446 lines
66 KiB
XML
446 lines
66 KiB
XML
<document id="9FDA3B8A4654C3D28BCBBBBFFB9AAA48" ID-CLB-Dataset="20150" ID-DOI="10.11646/zootaxa.5116.4.7" ID-GBIF-Dataset="8517cc18-c1bd-4838-b6b2-6ae35bb74c34" ID-ISSN="1175-5326" ID-Zenodo-Dep="6375446" ID-ZooBank="DD7FA75E-9D8D-425B-BD39-A3891BF423F0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647941196103" checkinUser="plazi" docAuthor="Aswathy, Mathilakath Dasan & Sudhikumar, Ambalaparambil Vasu" docDate="2022" docId="150087DB4D7B2F37FCC1FF117CF05B9B" docLanguage="en" docName="zootaxa.5116.4.7.pdf" docOrigin="Zootaxa 5116 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Propyrgodesmus bulbogonopus Aswathy & Sudhikumar 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="597" masterDocId="E939FFA34D792F31FC56FFC8796C5C62" masterDocTitle="Review of the millipede genus Propyrgodesmus Silvestri, 1920, with the description of a new species from a sacred grove in Kerala, India (Diplopoda: Polydesmida Pyrgodesmidae)" masterLastPageNumber="599" masterPageNumber="591" pageNumber="593" updateTime="1699318449368" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E3CE1D8AC318D64C8C60E64B90EB0FE7">Review of the millipede genus Propyrgodesmus Silvestri, 1920, with the description of a new species from a sacred grove in Kerala, India (Diplopoda: Polydesmida Pyrgodesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="22AE5BFE2B6C3A2DE96A5D5A31CCFB99" type="personal">
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<mods:role id="4972FEA4455B5972995E35730F230E7C">
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<mods:roleTerm id="F42BC33AE54D7C76CF55945A3BE3DE75">Author</mods:roleTerm>
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<mods:namePart id="FDF35726062E729BC0AD54341B834474">Aswathy, Mathilakath Dasan</mods:namePart>
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<mods:nameIdentifier id="184840CF8BD249C41BB062B1D21BC8F9" type="ORCID">0000-0002-3904-8066</mods:nameIdentifier>
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<mods:affiliation id="C9EB82A4BD31DE460D9AC7513E5E3964">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Kerala, India. & aswathym. das 94 @ gmail. com; https: // orcid. org / 0000 - 0002 - 3904 - 8066</mods:affiliation>
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<mods:nameIdentifier id="298C23EA997E5DD8EE69E4727F8C416D" type="email">aswathym.das94@gmail.com</mods:nameIdentifier>
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<mods:name id="2EB774A6BB7DC3815303841D5DE54334" type="personal">
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<mods:roleTerm id="4A4D33C0F988EABDB6E5D7EC7DA4D11F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="97B789AAAAC65BE76702FFDF26744DDF">Sudhikumar, Ambalaparambil Vasu</mods:namePart>
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<mods:affiliation id="AA3E7EF3A2CB991A7B4A4B1893FCAE63">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Kerala, India.</mods:affiliation>
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<mods:typeOfResource id="8B47538D61082441D14ABFEF1AAED1E0">text</mods:typeOfResource>
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<mods:title id="DF808092A2251A1F657EB3225F5A8E10">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3F2689B54F8C0C90235F6393CEB3CFA9">
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<mods:date id="90F9F4684BCCBE765BA06105E9163B0E">2022</mods:date>
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<mods:number id="FF1732429F719310228F975F67F2A7DE">2022-03-22</mods:number>
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<mods:number id="9A63BBD63E28FFC5B96D46178B926FBD">5116</mods:number>
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<mods:detail id="EC92371539A34C96C7FC1C3484CD7461" type="issue">
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<mods:number id="0AF5FC948B002261B248E5513DC383D1">4</mods:number>
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<mods:start id="6974C35BE30AB52344C4A5820E197FC6">591</mods:start>
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<mods:classification id="EF31CE831B1EC1A2A62D0312FB8D9E1C">journal article</mods:classification>
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<mods:identifier id="0FE4025FF6C3A668581048B0A4CD8BEA" type="CLB-Dataset">20150</mods:identifier>
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<mods:identifier id="2E6CA7333C65252EC7EBCA952520A12D" type="DOI">10.11646/zootaxa.5116.4.7</mods:identifier>
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<mods:identifier id="98C4438FD55F8DABB20AFC226AA9DEB2" type="GBIF-Dataset">8517cc18-c1bd-4838-b6b2-6ae35bb74c34</mods:identifier>
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<mods:identifier id="23C04913B8A72BA7DD5EDA932A8345E4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="150087DB4D7B2F37FCC1FF117CF05B9B" ID-DOI="http://doi.org/10.5281/zenodo.6377305" ID-GBIF-Taxon="194203837" ID-Zenodo-Dep="6377305" LSID="urn:lsid:plazi:treatment:150087DB4D7B2F37FCC1FF117CF05B9B" httpUri="http://treatment.plazi.org/id/150087DB4D7B2F37FCC1FF117CF05B9B" lastPageId="6" lastPageNumber="597" pageId="2" pageNumber="593">
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<subSubSection id="D5B365464D7B2F33FCC1FF117B155C96" box="[151,633,217,244]" pageId="2" pageNumber="593" type="nomenclature">
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<paragraph id="9D1636CD4D7B2F33FCC1FF117B155C96" blockId="2.[151,633,217,280]" box="[151,633,217,244]" pageId="2" pageNumber="593">
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<heading id="C65E81A14D7B2F33FCC1FF117B155C96" bold="true" box="[151,633,217,244]" fontSize="11" level="1" pageId="2" pageNumber="593" reason="1">
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<emphasis id="AFDDEADF4D7B2F33FCC1FF117B155C96" bold="true" box="[151,633,217,244]" pageId="2" pageNumber="593">
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<taxonomicName id="5AA94D4E4D7B2F33FCC1FF117B785C96" authority="Aswathy & Sudhikumar, 2022" authorityName="Aswathy & Sudhikumar" authorityYear="2022" box="[151,532,217,244]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="bulbogonopus" status="sp. nov.">
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<emphasis id="AFDDEADF4D7B2F33FCC1FF117B785C96" bold="true" box="[151,532,217,244]" italics="true" pageId="2" pageNumber="593">Propyrgodesmus bulbogonopus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B4EE57A44D7B2F33FE4DFF127B155C96" box="[539,633,218,244]" pageId="2" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D5B365464D7B2F33FCC1FF3679955D7A" box="[151,249,254,280]" pageId="2" pageNumber="593" type="description">
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<paragraph id="9D1636CD4D7B2F33FCC1FF3679955D7A" blockId="2.[151,633,217,280]" box="[151,249,254,280]" pageId="2" pageNumber="593">
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<figureCitation id="05922A484D7B2F33FCC1FF3679955D7A" box="[151,249,254,280]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1613,1637]" captionStartId-1="3.[151,250,1942,1966]" captionStartId-2="4.[151,250,1866,1890]" captionStartId-3="5.[151,250,1909,1933]" captionTargetBox-0="[151,1436,548,1587]" captionTargetBox-1="[164,1423,273,1919]" captionTargetBox-2="[151,1435,501,1843]" captionTargetBox-3="[242,1357,252,1882]" captionTargetId-0="figure-104@2.[151,1436,548,1587]" captionTargetId-1="figure-59@3.[164,1423,273,1919]" captionTargetId-2="figure-161@4.[151,1435,453,1843]" captionTargetId-3="figure-17@5.[191,1396,181,1885]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Propyrgodesmus bulbogonopus sp. nov., habitus. A, C. Holotype ♂ (CATE-5503A), dorsal and ventral views, respectively. B. Female paratype (CATE-5503C), dorsal view. Scale bar: 1 mm." captionText-1="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." captionText-2="FIGURE 3. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A. Left gonopod, posterior view. B. Right gonopod, anterior view. C, D. Both gonopods, posterior and anterior views, respectively. Abbreviations: b: medial branch; cx: coxite; h: hook like structure of telopodite; sph: solenophore; sl: solenomere. Scale bar: 0.2 mm (for C and D). A, B not to scale." captionText-3="FIGURE 4. Propyrgodesmus spp. A–C. Propyrgodesmus lobulatus Silvestri, 1920, holotype ♀ after Silvestri (1920). A, B. Anterior and posterior parts of the body, dorsal and lateral views, respectively. C. Right antenna, anterior view. D–F. Propyrgodesmus frater Carl, 1932 syntype ♂, after Carl (1932). D, E. Gonopods, anterior and posterior views, respectively. F. Antenna. G. Propyrgodesmus bulbogonopus sp. nov., holotype ♂ (CATE-5503A), right antenna, anterior view. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.6375448" figureDoi-1="http://doi.org/10.5281/zenodo.6375450" figureDoi-2="http://doi.org/10.5281/zenodo.6375452" figureDoi-3="http://doi.org/10.5281/zenodo.6375454" httpUri-0="https://zenodo.org/record/6375448/files/figure.png" httpUri-1="https://zenodo.org/record/6375450/files/figure.png" httpUri-2="https://zenodo.org/record/6375452/files/figure.png" httpUri-3="https://zenodo.org/record/6375454/files/figure.png" pageId="2" pageNumber="593">Figs 1–4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D5B365464D7B2F33FCC1FE887D995E6C" pageId="2" pageNumber="593" type="materials_examined">
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<paragraph id="9D1636CD4D7B2F33FCC1FE8878945DC0" blockId="2.[151,1436,320,527]" pageId="2" pageNumber="593">
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<emphasis id="AFDDEADF4D7B2F33FCC1FE8878EE5D38" bold="true" box="[151,386,320,346]" pageId="2" pageNumber="593">Material examined.</emphasis>
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<materialsCitation id="2DC13C904D7B2F33FDDCFE8978945DC0" ID-GBIF-Occurrence="3712376302" collectingDate="2021-08-10" collectorName="M. D. Aswathy" country="India" county="Muchukunnu" elevation="9" latitude="11.495" location="Sacred" longLatPrecision="21" longitude="75.665276" municipality="Kottayil Siva Kshethram" pageId="2" pageNumber="593" specimenCount="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="holotype">
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<emphasis id="AFDDEADF4D7B2F33FDDCFE89789B5D39" bold="true" box="[394,503,321,347]" pageId="2" pageNumber="593">
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<typeStatus id="4212886F4D7B2F33FDDCFE89789B5D39" box="[394,503,321,347]" pageId="2" pageNumber="593" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
<specimenCount id="8BAFFD444D7B2F33FE56FE897B795D39" box="[512,533,321,347]" pageId="2" pageNumber="593" type="male">♂</specimenCount>
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(CATE-5503A),
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<location id="987660164D7B2F33FE8DFE887A4B5D38" LSID="urn:lsid:plazi:treatment:150087DB4D7B2F37FCC1FF117CF05B9B:987660164D7B2F33FE8DFE887A4B5D38" box="[731,807,320,346]" country="India" county="Muchukunnu" latitude="11.495" longLatPrecision="21" longitude="75.665276" municipality="Kottayil Siva Kshethram" name="Sacred" pageId="2" pageNumber="593" stateProvince="Kerala">Sacred</location>
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groves of
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<collectingMunicipality id="7D72ACB74D7B2F33FFF5FE887DD35D38" box="[931,1215,320,346]" pageId="2" pageNumber="593">Kottayil Siva Kshethram</collectingMunicipality>
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,
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<collectingCounty id="74774E414D7B2F33F89AFE887C085D38" box="[1228,1380,320,346]" pageId="2" pageNumber="593">Muchukunnu</collectingCounty>
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, Kozhikode district,
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<collectingRegion id="5F6DF82F4D7B2F33FD03FEAC78CE5D1C" box="[341,418,356,382]" country="India" name="Kerala" pageId="2" pageNumber="593">Kerala</collectingRegion>
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,
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<collectingCountry id="E5BE765D4D7B2F33FDF9FEAC78855D1C" box="[431,489,356,382]" name="India" pageId="2" pageNumber="593">India</collectingCountry>
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<geoCoordinate id="F89D500A4D7B2F33FDA6FEAD7B135D1D" box="[496,639,357,383]" degrees="11" direction="north" minutes="29" orientation="latitude" pageId="2" pageNumber="593" precision="15" seconds="42" value="11.495">11°29’42’’N</geoCoordinate>
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,
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<geoCoordinate id="F89D500A4D7B2F33FEDAFEAC7A745D1D" box="[652,792,356,383]" degrees="75" direction="east" minutes="39" orientation="longitude" pageId="2" pageNumber="593" precision="15" seconds="55" value="75.665276">75°39’55’’E</geoCoordinate>
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,
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<elevation id="1684D1FE4D7B2F33FF73FEAD7AE35D1C" box="[805,911,356,383]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="593" unit="m" value="9.0">
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<quantity id="5A519B284D7B2F33FF73FEAD7A3D5D1D" box="[805,849,357,383]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="593" unit="m" value="9.0">9 m</quantity>
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a.s.l.
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</elevation>
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, collected from rotting wood, hand collection,
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<date id="E917100D4D7B2F33FCC1FE41787B5DC1" box="[151,279,393,419]" pageId="2" pageNumber="593" value="2021-08-10">
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<collectingDate id="F953E9E54D7B2F33FCC1FE41787B5DC1" box="[151,279,393,419]" pageId="2" pageNumber="593" value="2021-08-10">08.10.2021</collectingDate>
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</date>
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,
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<collectorName id="305C531B4D7B2F33FD75FE4178A45DC0" box="[291,456,392,419]" pageId="2" pageNumber="593">M.D. Aswathy</collectorName>
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leg.
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</materialsCitation>
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</paragraph>
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<paragraph id="9D1636CD4D7B2F33FC91FE657A7D5D88" blockId="2.[151,1436,320,527]" pageId="2" pageNumber="593">
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<emphasis id="AFDDEADF4D7B2F33FC91FE6578245DA5" bold="true" box="[199,328,429,455]" pageId="2" pageNumber="593">
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<typeStatus id="4212886F4D7B2F33FC91FE65782F5DA5" box="[199,323,429,455]" pageId="2" pageNumber="593" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCount id="8BAFFD444D7B2F33FD19FE6578155DA5" box="[335,377,429,455]" pageId="2" pageNumber="593" type="male">1 ♂</specimenCount>
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(CATE-5503B),
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<specimenCount id="8BAFFD444D7B2F33FE6CFE657B155DA5" box="[570,633,429,455]" pageId="2" pageNumber="593" type="female">2 ♀♀</specimenCount>
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(CATE-5503C, D),
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<materialsCitation id="2DC13C904D7B2F33FF0AFE657A625D88" ID-GBIF-Occurrence="3712376301" pageId="2" pageNumber="593" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
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<specimenCount id="8BAFFD444D7B2F33FF0AFE657AF65DA5" box="[860,922,429,455]" pageId="2" pageNumber="593" type="female">3 ♀♀</specimenCount>
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juveniles (CATE-5503E, F, G), same locality, hand-collected together with
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<typeStatus id="4212886F4D7B2F33FD89FE187B285D88" box="[479,580,464,490]" pageId="2" pageNumber="593" type="holotype">holotype</typeStatus>
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, but from top soil
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="9D1636CD4D7B2F33FC91FE3D7D995E6C" blockId="2.[151,1436,320,527]" box="[199,1269,500,527]" pageId="2" pageNumber="593">
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<emphasis id="AFDDEADF4D7B2F33FC91FE3D78785E6D" bold="true" box="[199,276,501,527]" pageId="2" pageNumber="593">Name.</emphasis>
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To emphasize the bulbous appearance of the medial branch of the gonopodal telopodite.
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</paragraph>
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</subSubSection>
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<caption id="C9D666454D7B2F33FCC1F9857AAB5AEB" ID-DOI="http://doi.org/10.5281/zenodo.6375448" ID-Zenodo-Dep="6375448" httpUri="https://zenodo.org/record/6375448/files/figure.png" pageId="2" pageNumber="593" startId="2.[151,250,1613,1637]" targetBox="[151,1436,548,1587]" targetPageId="2">
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<paragraph id="9D1636CD4D7B2F33FCC1F9857AAB5AEB" blockId="2.[151,1436,1613,1674]" pageId="2" pageNumber="593">
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||
<emphasis id="AFDDEADF4D7B2F33FCC1F985787A5A07" bold="true" box="[151,278,1613,1637]" pageId="2" pageNumber="593">FIGURE 1.</emphasis>
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<taxonomicName id="5AA94D4E4D7B2F33FD48F9857B0D5A07" authority="Aswathy & Sudhikumar, 2022" authorityName="Aswathy & Sudhikumar" authorityYear="2022" box="[286,609,1613,1637]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="bulbogonopus" status="sp. nov.">
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<emphasis id="AFDDEADF4D7B2F33FD48F9857B0D5A07" box="[286,609,1613,1637]" italics="true" pageId="2" pageNumber="593">Propyrgodesmus bulbogonopus</emphasis>
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</taxonomicName>
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<emphasis id="AFDDEADF4D7B2F33FE3CF9857BD15A07" bold="true" box="[618,701,1613,1637]" pageId="2" pageNumber="593">
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||
<taxonomicNameLabel id="B4EE57A44D7B2F33FE3CF9857BD15A07" box="[618,701,1613,1637]" pageId="2" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, habitus.
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<emphasis id="AFDDEADF4D7B2F33FF73F9857A335A07" bold="true" box="[805,863,1613,1637]" pageId="2" pageNumber="593">A, C.</emphasis>
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Holotype ♂ (CATE-5503A), dorsal and ventral views, respectively.
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<emphasis id="AFDDEADF4D7B2F33FD4BF9B978585AEB" bold="true" box="[285,308,1649,1673]" pageId="2" pageNumber="593">B.</emphasis>
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Female paratype (CATE-5503C), dorsal view.
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<emphasis id="AFDDEADF4D7B2F33FF44F9B97A135AEB" bold="true" box="[786,895,1649,1673]" pageId="2" pageNumber="593">Scale bar:</emphasis>
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1 mm.
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</paragraph>
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</caption>
|
||
<subSubSection id="D5B365464D7B2F33FC91F97F7A5A5BCB" pageId="2" pageNumber="593" type="diagnosis">
|
||
<paragraph id="9D1636CD4D7B2F33FC91F97F7A5A5BCB" blockId="2.[151,1437,1719,2034]" pageId="2" pageNumber="593">
|
||
<emphasis id="AFDDEADF4D7B2F33FC91F97F782E5AB3" bold="true" box="[199,322,1719,1745]" pageId="2" pageNumber="593">Diagnosis.</emphasis>
|
||
When comparing to
|
||
<emphasis id="AFDDEADF4D7B2F33FE78F9707BD55AB3" box="[558,697,1719,1745]" italics="true" pageId="2" pageNumber="593">
|
||
<taxonomicName id="5AA94D4E4D7B2F33FE78F9707BD95AB3" authorityName="Silvestri" authorityYear="1920" box="[558,693,1719,1745]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="lobulatus">P. lobulatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the new species shows major differences in the tergal ornamentation by having most paraterga only vaguely trilobulate laterally (
|
||
<figureCitation id="05922A484D7B2F33FF33F9137AB75A94" box="[869,987,1755,1782]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="2" pageNumber="593">Fig. 2B, E</figureCitation>
|
||
),
|
||
<emphasis id="AFDDEADF4D7B2F33FFB9F9147D6A5A97" box="[1007,1030,1756,1781]" italics="true" pageId="2" pageNumber="593">vs</emphasis>
|
||
clearly trilobate (
|
||
<figureCitation id="05922A484D7B2F33F89DF9137C2E5A94" box="[1227,1346,1755,1782]" captionStart="FIGURE 4" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[242,1357,252,1882]" captionTargetId="figure-17@5.[191,1396,181,1885]" captionTargetPageId="5" captionText="FIGURE 4. Propyrgodesmus spp. A–C. Propyrgodesmus lobulatus Silvestri, 1920, holotype ♀ after Silvestri (1920). A, B. Anterior and posterior parts of the body, dorsal and lateral views, respectively. C. Right antenna, anterior view. D–F. Propyrgodesmus frater Carl, 1932 syntype ♂, after Carl (1932). D, E. Gonopods, anterior and posterior views, respectively. F. Antenna. G. Propyrgodesmus bulbogonopus sp. nov., holotype ♂ (CATE-5503A), right antenna, anterior view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6375454" httpUri="https://zenodo.org/record/6375454/files/figure.png" pageId="2" pageNumber="593">Fig. 4A, B</figureCitation>
|
||
) and by the presence of a ventral cone on antennomere 4 bearing a long sensillum trichodeum (
|
||
<figureCitation id="05922A484D7B2F33F823F9377DBE5B78" box="[1141,1234,1791,1818]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="2" pageNumber="593">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="05922A484D7B2F33F889F8C87C685B78" box="[1247,1284,1792,1818]" captionStart="FIGURE 4" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[242,1357,252,1882]" captionTargetId="figure-17@5.[191,1396,181,1885]" captionTargetPageId="5" captionText="FIGURE 4. Propyrgodesmus spp. A–C. Propyrgodesmus lobulatus Silvestri, 1920, holotype ♀ after Silvestri (1920). A, B. Anterior and posterior parts of the body, dorsal and lateral views, respectively. C. Right antenna, anterior view. D–F. Propyrgodesmus frater Carl, 1932 syntype ♂, after Carl (1932). D, E. Gonopods, anterior and posterior views, respectively. F. Antenna. G. Propyrgodesmus bulbogonopus sp. nov., holotype ♂ (CATE-5503A), right antenna, anterior view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6375454" httpUri="https://zenodo.org/record/6375454/files/figure.png" pageId="2" pageNumber="593">4G</figureCitation>
|
||
),
|
||
<emphasis id="AFDDEADF4D7B2F33F94CF8C87C5D5B7B" box="[1306,1329,1792,1817]" italics="true" pageId="2" pageNumber="593">vs</emphasis>
|
||
a ventral cone on antennomere 4 absent (
|
||
<figureCitation id="05922A484D7B2F33FE50F8EB7B0D5B5C" box="[518,609,1827,1854]" captionStart="FIGURE 4" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[242,1357,252,1882]" captionTargetId="figure-17@5.[191,1396,181,1885]" captionTargetPageId="5" captionText="FIGURE 4. Propyrgodesmus spp. A–C. Propyrgodesmus lobulatus Silvestri, 1920, holotype ♀ after Silvestri (1920). A, B. Anterior and posterior parts of the body, dorsal and lateral views, respectively. C. Right antenna, anterior view. D–F. Propyrgodesmus frater Carl, 1932 syntype ♂, after Carl (1932). D, E. Gonopods, anterior and posterior views, respectively. F. Antenna. G. Propyrgodesmus bulbogonopus sp. nov., holotype ♂ (CATE-5503A), right antenna, anterior view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6375454" httpUri="https://zenodo.org/record/6375454/files/figure.png" pageId="2" pageNumber="593">Fig. 4C</figureCitation>
|
||
). On the contrary, the new species shows some similarities to the tergal ornamentation of
|
||
<taxonomicName id="5AA94D4E4D7B2F33FD34F88078D35B03" authorityName="Carl" authorityYear="1932" box="[354,447,1863,1889]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="frater">
|
||
<emphasis id="AFDDEADF4D7B2F33FD34F88078D35B03" box="[354,447,1863,1889]" italics="true" pageId="2" pageNumber="593">P. frater</emphasis>
|
||
</taxonomicName>
|
||
, but differs significantly in the biramous gonopod with a bulbous lateral branch of the telopodite (
|
||
<figureCitation id="05922A484D7B2F33FD40F8A378E15BE4" box="[278,397,1899,1926]" captionStart="FIGURE 3" captionStartId="4.[151,250,1866,1890]" captionTargetBox="[151,1435,501,1843]" captionTargetId="figure-161@4.[151,1435,453,1843]" captionTargetPageId="4" captionText="FIGURE 3. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A. Left gonopod, posterior view. B. Right gonopod, anterior view. C, D. Both gonopods, posterior and anterior views, respectively. Abbreviations: b: medial branch; cx: coxite; h: hook like structure of telopodite; sph: solenophore; sl: solenomere. Scale bar: 0.2 mm (for C and D). A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.6375452" httpUri="https://zenodo.org/record/6375452/files/figure.png" pageId="2" pageNumber="593">Fig. 3A, B</figureCitation>
|
||
)
|
||
<emphasis id="AFDDEADF4D7B2F33FDCAF8A478DF5BE7" box="[412,435,1900,1925]" italics="true" pageId="2" pageNumber="593">vs</emphasis>
|
||
with prong—and lobe-like structures on the main branch of the telopodite (
|
||
<figureCitation id="05922A484D7B2F33F8A9F8A37C145BE4" box="[1279,1400,1899,1926]" captionStart="FIGURE 4" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[242,1357,252,1882]" captionTargetId="figure-17@5.[191,1396,181,1885]" captionTargetPageId="5" captionText="FIGURE 4. Propyrgodesmus spp. A–C. Propyrgodesmus lobulatus Silvestri, 1920, holotype ♀ after Silvestri (1920). A, B. Anterior and posterior parts of the body, dorsal and lateral views, respectively. C. Right antenna, anterior view. D–F. Propyrgodesmus frater Carl, 1932 syntype ♂, after Carl (1932). D, E. Gonopods, anterior and posterior views, respectively. F. Antenna. G. Propyrgodesmus bulbogonopus sp. nov., holotype ♂ (CATE-5503A), right antenna, anterior view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6375454" httpUri="https://zenodo.org/record/6375454/files/figure.png" pageId="2" pageNumber="593">Fig. 4D, E</figureCitation>
|
||
). A key to all three existing species of the genus is given below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5B365464D7B2F37FC91F87B7D0C5B05" lastPageId="6" lastPageNumber="597" pageId="2" pageNumber="593" type="description">
|
||
<paragraph id="9D1636CD4D7B2F32FC91F87B7BBE5C9B" blockId="2.[151,1437,1719,2034]" lastBlockId="3.[151,1437,150,249]" lastPageId="3" lastPageNumber="594" pageId="2" pageNumber="593">
|
||
<emphasis id="AFDDEADF4D7B2F33FC91F87B783D5BAF" bold="true" box="[199,337,1971,1997]" pageId="2" pageNumber="593">Description</emphasis>
|
||
.
|
||
<typeStatus id="4212886F4D7B2F33FD09F87B78AB5BAF" box="[351,455,1971,1997]" pageId="2" pageNumber="593" type="holotype">Holotype</typeStatus>
|
||
ca
|
||
<quantity id="5A519B284D7B2F33FDBAF87B7B2D5BAC" box="[492,577,1971,1998]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="593" unit="mm" value="7.6">7.6 mm</quantity>
|
||
long, width of mid-body pro—and metazonae 0.78 and
|
||
<quantity id="5A519B284D7B2F33F8F8F87B7C695BAC" box="[1198,1285,1971,1998]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="593" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, respectively. Male
|
||
<typeStatus id="4212886F4D7B2F33FC8EF81078545B90" box="[216,312,2008,2034]" pageId="2" pageNumber="593" type="paratype">paratype</typeStatus>
|
||
ca
|
||
<quantity id="5A519B284D7B2F33FD09F81F78D45B90" box="[351,440,2007,2034]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="593" unit="mm" value="7.5">7.5 mm</quantity>
|
||
, width of pro—and metazonae 0.78 and
|
||
<quantity id="5A519B284D7B2F33FFD7F81F7AB55B90" box="[897,985,2007,2034]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="593" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, respectively. Adult female
|
||
<typeStatus id="4212886F4D7B2F33F947F8107C105B90" box="[1297,1404,2008,2034]" pageId="2" pageNumber="593" type="paratype">paratypes</typeStatus>
|
||
ca 7.5 and
|
||
<quantity id="5A519B284D7A2F32FCA4FF5F78585CD3" box="[242,308,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="594" unit="mm" value="9.0">9 mm</quantity>
|
||
long, width of pro—and metazonae 0.76–0.85 and
|
||
<quantity id="5A519B284D7A2F32FF2BFF5E7D6B5CD3" box="[893,1031,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="3" pageNumber="594" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
|
||
, respectively. Live colouration dark brown, dorsum encrusted with soil particles; antennae, legs and venter pale whitish (
|
||
<figureCitation id="05922A484D7A2F32F800FF737DBD5CB7" box="[1110,1233,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1613,1637]" captionTargetBox="[151,1436,548,1587]" captionTargetId="figure-104@2.[151,1436,548,1587]" captionTargetPageId="2" captionText="FIGURE 1. Propyrgodesmus bulbogonopus sp. nov., habitus. A, C. Holotype ♂ (CATE-5503A), dorsal and ventral views, respectively. B. Female paratype (CATE-5503C), dorsal view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6375448" httpUri="https://zenodo.org/record/6375448/files/figure.png" pageId="3" pageNumber="594">Fig. 1A, B</figureCitation>
|
||
). Both vertex and frons light brown, clypeolabrum whitish (
|
||
<figureCitation id="05922A484D7A2F32FE3CFF177BA95C9B" box="[618,709,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="3" pageNumber="594">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="C9D666454D7A2F32FCC1F85E7D9D5B94" ID-DOI="http://doi.org/10.5281/zenodo.6375450" ID-Zenodo-Dep="6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="3" pageNumber="594" startId="3.[151,250,1942,1966]" targetBox="[164,1423,273,1919]" targetPageId="3">
|
||
<paragraph id="9D1636CD4D7A2F32FCC1F85E7D9D5B94" blockId="3.[151,1437,1942,2038]" pageId="3" pageNumber="594">
|
||
<emphasis id="AFDDEADF4D7A2F32FCC1F85E787E5BCC" bold="true" box="[151,274,1942,1966]" pageId="3" pageNumber="594">FIGURE 2.</emphasis>
|
||
<taxonomicName id="5AA94D4E4D7A2F32FD41F85E7B3A5BCC" authority="Aswathy & Sudhikumar, 2022" authorityName="Aswathy & Sudhikumar" authorityYear="2022" box="[279,598,1942,1966]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="bulbogonopus" status="sp. nov.">
|
||
<emphasis id="AFDDEADF4D7A2F32FD41F85E7B3A5BCC" box="[279,598,1942,1966]" italics="true" pageId="3" pageNumber="594">Propyrgodesmus bulbogonopus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AFDDEADF4D7A2F32FE0DF85E7BC65BCC" bold="true" box="[603,682,1942,1966]" pageId="3" pageNumber="594">
|
||
<taxonomicNameLabel id="B4EE57A44D7A2F32FE0DF85E7BC65BCC" box="[603,682,1942,1966]" pageId="3" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype ♂ (CATE-5503B).
|
||
<emphasis id="AFDDEADF4D7A2F32FF84F85E7D6B5BCC" bold="true" box="[978,1031,1942,1966]" pageId="3" pageNumber="594">A, B.</emphasis>
|
||
Anterior part of body, ventral and dorsal views, respectively.
|
||
<emphasis id="AFDDEADF4D7A2F32FD32F87278115BB0" bold="true" box="[356,381,1978,2002]" pageId="3" pageNumber="594">C.</emphasis>
|
||
Right antenna, anterior view.
|
||
<emphasis id="AFDDEADF4D7A2F32FEF8F8727B8C5BB0" bold="true" box="[686,736,1978,2002]" pageId="3" pageNumber="594">D–F.</emphasis>
|
||
Posterior part of body, ventral, dorsal and lateral views, respectively.
|
||
<emphasis id="AFDDEADF4D7A2F32FCC1F81679DD5B94" bold="true" box="[151,177,2014,2038]" pageId="3" pageNumber="594">G.</emphasis>
|
||
Right midbody leg, anterior view.
|
||
<emphasis id="AFDDEADF4D7A2F32FE42F8167BE75B94" bold="true" box="[532,651,2014,2038]" pageId="3" pageNumber="594">Scale bars:</emphasis>
|
||
0.5 mm (
|
||
<emphasis id="AFDDEADF4D7A2F32FEBBF8167A575B94" bold="true" box="[749,827,2014,2038]" pageId="3" pageNumber="594">A, B, E</emphasis>
|
||
), 0.4 mm (
|
||
<emphasis id="AFDDEADF4D7A2F32FFFAF8167AAA5B94" bold="true" box="[940,966,2014,2038]" pageId="3" pageNumber="594">D)</emphasis>
|
||
, 0.2 mm (
|
||
<emphasis id="AFDDEADF4D7A2F32F879F8167D265B94" bold="true" box="[1071,1098,2014,2038]" pageId="3" pageNumber="594">C)</emphasis>
|
||
, 0.1 mm (
|
||
<emphasis id="AFDDEADF4D7A2F32F8E5F8167DAE5B94" bold="true" box="[1203,1218,2014,2038]" pageId="3" pageNumber="594">F</emphasis>
|
||
,
|
||
<emphasis id="AFDDEADF4D7A2F32F899F8167D8F5B94" bold="true" box="[1231,1251,2014,2038]" pageId="3" pageNumber="594">G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9D1636CD4D7D2F35FC91FF5F7D6E5DCF" blockId="4.[151,1437,151,429]" pageId="4" pageNumber="595">
|
||
Body broad due to large paraterga (
|
||
<figureCitation id="05922A484D7D2F35FE34FF5F7BD15CD3" box="[610,701,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1613,1637]" captionTargetBox="[151,1436,548,1587]" captionTargetId="figure-104@2.[151,1436,548,1587]" captionTargetPageId="2" captionText="FIGURE 1. Propyrgodesmus bulbogonopus sp. nov., habitus. A, C. Holotype ♂ (CATE-5503A), dorsal and ventral views, respectively. B. Female paratype (CATE-5503C), dorsal view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6375448" httpUri="https://zenodo.org/record/6375448/files/figure.png" pageId="4" pageNumber="595">Fig. 1A</figureCitation>
|
||
), with 20 body rings in both sexes (
|
||
<figureCitation id="05922A484D7D2F35F837FF5F7DB35CD3" box="[1121,1247,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1613,1637]" captionTargetBox="[151,1436,548,1587]" captionTargetId="figure-104@2.[151,1436,548,1587]" captionTargetPageId="2" captionText="FIGURE 1. Propyrgodesmus bulbogonopus sp. nov., habitus. A, C. Holotype ♂ (CATE-5503A), dorsal and ventral views, respectively. B. Female paratype (CATE-5503C), dorsal view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6375448" httpUri="https://zenodo.org/record/6375448/files/figure.png" pageId="4" pageNumber="595">Fig. 1A, B</figureCitation>
|
||
). In width, head << collum ≤ 2 <3–16, body tapering thereafter towards telson (
|
||
<figureCitation id="05922A484D7D2F35FF20FF737A985CB7" box="[886,1012,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1613,1637]" captionTargetBox="[151,1436,548,1587]" captionTargetId="figure-104@2.[151,1436,548,1587]" captionTargetPageId="2" captionText="FIGURE 1. Propyrgodesmus bulbogonopus sp. nov., habitus. A, C. Holotype ♂ (CATE-5503A), dorsal and ventral views, respectively. B. Female paratype (CATE-5503C), dorsal view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6375448" httpUri="https://zenodo.org/record/6375448/files/figure.png" pageId="4" pageNumber="595">Fig. 1A–C</figureCitation>
|
||
). Head ovoid, slightly transverse and sparsely setose in clypeolabral region, granulate above antennal sockets either side of epicranial suture. Genae subquadrate and granulate. Interantennal isthmus twice as wide as diameter of antennal socket. Antenna short, weakly clavate, reaching up to body ring 3 when stretched ventrolaterally. In length, antennomere 7<6<1<3<4<2<<5 (
|
||
<figureCitation id="05922A484D7D2F35F932FEEF79B05D07" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="4" pageNumber="595">Figs. 2A, C</figureCitation>
|
||
,
|
||
<figureCitation id="05922A484D7D2F35FCB0FE8378615D07" box="[230,269,331,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[242,1357,252,1882]" captionTargetId="figure-17@5.[191,1396,181,1885]" captionTargetPageId="5" captionText="FIGURE 4. Propyrgodesmus spp. A–C. Propyrgodesmus lobulatus Silvestri, 1920, holotype ♀ after Silvestri (1920). A, B. Anterior and posterior parts of the body, dorsal and lateral views, respectively. C. Right antenna, anterior view. D–F. Propyrgodesmus frater Carl, 1932 syntype ♂, after Carl (1932). D, E. Gonopods, anterior and posterior views, respectively. F. Antenna. G. Propyrgodesmus bulbogonopus sp. nov., holotype ♂ (CATE-5503A), right antenna, anterior view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6375454" httpUri="https://zenodo.org/record/6375454/files/figure.png" pageId="4" pageNumber="595">4G</figureCitation>
|
||
). Dense and sharply delimited fields of sensilla basiconica located distodorsaly on antennomeres 5 and 6. Sensory field of 5
|
||
<superScript id="6ADC9B854D7D2F35FD0DFEA678045D1E" attach="left" box="[347,360,366,380]" fontSize="6" pageId="4" pageNumber="595">th</superScript>
|
||
antennomere not extending up to base of long sensillum trichodeum.Antennomere 4 with ventral sensillum trichodeum borne on a small, but evident cone/hump (
|
||
<figureCitation id="05922A484D7D2F35FF31FE5B7AAF5DCF" box="[871,963,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="4" pageNumber="595">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="05922A484D7D2F35FF99FE5B7A995DCF" box="[975,1013,403,429]" captionStart="FIGURE 4" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[242,1357,252,1882]" captionTargetId="figure-17@5.[191,1396,181,1885]" captionTargetPageId="5" captionText="FIGURE 4. Propyrgodesmus spp. A–C. Propyrgodesmus lobulatus Silvestri, 1920, holotype ♀ after Silvestri (1920). A, B. Anterior and posterior parts of the body, dorsal and lateral views, respectively. C. Right antenna, anterior view. D–F. Propyrgodesmus frater Carl, 1932 syntype ♂, after Carl (1932). D, E. Gonopods, anterior and posterior views, respectively. F. Antenna. G. Propyrgodesmus bulbogonopus sp. nov., holotype ♂ (CATE-5503A), right antenna, anterior view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6375454" httpUri="https://zenodo.org/record/6375454/files/figure.png" pageId="4" pageNumber="595">4G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="C9D666454D7D2F35FCC1F88279BD5BAC" ID-DOI="http://doi.org/10.5281/zenodo.6375452" ID-Zenodo-Dep="6375452" httpUri="https://zenodo.org/record/6375452/files/figure.png" pageId="4" pageNumber="595" startId="4.[151,250,1866,1890]" targetBox="[151,1435,501,1843]" targetPageId="4">
|
||
<paragraph id="9D1636CD4D7D2F35FCC1F88279BD5BAC" blockId="4.[151,1436,1866,1998]" pageId="4" pageNumber="595">
|
||
<emphasis id="AFDDEADF4D7D2F35FCC1F88278635B00" bold="true" box="[151,271,1866,1890]" pageId="4" pageNumber="595">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="5AA94D4E4D7D2F35FD4BF8827B325B00" authority="Aswathy & Sudhikumar, 2022" authorityName="Aswathy & Sudhikumar" authorityYear="2022" box="[285,606,1866,1890]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="bulbogonopus" status="sp. nov.">
|
||
<emphasis id="AFDDEADF4D7D2F35FD4BF8827B325B00" box="[285,606,1866,1890]" italics="true" pageId="4" pageNumber="595">Propyrgodesmus bulbogonopus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AFDDEADF4D7D2F35FE30F8827BD45B00" bold="true" box="[614,696,1866,1890]" pageId="4" pageNumber="595">
|
||
<taxonomicNameLabel id="B4EE57A44D7D2F35FE30F8827BD45B00" box="[614,696,1866,1890]" pageId="4" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype ♂ (CATE-5503B).
|
||
<emphasis id="AFDDEADF4D7D2F35FFBDF8827D685B00" bold="true" box="[1003,1028,1866,1890]" pageId="4" pageNumber="595">A.</emphasis>
|
||
Left gonopod, posterior view.
|
||
<emphasis id="AFDDEADF4D7D2F35F913F8827C305B00" bold="true" box="[1349,1372,1866,1890]" pageId="4" pageNumber="595">B.</emphasis>
|
||
Right gonopod, anterior view.
|
||
<emphasis id="AFDDEADF4D7D2F35FDDBF8A678A85BE4" bold="true" box="[397,452,1902,1926]" pageId="4" pageNumber="595">C, D.</emphasis>
|
||
Both gonopods, posterior and anterior views, respectively.
|
||
<emphasis id="AFDDEADF4D7D2F35F84DF8A67DB55BE4" bold="true" box="[1051,1241,1902,1926]" pageId="4" pageNumber="595">Abbreviations: b:</emphasis>
|
||
medial branch;
|
||
<emphasis id="AFDDEADF4D7D2F35F92AF8A67CF05BE4" bold="true" box="[1404,1436,1902,1926]" pageId="4" pageNumber="595">cx:</emphasis>
|
||
coxite;
|
||
<emphasis id="AFDDEADF4D7D2F35FCB2F85A799E5BC8" bold="true" box="[228,242,1938,1962]" pageId="4" pageNumber="595">h</emphasis>
|
||
: hook like structure of telopodite;
|
||
<emphasis id="AFDDEADF4D7D2F35FE0CF85A7BE45BC8" bold="true" box="[602,648,1938,1962]" pageId="4" pageNumber="595">sph:</emphasis>
|
||
solenophore;
|
||
<emphasis id="AFDDEADF4D7D2F35FF4DF85A7A5F5BC8" bold="true" box="[795,819,1938,1962]" pageId="4" pageNumber="595">sl:</emphasis>
|
||
solenomere.
|
||
<emphasis id="AFDDEADF4D7D2F35FFE8F85A7D415BC8" bold="true" box="[958,1069,1938,1962]" pageId="4" pageNumber="595">Scale bar:</emphasis>
|
||
0.2 mm (for
|
||
<emphasis id="AFDDEADF4D7D2F35F8EFF85A7DA75BC8" bold="true" box="[1209,1227,1938,1962]" pageId="4" pageNumber="595">C</emphasis>
|
||
and
|
||
<emphasis id="AFDDEADF4D7D2F35F8A9F85A7C7D5BC8" bold="true" box="[1279,1297,1938,1962]" pageId="4" pageNumber="595">D</emphasis>
|
||
).
|
||
<emphasis id="AFDDEADF4D7D2F35F97EF85A7C355BC8" bold="true" box="[1320,1369,1938,1962]" pageId="4" pageNumber="595">A, B</emphasis>
|
||
not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C9D666454D7C2F34FCC1F8BD7C6D5B9B" ID-DOI="http://doi.org/10.5281/zenodo.6375454" ID-Zenodo-Dep="6375454" httpUri="https://zenodo.org/record/6375454/files/figure.png" pageId="5" pageNumber="596" startId="5.[151,250,1909,1933]" targetBox="[242,1357,252,1882]" targetPageId="5">
|
||
<paragraph id="9D1636CD4D7C2F34FCC1F8BD7C6D5B9B" blockId="5.[151,1436,1909,2041]" pageId="5" pageNumber="596">
|
||
<emphasis id="AFDDEADF4D7C2F34FCC1F8BD787E5BEF" bold="true" box="[151,274,1909,1933]" pageId="5" pageNumber="596">FIGURE 4.</emphasis>
|
||
<taxonomicName id="5AA94D4E4D7C2F34FD4EF8BD7B665BEF" box="[280,522,1909,1933]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="undefined-a">
|
||
<emphasis id="AFDDEADF4D7C2F34FD4EF8BD78AE5BEF" box="[280,450,1909,1933]" italics="true" pageId="5" pageNumber="596">Propyrgodesmus</emphasis>
|
||
spp.
|
||
<emphasis id="AFDDEADF4D7C2F34FDA1F8BD7B665BEF" bold="true" box="[503,522,1909,1933]" pageId="5" pageNumber="596">A</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AFDDEADF4D7C2F34FE5CF8BD7B435BEF" bold="true" box="[522,559,1909,1933]" pageId="5" pageNumber="596">–C.</emphasis>
|
||
<taxonomicName id="5AA94D4E4D7C2F34FE62F8BD7AB15BEF" authority="Silvestri, 1920" authorityName="Silvestri" authorityYear="1920" box="[564,989,1909,1933]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="lobulatus">
|
||
<emphasis id="AFDDEADF4D7C2F34FE62F8BD7A2D5BEF" box="[564,833,1909,1933]" italics="true" pageId="5" pageNumber="596">Propyrgodesmus lobulatus</emphasis>
|
||
<bibRefCitation id="F9384B3C4D7C2F34FF11F8BD7AB15BEF" author="Silvestri, F." box="[839,989,1909,1933]" pageId="5" pageNumber="596" pagination="117 - 135" refId="ref4139" refString="Silvestri, F. (1920) Descriptions of some Oriental Diplopoda Polydesmoidea of the subfamily Pyrgodesminae. Records of the Indian Museum, 19 (4), 117 - 135." type="journal article" year="1920">Silvestri, 1920</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype ♀ after
|
||
<bibRefCitation id="F9384B3C4D7C2F34F8C7F8BD7C5F5BEF" author="Silvestri, F." box="[1169,1331,1909,1933]" pageId="5" pageNumber="596" pagination="117 - 135" refId="ref4139" refString="Silvestri, F. (1920) Descriptions of some Oriental Diplopoda Polydesmoidea of the subfamily Pyrgodesminae. Records of the Indian Museum, 19 (4), 117 - 135." type="journal article" year="1920">Silvestri (1920)</bibRefCitation>
|
||
.
|
||
<emphasis id="AFDDEADF4D7C2F34F96DF8BD7C1C5BEF" bold="true" box="[1339,1392,1909,1933]" pageId="5" pageNumber="596">A, B.</emphasis>
|
||
Anterior and posterior parts of the body, dorsal and lateral views, respectively.
|
||
<emphasis id="AFDDEADF4D7C2F34FFC4F8517AC75BD3" bold="true" box="[914,939,1945,1969]" pageId="5" pageNumber="596">C.</emphasis>
|
||
Right antenna, anterior view.
|
||
<emphasis id="AFDDEADF4D7C2F34F88CF8517C605BD3" bold="true" box="[1242,1292,1945,1969]" pageId="5" pageNumber="596">D–F.</emphasis>
|
||
<taxonomicName id="5AA94D4E4D7C2F34F945F851781E5BB7" authority="Carl, 1932" authorityName="Carl" authorityYear="1932" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="frater">
|
||
<emphasis id="AFDDEADF4D7C2F34F945F851786D5BB7" italics="true" pageId="5" pageNumber="596">Propyrgodesmus frater</emphasis>
|
||
<bibRefCitation id="F9384B3C4D7C2F34FD51F875781E5BB7" author="Carl, J." box="[263,370,1981,2005]" pageId="5" pageNumber="596" pagination="411 - 529" refId="ref3654" refString="Carl, J. (1932) Diplopoden aus Sud-Indien und Ceylon. 1. Teil. Polydesmoidea. Revue suisse de Zoologie, 39 (17), 411 - 529. https: // doi. org / 10.5962 / bhl. part. 118948" type="journal article" year="1932">Carl, 1932</bibRefCitation>
|
||
</taxonomicName>
|
||
syntype ♂, after
|
||
<bibRefCitation id="F9384B3C4D7C2F34FE74F8757BF05BB7" author="Carl, J." box="[546,668,1981,2005]" pageId="5" pageNumber="596" pagination="411 - 529" refId="ref3654" refString="Carl, J. (1932) Diplopoden aus Sud-Indien und Ceylon. 1. Teil. Polydesmoidea. Revue suisse de Zoologie, 39 (17), 411 - 529. https: // doi. org / 10.5962 / bhl. part. 118948" type="journal article" year="1932">Carl (1932)</bibRefCitation>
|
||
.
|
||
<emphasis id="AFDDEADF4D7C2F34FEF3F8757BB75BB7" bold="true" box="[677,731,1981,2005]" pageId="5" pageNumber="596">D, E.</emphasis>
|
||
Gonopods, anterior and posterior views, respectively.
|
||
<emphasis id="AFDDEADF4D7C2F34F953F8757C755BB7" bold="true" box="[1285,1305,1981,2005]" pageId="5" pageNumber="596">F.</emphasis>
|
||
Antenna.
|
||
<emphasis id="AFDDEADF4D7C2F34F9D4F8757CF05BB7" bold="true" box="[1410,1436,1981,2005]" pageId="5" pageNumber="596">G.</emphasis>
|
||
<taxonomicName id="5AA94D4E4D7C2F34FCC1F82978BB5B9B" authority="Aswathy & Sudhikumar, 2022" authorityName="Aswathy & Sudhikumar" authorityYear="2022" box="[151,471,2017,2041]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="bulbogonopus" status="sp. nov.">
|
||
<emphasis id="AFDDEADF4D7C2F34FCC1F82978BB5B9B" box="[151,471,2017,2041]" italics="true" pageId="5" pageNumber="596">Propyrgodesmus bulbogonopus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AFDDEADF4D7C2F34FD8BF8297B425B9B" bold="true" box="[477,558,2017,2041]" pageId="5" pageNumber="596">
|
||
<taxonomicNameLabel id="B4EE57A44D7C2F34FD8BF8297B425B9B" box="[477,558,2017,2041]" pageId="5" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype ♂ (CATE-5503A), right antenna, anterior view. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C9D666454D7F2F37FCC1FBD07A0B5852" ID-DOI="http://doi.org/10.5281/zenodo.6375456" ID-Zenodo-Dep="6375456" box="[151,871,1048,1072]" httpUri="https://zenodo.org/record/6375456/files/figure.png" pageId="6" pageNumber="597" startId="6.[151,250,1048,1072]" targetBox="[218,1370,181,1024]" targetPageId="6">
|
||
<paragraph id="9D1636CD4D7F2F37FCC1FBD07A0B5852" blockId="6.[151,871,1048,1072]" box="[151,871,1048,1072]" pageId="6" pageNumber="597">
|
||
<emphasis id="AFDDEADF4D7F2F37FCC1FBD0787F5852" bold="true" box="[151,275,1048,1072]" pageId="6" pageNumber="597">FIGURE 5.</emphasis>
|
||
Distribution of the genus
|
||
<taxonomicName id="5AA94D4E4D7F2F37FE4AFBD07A085852" authority="Silvestri, 1920" authorityName="Silvestri" authorityYear="1920" box="[540,868,1048,1072]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="597" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AFDDEADF4D7F2F37FE4AFBD07BAA5852" box="[540,710,1048,1072]" italics="true" pageId="6" pageNumber="597">Propyrgodesmus</emphasis>
|
||
<bibRefCitation id="F9384B3C4D7F2F37FE9AFBD07A085852" author="Silvestri, F." box="[716,868,1048,1072]" pageId="6" pageNumber="597" pagination="117 - 135" refId="ref4139" refString="Silvestri, F. (1920) Descriptions of some Oriental Diplopoda Polydesmoidea of the subfamily Pyrgodesminae. Records of the Indian Museum, 19 (4), 117 - 135." type="journal article" year="1920">Silvestri, 1920</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9D1636CD4D7F2F37FC91FB9678DF5A67" blockId="6.[151,1437,1118,1829]" pageId="6" pageNumber="597">
|
||
Collum flabellate (
|
||
<figureCitation id="05922A484D7F2F37FDCDFB967B7B581B" box="[411,535,1118,1145]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="6" pageNumber="597">Fig. 2A, B</figureCitation>
|
||
), completely covering the head from above, fore margin with 5+5 distinct, subequal, faintly incised, rounded lobulations, middle with 4+4 distinct tubercles (
|
||
<figureCitation id="05922A484D7F2F37F847FB4A7D0658FE" box="[1041,1130,1154,1180]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="6" pageNumber="597">Fig. 2B</figureCitation>
|
||
). Dorsum strongly convex, paraterga lying below mid-body height, moderately sloping down at about 20–25°. Tegument covered with a microspiculate cerotegument. Postcollum paraterga thick and rather indistinctly lobulated laterally (
|
||
<figureCitation id="05922A484D7F2F37F88EFB027C3C5887" box="[1240,1360,1226,1253]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="6" pageNumber="597">Fig. 2B, E</figureCitation>
|
||
), three lobulations on 2
|
||
<superScript id="6ADC9B854D7F2F37FD1AFB267830589E" attach="left" box="[332,348,1262,1276]" fontSize="6" pageId="6" pageNumber="597">nd</superScript>
|
||
and pore-bearing rings,
|
||
<emphasis id="AFDDEADF4D7F2F37FE26FB277BEB596A" box="[624,647,1263,1288]" italics="true" pageId="6" pageNumber="597">vs</emphasis>
|
||
two lobes on poreless ones (
|
||
<figureCitation id="05922A484D7F2F37FF9DFB267D0C596B" box="[971,1120,1262,1289]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="6" pageNumber="597">Fig. 2B, E, F</figureCitation>
|
||
). Pore formula: 5, 9, 12, 15, 17–19, ozopores borne on porosteles behind second lobulation of paraterga (
|
||
<figureCitation id="05922A484D7F2F37FFBAFADA7D2F594E" box="[1004,1091,1298,1324]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="6" pageNumber="597">Fig. 2B</figureCitation>
|
||
). All paraterga starting with 5
|
||
<superScript id="6ADC9B854D7F2F37F9C6FADA7CF15942" attach="none" box="[1424,1437,1298,1312]" fontSize="6" pageId="6" pageNumber="597">th</superScript>
|
||
directed caudad. Metaterga behind collum with 2+2 longitudinal rows of 3+3 larger and fused tubercles, those before ring 19 slightly inclined caudad. Paramedian tubercle on ring 19 abruptly modified to form an elongated ridge overhanging and concealing the epiproct (
|
||
<figureCitation id="05922A484D7F2F37FE2EFAB67BBE59FA" box="[632,722,1406,1432]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="6" pageNumber="597">Fig. 2E</figureCitation>
|
||
). Epiproct directed ventrad, with four setae on top. Hypoproct triangular, caudal edge with two strong and well separated setae on knobs (
|
||
<figureCitation id="05922A484D7F2F37FFB6FA6A7D5759DE" box="[992,1083,1442,1468]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="6" pageNumber="597">Fig. 2D</figureCitation>
|
||
). Legs with neither adenostyles nor tarsal brushes, short, stout and enlarged, femur the longest (
|
||
<figureCitation id="05922A484D7F2F37FF26FA0E7AA05982" box="[880,972,1478,1504]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="6" pageNumber="597">Fig. 2G</figureCitation>
|
||
). In length, femur> prefemur> tarsus> postfemur> tibia> coxa.
|
||
</paragraph>
|
||
<paragraph id="9D1636CD4D7F2F37FC91F9C67C665B47" blockId="6.[151,1437,1118,1829]" pageId="6" pageNumber="597">
|
||
Gonopods taking up nearly entire ventral surface of metazonum 7, almost reaching the base of paraterga (
|
||
<figureCitation id="05922A484D7F2F37F939F9C679D15A2E" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[164,1423,273,1919]" captionTargetId="figure-59@3.[164,1423,273,1919]" captionTargetPageId="3" captionText="FIGURE 2. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A, B. Anterior part of body, ventral and dorsal views, respectively. C. Right antenna, anterior view. D–F. Posterior part of body, ventral, dorsal and lateral views, respectively. G. Right midbody leg, anterior view. Scale bars: 0.5 mm (A, B, E), 0.4 mm (D), 0.2 mm (C), 0.1 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6375450" httpUri="https://zenodo.org/record/6375450/files/figure.png" pageId="6" pageNumber="597">Fig. 2A</figureCitation>
|
||
). Gonopods relatively simple, placed inside a transversely oblong gonopodal aperture, with thin and slightly elevated caudal and lateral margins. Both gonopods held parallel to each other, strongly exposed, telopodite tips directed caudally. Coxites (
|
||
<emphasis id="AFDDEADF4D7F2F37FD91F9B2788D5AF6" bold="true" box="[455,481,1658,1684]" pageId="6" pageNumber="597">cx</emphasis>
|
||
) voluminous, transverse and microgranulate laterally (
|
||
<figureCitation id="05922A484D7F2F37F815F9B27DD75AF7" box="[1091,1211,1658,1685]" captionStart="FIGURE 3" captionStartId="4.[151,250,1866,1890]" captionTargetBox="[151,1435,501,1843]" captionTargetId="figure-161@4.[151,1435,453,1843]" captionTargetPageId="4" captionText="FIGURE 3. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A. Left gonopod, posterior view. B. Right gonopod, anterior view. C, D. Both gonopods, posterior and anterior views, respectively. Abbreviations: b: medial branch; cx: coxite; h: hook like structure of telopodite; sph: solenophore; sl: solenomere. Scale bar: 0.2 mm (for C and D). A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.6375452" httpUri="https://zenodo.org/record/6375452/files/figure.png" pageId="6" pageNumber="597">Fig. 3A, B</figureCitation>
|
||
). Gonocoel distinct, but not too deep; telopodites short, simple, biramous and distally carrying a long, in situ caudally directed and slightly bifurcate lamina with fringed/microspiculate lamellae; solenomere
|
||
<emphasis id="AFDDEADF4D7F2F37FFB2F90A7D655ABE" bold="true" box="[996,1033,1730,1756]" pageId="6" pageNumber="597">(sl)</emphasis>
|
||
slender and barely exposed; medial branch (
|
||
<emphasis id="AFDDEADF4D7F2F37FCA3F92E78695B62" bold="true" box="[245,261,1766,1792]" pageId="6" pageNumber="597">b</emphasis>
|
||
) stalked, with a short bulbous structure emerging distolaterally (
|
||
<figureCitation id="05922A484D7F2F37FFB4F92E7D335B63" box="[994,1119,1766,1793]" captionStart="FIGURE 3" captionStartId="4.[151,250,1866,1890]" captionTargetBox="[151,1435,501,1843]" captionTargetId="figure-161@4.[151,1435,453,1843]" captionTargetPageId="4" captionText="FIGURE 3. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A. Left gonopod, posterior view. B. Right gonopod, anterior view. C, D. Both gonopods, posterior and anterior views, respectively. Abbreviations: b: medial branch; cx: coxite; h: hook like structure of telopodite; sph: solenophore; sl: solenomere. Scale bar: 0.2 mm (for C and D). A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.6375452" httpUri="https://zenodo.org/record/6375452/files/figure.png" pageId="6" pageNumber="597">Fig. 3C, D</figureCitation>
|
||
). A laminar sheath covering the postfemoral part of telopodite with a caudally directed hook-shaped/unciform tip
|
||
<emphasis id="AFDDEADF4D7F2F37F807F8C27D185B46" bold="true" box="[1105,1140,1802,1828]" pageId="6" pageNumber="597">(h)</emphasis>
|
||
(
|
||
<figureCitation id="05922A484D7F2F37F8D5F8C27D905B47" box="[1155,1276,1802,1829]" captionStart="FIGURE 3" captionStartId="4.[151,250,1866,1890]" captionTargetBox="[151,1435,501,1843]" captionTargetId="figure-161@4.[151,1435,453,1843]" captionTargetPageId="4" captionText="FIGURE 3. Propyrgodesmus bulbogonopus sp. nov., paratype ♂ (CATE-5503B). A. Left gonopod, posterior view. B. Right gonopod, anterior view. C, D. Both gonopods, posterior and anterior views, respectively. Abbreviations: b: medial branch; cx: coxite; h: hook like structure of telopodite; sph: solenophore; sl: solenomere. Scale bar: 0.2 mm (for C and D). A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.6375452" httpUri="https://zenodo.org/record/6375452/files/figure.png" pageId="6" pageNumber="597">Fig. 3A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9D1636CD4D7F2F37FCC1F8857D0C5B05" blockId="6.[151,1120,1869,1895]" box="[151,1120,1869,1895]" pageId="6" pageNumber="597">
|
||
The following key can be proposed to separate the known species of
|
||
<taxonomicName id="5AA94D4E4D7F2F37FFCAF8857D345B05" authorityName="Silvestri" authorityYear="1920" box="[924,1112,1869,1895]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="597" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AFDDEADF4D7F2F37FFCAF8857D345B05" box="[924,1112,1869,1895]" italics="true" pageId="6" pageNumber="597">Propyrgodesmus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5B365464D7F2F37FCC1F8437CF05B9B" pageId="6" pageNumber="597" type="key">
|
||
<key id="E93861FC4D7F2F37FCC1F8437CF05B9B" pageId="6" pageNumber="597">
|
||
<keyStep id="265D2E684D7F2F37FCC1F8437CF05BDF" pageId="6" pageNumber="597">
|
||
<paragraph id="9D1636CD4D7F2F37FCC1F8437CF05BC2" blockId="6.[151,1437,1930,2041]" box="[151,1436,1930,1953]" pageId="6" pageNumber="597">
|
||
<keyLead id="265895F84D7F2F37FCC1F8437CF05BC2" box="[151,1436,1930,1953]" pageId="6" pageNumber="597">
|
||
1 Sensillum trichodeum of the 4
|
||
<superScript id="6ADC9B854D7F2F37FE52F8427B635BF4" attach="left" box="[516,527,1930,1942]" fontSize="5" pageId="6" pageNumber="597">th</superScript>
|
||
antennomere borne on a hump................................................. 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="9D1636CD4D7F2F37FCC1F8617CF05BDF" blockId="6.[151,1437,1930,2041]" box="[151,1436,1960,1982]" pageId="6" pageNumber="597">
|
||
<keyLead id="265895F84D7F2F37FCC1F8617CF05BDF" box="[151,1436,1960,1982]" pageId="6" pageNumber="597">
|
||
- Sensillum trichodeum of the 4
|
||
<superScript id="6ADC9B854D7F2F37FE52F8607B635BD6" attach="left" box="[516,527,1960,1972]" fontSize="5" pageId="6" pageNumber="597">th</superScript>
|
||
antennomere not on a hump...........................................
|
||
<taxonomicName id="5AA94D4E4D7F2F37F97DF8617CF05BDF" ID-CoL="789F9" authorityName="Silvestri" authorityYear="1920" box="[1323,1436,1960,1982]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="lobulatus">
|
||
<emphasis id="AFDDEADF4D7F2F37F97DF8617CF05BDF" box="[1323,1436,1960,1982]" italics="true" pageId="6" pageNumber="597">P. lobulatus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="265D2E684D7F2F37FCC1F80E7CF05B9B" pageId="6" pageNumber="597">
|
||
<paragraph id="9D1636CD4D7F2F37FCC1F80E7CF15BB9" blockId="6.[151,1437,1930,2041]" box="[151,1437,1990,2012]" pageId="6" pageNumber="597">
|
||
<keyLead id="265895F84D7F2F37FCC1F80E7CF15BB9" box="[151,1437,1990,2012]" pageId="6" pageNumber="597">
|
||
2 Gonopod telopodite with a laterally directed prong-like structure..........................................
|
||
<taxonomicName id="5AA94D4E4D7F2F37F918F80E7CF15BB9" ID-CoL="77XF9" authorityName="Carl" authorityYear="1932" box="[1358,1437,1990,2011]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="frater">
|
||
<emphasis id="AFDDEADF4D7F2F37F918F80E7CF15BB9" box="[1358,1437,1990,2011]" italics="true" pageId="6" pageNumber="597">P. frater</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="9D1636CD4D7F2F37FCC1F82C7CF05B9B" blockId="6.[151,1437,1930,2041]" box="[151,1436,2019,2041]" pageId="6" pageNumber="597">
|
||
<keyLead id="265895F84D7F2F37FCC1F82C7CF05B9B" box="[151,1436,2019,2041]" pageId="6" pageNumber="597">
|
||
- Gonopod telopodite with prominent bulb-like medial branch...............................
|
||
<taxonomicName id="5AA94D4E4D7F2F37F8FFF82C7C245B9A" authority="Aswathy & Sudhikumar, 2022" authorityName="Aswathy & Sudhikumar" authorityYear="2022" box="[1193,1352,2019,2041]" class="Diplopoda" family="Pyrgodesmidae" genus="Propyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="bulbogonopus" status="sp. nov.">
|
||
<emphasis id="AFDDEADF4D7F2F37F8FFF82C7C245B9A" box="[1193,1352,2019,2041]" italics="true" pageId="6" pageNumber="597">P. bulbogonopus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B4EE57A44D7F2F37F918F82B7CF05B9B" box="[1358,1436,2019,2041]" pageId="6" pageNumber="597" rank="species">
|
||
<emphasis id="AFDDEADF4D7F2F37F918F82B7CFA5B9A" bold="true" box="[1358,1430,2019,2040]" pageId="6" pageNumber="597">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |