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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFAEFFEF5AB6FB8745A4CAA0" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus dalatensis Wernham 1921" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571636" ID-Zenodo-Dep="7571636" LSID="urn:lsid:plazi:treatment:43284262FFAEFFEF5AB6FB8745A4CAA0" httpUri="http://treatment.plazi.org/id/43284262FFAEFFEF5AB6FB8745A4CAA0" pageId="49">
<subSubSection box="[136,1038,1058,1085]" pageId="49" type="nomenclature">
<paragraph blockId="49.[136,1452,1058,1156]" box="[136,1038,1058,1085]" pageId="49">
<emphasis bold="true" box="[136,171,1058,1084]" pageId="49">17.</emphasis>
<taxonomicName ID-CoL="6NYVB" authority="Wernham (1921: 138)" authorityName="Wernham" authorityPageNumber="138" authorityYear="1921" box="[178,681,1058,1085]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="49" phylum="Tracheophyta" rank="species" species="dalatensis">
<emphasis bold="true" box="[178,429,1058,1085]" italics="true" pageId="49">Lasianthus dalatensis</emphasis>
Wernham (1921: 138)
</taxonomicName>
<figureCitation box="[712,790,1058,1084]" captionStart="FIGURE 36" captionStartId="51.[136,229,1768,1790]" captionTargetBox="[169,1414,194,1738]" captionTargetId="figure-18@51.[165,1418,193,1744]" captionTargetPageId="51" captionText="FIGURE 36. Lasianthus dalatensis Wernham (from Dang 189, VNM): A. Branch with flower buds, B. Close-up of young branch and leaves, C. Stipules, D. Abaxial leaf surfaces, EF. Inflorescence showing side view flower buds, GH. Young fruits." figureDoi="http://doi.org/10.5281/zenodo.7571660" httpUri="https://zenodo.org/record/7571660/files/figure.png" pageId="49">Fig. 36</figureCitation>
(plate),
<figureCitation box="[884,964,1058,1084]" captionStart="FIGURE 37" captionStartId="52.[136,229,1789,1811]" captionTargetBox="[176,1405,205,1753]" captionTargetId="figure-25@52.[165,1418,193,1765]" captionTargetPageId="52" captionText="FIGURE 37. Distribution maps of Lasianthus dalatensis, L. dienkhanhensis, L. eberhartii and L. elevatineurus." figureDoi="http://doi.org/10.5281/zenodo.7571662" httpUri="https://zenodo.org/record/7571662/files/figure.png" pageId="49">Fig. 37</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="49" type="materials_examined">
<paragraph blockId="49.[136,1452,1058,1156]" pageId="49">
<emphasis bold="true" box="[136,194,1098,1119]" pageId="49">Type:</emphasis>
<materialsCitation collectingDate="1918-04" collectionCode="BM" country="Vietnam" elevation="1524" location="Langbian Plateau" pageId="49" specimenCode="BM000945601" specimenCount="1" stateProvince="Lam Dong" typeStatus="holotype">
<collectingCountry box="[219,332,1098,1119]" name="Vietnam" pageId="49">VIETNAM</collectingCountry>
.
<collectingRegion box="[352,571,1098,1120]" country="Vietnam" name="Lam Dong" pageId="49">Lam Dong Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFAEFFEF5AB6FB8745A4CAA0:CE5EA5AFFFAEFFEF5871FBEF4322C828" box="[591,772,1098,1120]" country="Vietnam" name="Langbian Plateau" pageId="49" stateProvince="Lam Dong">Langbian Plateau</location>
,
<quantity box="[793,877,1098,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="49" unit="m" value="1524.0">
<elevation box="[793,877,1098,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="49" unit="m" value="1524.0">1524 m</elevation>
</quantity>
elev.,
<date box="[959,1077,1098,1120]" pageId="49" value="1918-04">
<collectingDate box="[959,1077,1098,1120]" pageId="49" value="1918-04">April 1918</collectingDate>
</date>
,
<emphasis box="[1098,1280,1098,1119]" italics="true" pageId="49">Boden Kloss s.n.</emphasis>
(
<typeStatus box="[1303,1389,1098,1120]" pageId="49">holotype</typeStatus>
BM! [
<specimenCode box="[195,349,1134,1156]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="49">BM000945601</specimenCode>
])
</materialsCitation>
.
</paragraph>
<paragraph blockId="49.[136,1452,1202,1768]" pageId="49">
<emphasis bold="true" box="[136,392,1202,1228]" pageId="49">
<collectingRegion box="[136,262,1202,1228]" pageId="49">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="1939-05" collectingDateMax="2018-07-21" collectingDateMin="1939-05" collectionCode="FU, VNM" collectorName="Toyama H. &amp; Tagane S. &amp; Dang V. S. &amp; Nagamasu H. &amp; Naiki A. &amp; Tran H. &amp; Yang C. J. &amp; Hoang N. T. B. N. &amp; Dang" country="Vietnam" county="Bidoup Nui Ba National Park" elevation="1490" location="Iriomote Station" municipality="Dalat City" pageId="49" specimenCount="1" stateProvince="Lam Dong">
<collectingCountry box="[421,553,1202,1228]" name="Vietnam" pageId="49">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[562,663,1202,1228]" pageId="49">Central:</emphasis>
<collectingRegion box="[669,896,1202,1229]" country="Vietnam" name="Lam Dong" pageId="49">Lam Dong Province</collectingRegion>
,
<collectingCounty box="[906,1235,1202,1229]" pageId="49">Bidoup Nui Ba National Park</collectingCounty>
,
<quantity box="[1245,1314,1202,1229]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="49" unit="km" value="50.0">50 km</quantity>
northeast of
<collectingMunicipality box="[136,252,1238,1264]" pageId="49">Dalat City</collectingMunicipality>
,
<quantity box="[263,347,1238,1265]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="49" unit="m" value="1490.0">
<elevation box="[263,347,1238,1265]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="49" unit="m" value="1490.0">1490 m</elevation>
</quantity>
elev.,
<date box="[418,636,1238,1264]" pageId="49" value="2014-11-19">
<collectingDate box="[418,636,1238,1264]" pageId="49" value="2014-11-19">19 November 2014</collectingDate>
</date>
,
<emphasis italics="true" pageId="49">
<collectorName box="[646,770,1239,1264]" pageId="49">Toyama H.</collectorName>
,
<collectorName box="[781,891,1239,1265]" pageId="49">Tagane S.</collectorName>
,
<collectorName box="[903,1016,1239,1265]" pageId="49">Dang V.S.</collectorName>
,
<collectorName box="[1026,1182,1239,1264]" pageId="49">Nagamasu H.</collectorName>
,
<collectorName box="[1193,1285,1238,1265]" pageId="49">Naiki A.</collectorName>
,
<collectorName box="[1297,1386,1239,1264]" pageId="49">Tran H.</collectorName>
,
<collectorName pageId="49">Yang C.J.</collectorName>
<date box="[190,264,1274,1300]" pageId="49" value="1939-05">
<collectingDate box="[190,264,1274,1300]" pageId="49" value="1939-05">V1939</collectingDate>
</date>
</emphasis>
(
<collectionCode box="[280,320,1274,1300]" country="China" name="Fudan University, Department of Biology" pageId="49">FU</collectionCode>
,
<collectionCode box="[333,401,1275,1301]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="49" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFAEFFEF5AB6FB8745A4CAA0:CE5EA5AFFFAEFFEF5858FB5F4339C95C" box="[614,799,1274,1300]" country="Vietnam" county="Bidoup Nui Ba National Park" municipality="Dalat City" name="Iriomote Station" pageId="49" stateProvince="Lam Dong">Iriomote Station</location>
(Japan));
<collectingRegion box="[917,1158,1274,1301]" country="Vietnam" name="Khanh Hoa" pageId="49">Khanh Hoa Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFAEFFEF5AB6FB8745A4CAA0:CE5EA5AFFFAEFFEF5EAAFB5E4581C95D" box="[1172,1447,1275,1301]" country="Vietnam" county="Bidoup Nui Ba National Park" municipality="Dalat City" name="Hon Ba Nature Reserve" pageId="49" stateProvince="Lam Dong">Hon Ba Nature Reserve</location>
,
<quantity box="[136,221,1310,1337]" metricMagnitude="3" metricUnit="m" metricValue="1.512" pageId="49" unit="m" value="1512.0">
<elevation box="[136,221,1310,1337]" metricMagnitude="3" metricUnit="m" metricValue="1.512" pageId="49" unit="m" value="1512.0">1512 m</elevation>
</quantity>
elev.,
<date box="[294,441,1310,1336]" pageId="49" value="2018-07-21">
<collectingDate box="[294,441,1310,1336]" pageId="49" value="2018-07-21">21 July 2018</collectingDate>
</date>
,
<emphasis box="[452,903,1310,1337]" italics="true" pageId="49">
<collectorName box="[452,564,1311,1337]" pageId="49">Dang V.S.</collectorName>
&amp;
<collectorName box="[599,781,1311,1336]" pageId="49">Hoang N.T.B.N.</collectorName>
,
<collectorName box="[792,854,1311,1336]" pageId="49">Dang</collectorName>
189
</emphasis>
(
<collectionCode box="[918,989,1310,1336]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="49" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="49" type="distribution">
<paragraph blockId="49.[136,1452,1202,1768]" pageId="49">
<emphasis bold="true" box="[189,345,1346,1372]" pageId="49">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[510,605,1346,1372]" name="Vietnam" pageId="49">Vietnam</collectingCountry>
(currently known as Bidoup Nui Ba National Park in
<collectingRegion box="[1222,1452,1346,1373]" country="Vietnam" name="Lam Dong" pageId="49">Lam Dong Province</collectingRegion>
and Hon Ba Nature Reserve in
<collectingRegion box="[486,726,1382,1409]" country="Vietnam" name="Khanh Hoa" pageId="49">Khanh Hoa Province</collectingRegion>
).
</paragraph>
<paragraph blockId="49.[136,1452,1202,1768]" pageId="49">
<emphasis bold="true" box="[189,421,1418,1444]" pageId="49">Habitat &amp; Ecology:</emphasis>
—Growing in primary or secondary montane forests, rarely at the edge of a forest,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.6" metricValueMin="0.9" pageId="49" unit="m" value="1250.0" valueMax="1600.0" valueMin="900.0">9001600 m</quantity>
elev.
</paragraph>
<paragraph blockId="49.[136,1452,1202,1768]" box="[189,880,1490,1517]" pageId="49">
<emphasis bold="true" box="[189,323,1490,1516]" pageId="49">Phenology:</emphasis>
—Flowering and fruiting from July to November.
</paragraph>
<paragraph blockId="49.[136,1452,1202,1768]" box="[189,622,1526,1555]" pageId="49">
<emphasis bold="true" box="[189,405,1526,1552]" pageId="49">Vernacular name:</emphasis>
—Xú hưƠng đà lạt.
</paragraph>
<paragraph blockId="49.[136,1452,1202,1768]" pageId="49">
<emphasis bold="true" box="[189,265,1562,1588]" pageId="49">Notes:</emphasis>
<taxonomicName box="[294,539,1562,1588]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="49" phylum="Tracheophyta" rank="species" species="dalatensis">
<emphasis box="[294,539,1562,1588]" italics="true" pageId="49">Lasianthus dalatensis</emphasis>
</taxonomicName>
was described by Wernham in 1921 from a Vietnamese specimen (
<emphasis italics="true" pageId="49">Boden Kloss s.n.</emphasis>
). This species is characterized by having an elliptic-lanceolate leaf blade, glabrous on both surfaces, secondary veins 79 on each side of midrib, and flowers crimson to pink, and ultimately white.
</paragraph>
<paragraph blockId="49.[136,1452,1202,1768]" pageId="49">
<taxonomicName box="[189,433,1670,1696]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="49" phylum="Tracheophyta" rank="species" species="dalatensis">
<emphasis box="[189,433,1670,1696]" italics="true" pageId="49">Lasianthus dalatensis</emphasis>
</taxonomicName>
morphologically resembles
<taxonomicName authority="V. S. Dang, Tagane &amp; H. Tran (2015: 352)" authorityName="V. S. Dang, Tagane &amp; H. Tran" authorityPageNumber="352" authorityYear="2015" box="[749,1327,1670,1697]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="49" phylum="Tracheophyta" rank="species" species="yaharae">
<emphasis box="[749,870,1670,1696]" italics="true" pageId="49">L. yaharae</emphasis>
V.S.Dang, Tagane &amp; H.Tran (2015: 352)
</taxonomicName>
which was discovered in 2015 from Hon Ba Nature Reserve of
<collectingRegion box="[732,972,1706,1733]" country="Vietnam" name="Khanh Hoa" pageId="49">Khanh Hoa Province</collectingRegion>
, but differs from the latter in having more number of secondary veins, ultimately white flowers in anthesis and smaller calyx, corolla tube and corolla lobes.
</paragraph>
</subSubSection>
</treatment>
</document>