BJ, 2017. Five new synonymies in Haliplus subgen. Liaphlus Guignot, based on the variability of the left paramere (Coleoptera: Haliplidae). Koleopterologische Rundschau 87: 31 - 35" title="Five new synonymies in Haliplus subgen. Liaphlus Guignot, based on the variability of the left paramere (Coleoptera: Haliplidae)." volume="87" year="2017">van Vondel 2017</bibRefCitation>
|
|
); therefore, the record of
|
|
<taxonomicName id="F18AC8A0FB61A6E02EE3938AB1AFADFF" family="Haliplidae" kingdom="Animalia" lsidName="H. davidi" order="Coleoptera" pageId="0" pageNumber="267" rank="species" species="davidi">
|
|
<emphasis id="29921E5B43736F943E314A5A6CD4C822" italics="true" pageId="0" pageNumber="267">H. davidi</emphasis>
|
|
</taxonomicName>
|
|
from Japan is not
|
|
<taxonomicName id="EF5D3359D67AE1E41D1798B4FB67E6F1" family="Haliplidae" kingdom="Animalia" lsidName="H. diruptus" order="Coleoptera" pageId="0" pageNumber="267" rank="species" species="diruptus">
|
|
<emphasis id="E566CBFDCA85CB5C3C747500DD09954B" italics="true" pageId="0" pageNumber="267">H. diruptus</emphasis>
|
|
</taxonomicName>
|
|
but
|
|
<taxonomicName id="4DDDDD9C0A45060D140C599511F0100E" family="Haliplidae" kingdom="Animalia" lsidName="H. kotoshonis" order="Coleoptera" pageId="0" pageNumber="267" rank="species" species="kotoshonis">
|
|
<emphasis id="6F5209CC4E14914FFCC0FA3395AA5EB1" italics="true" pageId="0" pageNumber="267">H. kotoshonis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="17D7BA1C0EC40476B6767FB7FE50E069" pageId="0" pageNumber="267">
|
|
This study revealed that specimens of
|
|
<taxonomicName id="2BB9ABA39FDCEBF59EE2DCEB533647F6" authorityName="Re ́ gimbart" authorityYear="1892" class="Insecta" family="Characeae" genus="Haliplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haliplus angustifrons" order="Coleoptera" pageId="0" pageNumber="267" phylum="Arthropoda" rank="species" species="angustifrons">
|
|
<emphasis id="CD87E68A006B2F84830294CD60A38A54" italics="true" pageId="0" pageNumber="267">Haliplus angustifrons</emphasis>
|
|
</taxonomicName>
|
|
Régimbart, 1892 were mixed in with the specimens identified as "
|
|
<taxonomicName id="6CBB0DE501A22E66A49B9A2B3FFF7EBF" family="Haliplidae" kingdom="Animalia" lsidName="H. kotoshonis" order="Coleoptera" pageId="0" pageNumber="267" rank="species" species="kotoshonis">
|
|
<emphasis id="4FF7351C7B774254B59944ADFB97940D" italics="true" pageId="0" pageNumber="267">H. kotoshonis</emphasis>
|
|
</taxonomicName>
|
|
" from Japan. In a typical individual, the dorsal marks can easily distinguish between the two species. In particular, this species has a black mark on head, but the
|
|
<taxonomicName id="87C1B624523D4B7A8BBEDAFD64D5CA06" family="Haliplidae" kingdom="Animalia" lsidName="H. angustifrons" order="Coleoptera" pageId="0" pageNumber="267" rank="species" species="angustifrons">
|
|
<emphasis id="E15DE9A6EA99DFFEC2655F5F2D82BE0C" italics="true" pageId="0" pageNumber="267">H. angustifrons</emphasis>
|
|
</taxonomicName>
|
|
does not have such a mark.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="62EB5C69C66A321A2A7CF4129E9E85E3" pageId="0" pageNumber="267" type="distribution">
|
|
<paragraph id="FA6E6ED5FD5C3EDBC86177365F814B6E" pageId="0" pageNumber="267">Distribution.</paragraph>
|
|
<paragraph id="B1B427C0CAC259009C3FB33902E7AC18" pageId="0" pageNumber="267">Japan: Nansei shoto (Takara-jima of Tokara-retto, Okinawa-jima, Ishigaki-jima, Iriomote-jima, Yonaguni-jima); Taiwan, China, Vietnam, Myanmar.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |