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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFD5FF925AB6F8334587CEC5" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus hongiaoensis V. S. Dang &amp; Naiki 2021" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571720" ID-Zenodo-Dep="7571720" LSID="urn:lsid:plazi:treatment:43284262FFD5FF925AB6F8334587CEC5" httpUri="http://treatment.plazi.org/id/43284262FFD5FF925AB6F8334587CEC5" lastPageId="76" pageId="74">
<subSubSection box="[136,1151,1942,1969]" pageId="74" type="nomenclature">
<paragraph blockId="74.[136,1452,1942,2076]" box="[136,1151,1942,1969]" pageId="74">
<emphasis bold="true" box="[136,171,1942,1968]" pageId="74">32.</emphasis>
<taxonomicName authority="V. S. Dang &amp; Naiki (2021: 32)" authorityName="V. S. Dang &amp; Naiki" authorityPageNumber="32" authorityYear="2021" box="[178,802,1942,1969]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="74" phylum="Tracheophyta" rank="species" species="hongiaoensis">
<emphasis bold="true" box="[178,466,1942,1969]" italics="true" pageId="74">Lasianthus hongiaoensis</emphasis>
V.S.Dang &amp; Naiki (2021: 32)
</taxonomicName>
<figureCitation box="[833,910,1942,1969]" captionStart="FIGURE 55" captionStartId="75.[136,229,1781,1803]" captionTargetBox="[167,1418,194,1754]" captionTargetId="figure-17@75.[165,1418,193,1757]" captionTargetPageId="75" captionText="FIGURE 55. Lasianthus hongiaoensis V.S.Dang &amp; Naiki (from V4318, VNM): A. Branch with fruits, B. Close-up of expanding young leaves, C. Stipules, D. Abaxial leaf surfaces, E. Flower buds and young fruit, F. Inflorescence showing side view flowers, GH. Fruits, J. Seeds (EF: photos by Shuichiro Tagane)." figureDoi="http://doi.org/10.5281/zenodo.7571722" httpUri="https://zenodo.org/record/7571722/files/figure.png" pageId="74">Fig. 55</figureCitation>
(plate),
<figureCitation box="[1004,1077,1942,1969]" captionStart="FIGURE 52" captionStartId="71.[136,229,1789,1811]" captionTargetBox="[176,1411,204,1757]" captionTargetId="figure-17@71.[165,1418,193,1765]" captionTargetPageId="71" captionText="FIGURE 52. Distribution maps of Lasianthus hirsutus, L. hispidulus, L. honbaensis and L. hongiaoensis." figureDoi="http://doi.org/10.5281/zenodo.7571712" httpUri="https://zenodo.org/record/7571712/files/figure.png" pageId="74">Fig 52</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection lastPageId="76" pageId="74" type="materials_examined">
<paragraph blockId="74.[136,1452,1942,2076]" pageId="74">
<emphasis bold="true" box="[136,194,1982,2003]" pageId="74">Type:</emphasis>
<materialsCitation collectingDate="2016-02-24" collectorName="Tagane S. &amp; Nagamasu H. &amp; Naiki A. &amp; Dang V. S. &amp; Nguyen" country="Vietnam" elevation="1698" location="Bidoup Nui Ba National Park" pageId="74" specimenCount="1" stateProvince="Lam Dong">
<collectingCountry box="[219,332,1982,2003]" name="Vietnam" pageId="74">VIETNAM</collectingCountry>
.
<collectingRegion box="[340,536,1982,2004]" country="Vietnam" name="Lam Dong" pageId="74">Lam Dong Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFD5FF925AB6F8334587CEC5:CE5EA5AFFFD5FF94581FF81B431BCB9C" box="[545,829,1982,2004]" country="Vietnam" name="Bidoup Nui Ba National Park" pageId="74" stateProvince="Lam Dong">Bidoup Nui Ba National Park</location>
,
<quantity box="[838,910,1982,2004]" metricMagnitude="3" metricUnit="m" metricValue="1.698" pageId="74" unit="m" value="1698.0">
<elevation box="[838,910,1982,2004]" metricMagnitude="3" metricUnit="m" metricValue="1.698" pageId="74" unit="m" value="1698.0">1698 m</elevation>
</quantity>
elev.,
<date box="[971,1143,1982,2004]" pageId="74" value="2016-02-24">
<collectingDate box="[971,1143,1982,2004]" pageId="74" value="2016-02-24">24 February 2016</collectingDate>
</date>
,
<emphasis italics="true" pageId="74">
<collectorName box="[1152,1247,1982,2004]" pageId="74">Tagane S.</collectorName>
,
<collectorName box="[1257,1391,1983,2004]" pageId="74">Nagamasu H.</collectorName>
,
<collectorName pageId="74">Naiki A.</collectorName>
,
<collectorName box="[220,315,2018,2040]" pageId="74">Dang V.S.</collectorName>
,
<collectorName box="[322,395,2019,2040]" pageId="74">Nguyen</collectorName>
</emphasis>
</materialsCitation>
<materialsCitation collectionCode="VNM, FU" collectorName="Wai J. V" country="Japan" location="Iriomote Station" pageId="74" specimenCode="VNM00042898, VNM00042899, VNM00042900" specimenCount="1" typeStatus="holotype">
<emphasis box="[399,572,2018,2040]" italics="true" pageId="74">
<taxonomicNameLabel box="[399,442,2019,2040]" pageId="74" rank="variety">V.N.</taxonomicNameLabel>
,
<collectorName box="[449,525,2018,2040]" pageId="74">Wai J. V</collectorName>
4318
</emphasis>
(
<typeStatus box="[582,668,2018,2040]" pageId="74">holotype</typeStatus>
<collectionCode box="[671,732,2018,2039]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="74" type="Herbarium">VNM</collectionCode>
![
<specimenCode box="[745,905,2018,2039]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="74" type="Herbarium">VNM00042898</specimenCode>
]; isotypes
<collectionCode box="[1000,1037,2018,2039]" country="China" name="Fudan University, Department of Biology" pageId="74">FU</collectionCode>
!,
<collectionCode box="[1048,1109,2018,2039]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="74" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[1121,1278,2018,2039]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="74" type="Herbarium">VNM00042899</specimenCode>
,
<specimenCode box="[1285,1443,2018,2039]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="74" type="Herbarium">VNM00042900</specimenCode>
], the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFD5FF925AB6F8334587CEC5:CE5EA5AFFFD5FF945B5DF7A34226C454" box="[355,512,2054,2076]" country="Japan" name="Iriomote Station" pageId="74">Iriomote Station</location>
(
<collectingCountry box="[525,584,2054,2075]" name="Japan" pageId="74">Japan</collectingCountry>
))
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571722" ID-Zenodo-Dep="7571722" httpUri="https://zenodo.org/record/7571722/files/figure.png" pageId="75" startId="75.[136,229,1781,1803]" targetBox="[167,1418,194,1754]" targetPageId="75">
<paragraph blockId="75.[136,1452,1781,1875]" pageId="75">
<emphasis bold="true" box="[136,267,1781,1803]" pageId="75">FIGURE 55.</emphasis>
<taxonomicName authority="V. S. Dang &amp; Naiki" authorityName="V. S. Dang &amp; Naiki" box="[273,695,1781,1803]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="75" phylum="Tracheophyta" rank="species" species="hongiaoensis">
<emphasis box="[273,512,1781,1802]" italics="true" pageId="75">Lasianthus hongiaoensis</emphasis>
V.S.Dang &amp; Naiki
</taxonomicName>
(from
<emphasis box="[764,826,1781,1802]" italics="true" pageId="75">V4318</emphasis>
, VNM):
<emphasis bold="true" box="[916,940,1781,1802]" pageId="75">A.</emphasis>
Branch with fruits,
<emphasis bold="true" box="[1135,1158,1781,1802]" pageId="75">B.</emphasis>
Close-up of expanding young leaves,
<emphasis bold="true" box="[208,231,1817,1839]" pageId="75">C.</emphasis>
Stipules,
<emphasis bold="true" box="[326,349,1817,1838]" pageId="75">D.</emphasis>
Abaxial leaf surfaces,
<emphasis bold="true" box="[567,589,1817,1838]" pageId="75">E.</emphasis>
Flower buds and young fruit,
<emphasis bold="true" box="[878,897,1817,1838]" pageId="75">F.</emphasis>
Inflorescence showing side view flowers,
<emphasis bold="true" box="[1305,1360,1817,1839]" pageId="75">GH.</emphasis>
Fruits,
<emphasis bold="true" box="[1433,1452,1817,1838]" pageId="75">J.</emphasis>
Seeds (
<emphasis bold="true" box="[206,249,1853,1875]" pageId="75">EF</emphasis>
: photos by Shuichiro Tagane).
</paragraph>
</caption>
<paragraph blockId="75.[136,1451,1922,2057]" lastBlockId="76.[136,1452,158,653]" lastPageId="76" pageId="75">
<emphasis bold="true" box="[136,397,1922,1948]" pageId="75">
<collectingRegion box="[136,262,1922,1948]" pageId="75">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="2016-02-24" collectorName="Tagane S. &amp; Nagamasu H. &amp; Naiki A. &amp; Dang V. S. &amp; Nguyen" country="Vietnam" elevation="1698" latitude="12.164708" location="Bidoup Nui Ba National Park" longLatPrecision="1" longitude="108.53344" pageId="75" specimenCount="1" stateProvince="Lam Dong">
<collectingCountry box="[426,558,1922,1948]" name="Vietnam" pageId="75">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[571,672,1922,1948]" pageId="75">Central:</emphasis>
<collectingRegion box="[682,918,1922,1948]" country="Vietnam" name="Lam Dong" pageId="75">Lam Dong Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFD5FF925AB6F8334587CEC5:CE5EA5AFFFD4FF95599BF8274526CBD4" box="[933,1280,1922,1948]" country="Vietnam" latitude="12.164708" longLatPrecision="1" longitude="108.53344" name="Bidoup Nui Ba National Park" pageId="75" stateProvince="Lam Dong">Bidoup Nui Ba National Park</location>
,
<quantity box="[1294,1383,1922,1948]" metricMagnitude="3" metricUnit="m" metricValue="1.698" pageId="75" unit="m" value="1698.0">
<elevation box="[1294,1383,1922,1948]" metricMagnitude="3" metricUnit="m" metricValue="1.698" pageId="75" unit="m" value="1698.0">1698 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[136,315,1958,1985]" degrees="12" direction="north" minutes="09" orientation="latitude" pageId="75" precision="1" seconds="52.95" value="12.164708">12°0952.95N</geoCoordinate>
,
<geoCoordinate box="[329,519,1958,1984]" degrees="108" direction="east" minutes="32" orientation="longitude" pageId="75" precision="1" seconds="00.38" value="108.53344">108°3200.38E</geoCoordinate>
, in hill evergreen forests,
<date box="[824,1033,1958,1984]" pageId="75" value="2016-02-24">
<collectingDate box="[824,1033,1958,1984]" pageId="75" value="2016-02-24">24 February 2016</collectingDate>
</date>
,
<emphasis italics="true" pageId="75">
<collectorName box="[1047,1161,1959,1985]" pageId="75">Tagane S.</collectorName>
,
<collectorName box="[1176,1336,1959,1984]" pageId="75">Nagamasu H.</collectorName>
,
<collectorName box="[1350,1446,1958,1985]" pageId="75">Naiki A.</collectorName>
,
<collectorName box="[136,251,1995,2021]" pageId="75">Dang V.S.</collectorName>
,
<collectorName box="[263,349,1995,2020]" pageId="75">Nguyen</collectorName>
</emphasis>
</materialsCitation>
<materialsCitation collectingDate="1842-05" collectingDateMax="2014-02-19" collectingDateMin="1842-05" collectionCode="FU, VNM" collectorName="Wai J. V &amp; Tagane S. &amp; Dang V. S. &amp; Nagamasu H. &amp; Naiki A. &amp; Tran H. &amp; Yang C. J. &amp; Ngoc N. V. &amp; Cuong N. Q. &amp; Hieu H. N. P. &amp; Loi X. N." country="Japan" elevation="1490" lastPageId="76" latitude="12.174142" location="Iriomote Station" longLatPrecision="1" longitude="108.69832" municipality="Iriomote Station" pageId="75" specimenCount="1" stateProvince="Toyama">
<emphasis box="[357,570,1995,2021]" italics="true" pageId="75">
<taxonomicNameLabel box="[357,406,1995,2020]" pageId="75" rank="variety">V.N.</taxonomicNameLabel>
,
<collectorName box="[418,514,1995,2021]" pageId="75">Wai J. V</collectorName>
4318
</emphasis>
(
<collectionCode box="[586,626,1995,2021]" country="China" name="Fudan University, Department of Biology" pageId="75">FU</collectionCode>
,
<collectionCode box="[638,706,1995,2021]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="75" type="Herbarium">VNM</collectionCode>
, the herbarium of
<collectingMunicipality box="[916,1100,1994,2020]" pageId="75">Iriomote Station</collectingMunicipality>
(
<collectingCountry box="[1116,1185,1995,2021]" name="Japan" pageId="75">Japan</collectingCountry>
)); in the same location,
<quantity box="[136,221,2031,2057]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="75" unit="m" value="1490.0">
<elevation box="[136,221,2031,2057]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="75" unit="m" value="1490.0">1490 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[293,471,2031,2057]" degrees="12" direction="north" minutes="10" orientation="latitude" pageId="75" precision="1" seconds="26.91" value="12.174142">12°1026.91N</geoCoordinate>
,
<geoCoordinate box="[483,673,2030,2057]" degrees="108" direction="east" minutes="41" orientation="longitude" pageId="75" precision="1" seconds="53.94" value="108.69832">108°4153.94E</geoCoordinate>
, in evergreen forests,
<date box="[918,1120,2031,2057]" pageId="75" value="2014-02-19">
<collectingDate box="[918,1120,2031,2057]" pageId="75" value="2014-02-19">19 February 2014</collectingDate>
</date>
,
<emphasis italics="true" lastPageId="76" pageId="75">
<collectingRegion box="[1131,1219,2031,2056]" country="Japan" name="Toyama" pageId="75">Toyama</collectingRegion>
H.,
<collectorName box="[1266,1377,2031,2057]" pageId="75">Tagane S.</collectorName>
,
<collectorName lastPageId="76" pageId="75">Dang V.S.</collectorName>
,
<collectorName box="[191,346,159,184]" pageId="76">Nagamasu H.</collectorName>
,
<collectorName box="[357,449,158,185]" pageId="76">Naiki A.</collectorName>
,
<collectorName box="[460,549,159,184]" pageId="76">Tran H.</collectorName>
,
<collectorName box="[559,665,159,185]" pageId="76">Yang C.J.</collectorName>
with
<collectorName box="[725,840,159,184]" pageId="76">
Ngoc
<taxonomicNameLabel box="[791,840,159,184]" pageId="76" rank="variety">N.V.</taxonomicNameLabel>
</collectorName>
,
<collectorName box="[850,986,159,185]" pageId="76">Cuong N.Q.</collectorName>
,
<collectorName box="[996,1133,159,184]" pageId="76">Hieu H.N.P.</collectorName>
,
<collectorName box="[1143,1237,159,184]" pageId="76">Loi X.N.</collectorName>
<date box="[1242,1315,158,184]" pageId="76" value="1842-05">
<collectingDate box="[1242,1315,158,184]" pageId="76" value="1842-05">V1842</collectingDate>
</date>
</emphasis>
(
<collectionCode box="[1329,1369,158,184]" country="China" name="Fudan University, Department of Biology" pageId="76">FU</collectionCode>
,
<collectionCode box="[1379,1447,159,185]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="76" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFD5FF925AB6F8334587CEC5:CE5EA5AFFFD3FF925B74FF674226CC94" box="[330,512,194,220]" country="Japan" latitude="12.174142" longLatPrecision="1" longitude="108.69832" municipality="Iriomote Station" name="Iriomote Station" pageId="76" stateProvince="Toyama">Iriomote Station</location>
(
<collectingCountry box="[527,596,195,221]" name="Japan" pageId="76">Japan</collectingCountry>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="76" type="distribution">
<paragraph blockId="76.[136,1452,158,653]" box="[189,1451,230,257]" pageId="76">
<emphasis bold="true" box="[189,345,230,256]" pageId="76">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[508,603,230,256]" name="Vietnam" pageId="76">Vietnam</collectingCountry>
(known only from the Bidoup Nui Ba National Park,
<collectingRegion box="[1207,1439,230,257]" country="Vietnam" name="Lam Dong" pageId="76">Lam Dong Province</collectingRegion>
).
</paragraph>
<paragraph blockId="76.[136,1452,158,653]" box="[189,1039,266,293]" pageId="76">
<emphasis bold="true" box="[189,423,266,293]" pageId="76">Habitat &amp; Ecology:</emphasis>
—In hill evergreen forest at around
<quantity box="[825,980,267,293]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="76" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">14001700 m</quantity>
elev.
</paragraph>
<paragraph blockId="76.[136,1452,158,653]" box="[189,869,302,329]" pageId="76">
<emphasis bold="true" box="[189,323,303,329]" pageId="76">Phenology:</emphasis>
—Flowering and fruiting from February to June.
</paragraph>
<paragraph blockId="76.[136,1452,158,653]" box="[189,658,338,367]" pageId="76">
<emphasis bold="true" box="[189,405,339,365]" pageId="76">Vernacular name:</emphasis>
—Xú hưƠng hòn giao.
</paragraph>
<paragraph blockId="76.[136,1452,158,653]" pageId="76">
<emphasis bold="true" box="[189,255,375,401]" pageId="76">Note:</emphasis>
<taxonomicName box="[283,560,374,400]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="76" phylum="Tracheophyta" rank="species" species="hongiaoensis">
<emphasis box="[283,560,374,400]" italics="true" pageId="76">Lasianthus hongiaoensis</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName authority="(Drake) Pit. (1924: 382)" authorityName="Pit." authorityPageNumber="382" authorityYear="1924" baseAuthorityName="Drake" box="[894,1313,374,401]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="76" phylum="Tracheophyta" rank="species" species="hispidulus">
<emphasis box="[894,1038,374,400]" italics="true" pageId="76">L. hispidulus</emphasis>
(Drake) Pit. (1924: 382)
</taxonomicName>
in the shape of leaves, flower color at anthesis, and blue fruits, but differs from the latter in having densely pubescent at branches and branchlets (vs. densely brown-villous), longer stipules (
<quantity box="[799,912,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="76" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
long vs.
<quantity box="[1011,1075,447,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="76" unit="mm" value="1.0">1 mm</quantity>
long) and larger calyx (
<quantity box="[1338,1452,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="76" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
long vs.
<quantity box="[232,346,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="76" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
long for tubes and
<quantity box="[562,676,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="76" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
long vs.
<quantity box="[779,893,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="76" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
long for lobes).
</paragraph>
<paragraph blockId="76.[136,1452,158,653]" pageId="76">
It is also similar to
<taxonomicName authority="V. S. Dang &amp; Naiki" authorityName="V. S. Dang &amp; Naiki" box="[408,797,518,545]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="76" phylum="Tracheophyta" rank="species" species="nuichuaensis">
<emphasis box="[408,584,518,544]" italics="true" pageId="76">L. nuichuaensis</emphasis>
V.S.Dang &amp; Naiki
</taxonomicName>
which was described from Nui Chua National Park,
<collectingRegion country="Vietnam" name="Ninh Thuan" pageId="76">Ninh Thuan Province</collectingRegion>
, Southern
<collectingCountry box="[444,542,554,580]" name="Vietnam" pageId="76">Vietnam</collectingCountry>
, but differs from the latter by having plants covered by densely pubescent (vs. glabrous), larger leaves (79 ×
<quantity box="[483,608,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="76" unit="cm" value="3.0" valueMax="3.5" valueMin="2.5">2.53.5 cm</quantity>
vs. 57 ×
<quantity box="[724,831,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="76" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.53 cm</quantity>
), fewer secondary veins (45 on each side of midrib vs. 56 on each side of midrib), larger calyx lobes (
<quantity box="[672,788,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="76" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
long vs.
<quantity box="[890,1004,626,653]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="76" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
long) and fruits with 5 (vs. 4) pyrenes.
</paragraph>
</subSubSection>
</treatment>
</document>