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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFA6FFE55AB6FF3B41C7CE0C" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus fansipanensis V. S. Dang &amp; Naiki 2019" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571654" ID-Zenodo-Dep="7571654" LSID="urn:lsid:plazi:treatment:43284262FFA6FFE55AB6FF3B41C7CE0C" httpUri="http://treatment.plazi.org/id/43284262FFA6FFE55AB6FF3B41C7CE0C" lastPageId="59" pageId="57">
<subSubSection box="[136,1164,158,185]" pageId="57" type="nomenclature">
<paragraph blockId="57.[136,1452,158,292]" box="[136,1164,158,185]" pageId="57">
<emphasis bold="true" box="[136,171,158,184]" pageId="57">21.</emphasis>
<taxonomicName ID-CoL="8Y54V" authority="V. S. Dang &amp; Naiki (2019: 34)" authorityName="V. S. Dang &amp; Naiki" authorityPageNumber="34" authorityYear="2019" box="[178,809,158,185]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="57" phylum="Tracheophyta" rank="species" species="fansipanensis">
<emphasis bold="true" box="[178,473,158,185]" italics="true" pageId="57">Lasianthus fansipanensis</emphasis>
V.S.
<bibRefCitation author="Dang, V. S. &amp; Naiki, A." box="[522,809,158,184]" pageId="57" pagination="33 - 40" refId="ref54157" refString="Dang, V. S. &amp; Naiki, A. (2019) Two new species of Lasianthus (Rubiaceae) from Hoang Lien National Park, northern Vietnam. Phytotaxa 423 (1): 33 - 40. https: // doi. org / 10.11646 / phytotaxa. 423.1.4" type="journal article" year="2019">Dang &amp; Naiki (2019: 34)</bibRefCitation>
</taxonomicName>
<emphasis bold="true" box="[809,837,158,184]" pageId="57">—</emphasis>
<figureCitation box="[837,917,158,184]" captionStart="FIGURE 41" captionStartId="57.[136,229,1937,1959]" captionTargetBox="[170,1415,357,1908]" captionTargetId="figure-81@57.[166,1418,356,1912]" captionTargetPageId="57" captionText="FIGURE 41. Lasianthus fansipanensis V.S.Dang &amp; Naiki (from Dang 233, VNM and the herbarium of Iriomote Station (Japan)): A. Branch, B. Close-up of young branch and leaves, C. Stipules, D. Abaxial leaf surfaces, E–F. Inflorescence showing side view flower buds, G–H. Mature fruits, I. Seeds." figureDoi="http://doi.org/10.5281/zenodo.7571676" httpUri="https://zenodo.org/record/7571676/files/figure.png" pageId="57">Fig. 41</figureCitation>
(plate),
<figureCitation box="[1011,1090,158,184]" captionStart="FIGURE 42" captionStartId="58.[136,229,1789,1811]" captionTargetBox="[173,1409,203,1757]" captionTargetId="figure-25@58.[165,1419,193,1765]" captionTargetPageId="58" captionText="FIGURE 42. Distribution maps of Lasianthus fansipanensis, L. foetidissimus, L. fordii and L. formosensis." figureDoi="http://doi.org/10.5281/zenodo.7571678" httpUri="https://zenodo.org/record/7571678/files/figure.png" pageId="57">Fig. 42</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection lastPageId="58" pageId="57" type="materials_examined">
<paragraph blockId="57.[136,1452,158,292]" pageId="57">
<emphasis bold="true" box="[136,194,198,219]" pageId="57">Type:</emphasis>
—
<materialsCitation collectingDate="2018-11-11" collectionCode="VNM, HN" collectorName="Dang V. S. &amp; Naiki A. &amp; Dang" country="Vietnam" county="Sa Pa District" elevation="1906" location="Fansipan" municipality="Hoang Lien National Park" pageId="57" specimenCode="VNM00042888, VNM00042889, VNM00042890" specimenCount="1" stateProvince="Lao Cai" typeStatus="holotype">
<collectingCountry box="[219,332,198,219]" name="Vietnam" pageId="57">VIETNAM</collectingCountry>
.
<collectingRegion box="[341,511,198,220]" country="Vietnam" name="Lao Cai" pageId="57">Lao Cai Province</collectingRegion>
,
<collectingCounty box="[521,655,198,220]" pageId="57">Sa Pa District</collectingCounty>
,
<collectingMunicipality box="[664,919,198,220]" pageId="57">Hoang Lien National Park</collectingMunicipality>
, on the way to
<location LSID="urn:lsid:plazi:treatment:43284262FFA6FFE55AB6FF3B41C7CE0C:CE5EA5AFFFA6FFE75E16FF634459CC94" box="[1064,1151,198,220]" country="Vietnam" county="Sa Pa District" municipality="Hoang Lien National Park" name="Fansipan" pageId="57" stateProvince="Lao Cai">Fansipan</location>
peak from
<location LSID="urn:lsid:plazi:treatment:43284262FFA6FFE55AB6FF3B41C7CE0C:CE5EA5AFFFA6FFE75ED2FF63457FCC94" box="[1260,1369,198,220]" country="Vietnam" county="Sa Pa District" municipality="Hoang Lien National Park" name="Ton station" pageId="57" stateProvince="Lao Cai">Ton station</location>
,
<quantity box="[1379,1452,198,220]" metricMagnitude="3" metricUnit="m" metricValue="1.906" pageId="57" unit="m" value="1906.0">
<elevation box="[1379,1452,198,220]" metricMagnitude="3" metricUnit="m" metricValue="1.906" pageId="57" unit="m" value="1906.0">1906 m</elevation>
</quantity>
elev.,
<date box="[244,427,234,256]" pageId="57" value="2018-11-11">
<collectingDate box="[244,427,234,256]" pageId="57" value="2018-11-11">11 November 2018</collectingDate>
</date>
,
<emphasis box="[435,735,234,255]" italics="true" pageId="57">
<collectorName box="[435,529,234,255]" pageId="57">Dang V.S.</collectorName>
&amp;
<collectorName box="[555,634,234,255]" pageId="57">Naiki A.</collectorName>
,
<collectorName box="[642,695,234,255]" pageId="57">Dang</collectorName>
233
</emphasis>
(
<typeStatus box="[746,832,234,256]" pageId="57">holotype</typeStatus>
<collectionCode box="[835,896,234,255]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="57" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[909,1068,234,255]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="57" type="Herbarium">VNM00042888</specimenCode>
]; isotypes
<collectionCode box="[1165,1206,234,255]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="57" type="Herbarium">HN</collectionCode>
!,
<collectionCode box="[1217,1278,234,255]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="57" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[1291,1447,234,255]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="57" type="Herbarium">VNM00042889</specimenCode>
,
<specimenCode box="[189,347,270,291]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="57" type="Herbarium">VNM00042890</specimenCode>
], the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFA6FFE55AB6FF3B41C7CE0C:CE5EA5AFFFA6FFE75831FEAB4288CD6C" box="[527,686,270,292]" country="Vietnam" county="Sa Pa District" municipality="Hoang Lien National Park" name="Iriomote Station" pageId="57" stateProvince="Lao Cai">Iriomote Station</location>
, (Japan))
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571676" ID-Zenodo-Dep="7571676" httpUri="https://zenodo.org/record/7571676/files/figure.png" pageId="57" startId="57.[136,229,1937,1959]" targetBox="[170,1415,357,1908]" targetPageId="57">
<paragraph blockId="57.[136,1452,1937,2031]" pageId="57">
<emphasis bold="true" box="[136,267,1937,1959]" pageId="57">FIGURE 41.</emphasis>
<taxonomicName authority="V. S. Dang &amp; Naiki" authorityName="V. S. Dang &amp; Naiki" box="[276,707,1937,1959]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="57" phylum="Tracheophyta" rank="species" species="fansipanensis">
<emphasis box="[276,520,1937,1958]" italics="true" pageId="57">Lasianthus fansipanensis</emphasis>
V.S.Dang &amp; Naiki
</taxonomicName>
(from
<emphasis box="[778,875,1937,1959]" italics="true" pageId="57">Dang 233</emphasis>
, VNM and the herbarium of Iriomote Station (Japan)):
<emphasis bold="true" box="[1428,1452,1937,1958]" pageId="57">A.</emphasis>
Branch,
<emphasis bold="true" box="[217,239,1973,1994]" pageId="57">B.</emphasis>
Close-up of young branch and leaves,
<emphasis bold="true" box="[607,631,1973,1995]" pageId="57">C.</emphasis>
Stipules,
<emphasis bold="true" box="[725,748,1973,1994]" pageId="57">D.</emphasis>
Abaxial leaf surfaces,
<emphasis bold="true" box="[964,1010,1973,1994]" pageId="57">E–F.</emphasis>
Inflorescence showing side view flower buds,
<emphasis bold="true" box="[136,191,2009,2031]" pageId="57">G–H.</emphasis>
Mature fruits,
<emphasis bold="true" box="[335,351,2009,2030]" pageId="57">I.</emphasis>
Seeds.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571678" ID-Zenodo-Dep="7571678" box="[136,1163,1789,1811]" httpUri="https://zenodo.org/record/7571678/files/figure.png" pageId="58" startId="58.[136,229,1789,1811]" targetBox="[173,1409,203,1757]" targetPageId="58">
<paragraph blockId="58.[136,1163,1789,1811]" box="[136,1163,1789,1811]" pageId="58">
<emphasis bold="true" box="[136,265,1789,1811]" pageId="58">FIGURE 42.</emphasis>
Distribution maps of
<taxonomicName box="[476,718,1789,1810]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="58" phylum="Tracheophyta" rank="species" species="fansipanensis">
<emphasis box="[476,718,1789,1810]" italics="true" pageId="58">Lasianthus fansipanensis</emphasis>
</taxonomicName>
,
<taxonomicName box="[730,880,1789,1811]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="58" phylum="Tracheophyta" rank="species" species="foetidissimus">
<emphasis box="[730,880,1789,1811]" italics="true" pageId="58">L. foetidissimus</emphasis>
</taxonomicName>
,
<taxonomicName box="[893,970,1789,1811]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="58" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis box="[893,970,1789,1811]" italics="true" pageId="58">L. fordii</emphasis>
</taxonomicName>
and
<taxonomicName box="[1017,1158,1789,1811]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="58" phylum="Tracheophyta" rank="species" species="formosensis">
<emphasis box="[1017,1158,1789,1811]" italics="true" pageId="58">L. formosensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="58.[136,1452,1857,2027]" pageId="58">
<emphasis bold="true" box="[136,394,1857,1883]" pageId="58">
<collectingRegion box="[136,262,1857,1883]" pageId="58">Specimens</collectingRegion>
examined:
</emphasis>
—
<materialsCitation collectingDate="2016-04-29" collectingDateMax="2018-11-11" collectingDateMin="2016-04-29" collectionCode="VNM" collectorName="Dang V. S. &amp; Naiki A. &amp; Yahara T. &amp; Tagane S. &amp; Nagamasu H. &amp; Nguyen" country="Vietnam" county="Sa Pa District" elevation="1906" location="Fansipan" municipality="Hoang Lien National Park" pageId="58" specimenCount="1" stateProvince="Lao Cai">
<collectingCountry box="[423,555,1857,1883]" name="Vietnam" pageId="58">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[565,686,1857,1883]" pageId="58">Northern:</emphasis>
<collectingRegion box="[693,892,1857,1884]" country="Vietnam" name="Lao Cai" pageId="58">Lao Cai Province</collectingRegion>
,
<collectingCounty box="[904,1061,1857,1884]" pageId="58">Sa Pa District</collectingCounty>
,
<collectingMunicipality box="[1072,1370,1857,1884]" pageId="58">Hoang Lien National Park</collectingMunicipality>
, on the way to
<location LSID="urn:lsid:plazi:treatment:43284262FFA6FFE55AB6FF3B41C7CE0C:CE5EA5AFFFA5FFE45AE4F8C04119CB37" box="[218,319,1893,1919]" country="Vietnam" county="Sa Pa District" municipality="Hoang Lien National Park" name="Fansipan" pageId="58" stateProvince="Lao Cai">Fansipan</location>
peak from
<location LSID="urn:lsid:plazi:treatment:43284262FFA6FFE55AB6FF3B41C7CE0C:CE5EA5AFFFA5FFE45B80F8C3421BCB37" box="[446,573,1893,1920]" country="Vietnam" county="Sa Pa District" municipality="Hoang Lien National Park" name="Ton station" pageId="58" stateProvince="Lao Cai">Ton station</location>
,
<quantity box="[584,669,1893,1920]" metricMagnitude="3" metricUnit="m" metricValue="1.906" pageId="58" unit="m" value="1906.0">
<elevation box="[584,669,1893,1920]" metricMagnitude="3" metricUnit="m" metricValue="1.906" pageId="58" unit="m" value="1906.0">1906 m</elevation>
</quantity>
elev.,
<date box="[740,958,1893,1919]" pageId="58" value="2018-11-11">
<collectingDate box="[740,958,1893,1919]" pageId="58" value="2018-11-11">11 November 2018</collectingDate>
</date>
,
<emphasis box="[969,1322,1893,1920]" italics="true" pageId="58">
<collectorName box="[969,1082,1894,1920]" pageId="58">Dang V.S.</collectorName>
,
<collectorName box="[1093,1186,1893,1920]" pageId="58">Naiki A.</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA6FFE55AB6FF3B41C7CE0C:CE5EA5AFFFA5FFE45E93F8C344CDCB37" box="[1197,1259,1894,1919]" country="Vietnam" county="Sa Pa District" municipality="Hoang Lien National Park" name="Dang" pageId="58" stateProvince="Lao Cai">Dang</location>
233a
</emphasis>
(
<collectionCode box="[1337,1407,1893,1919]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="58" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFA6FFE55AB6FF3B41C7CE0C:CE5EA5AFFFA5FFE45B21F82C41F3CBEB" box="[287,469,1929,1955]" country="Vietnam" county="Sa Pa District" municipality="Hoang Lien National Park" name="Iriomote Station" pageId="58" stateProvince="Lao Cai">Iriomote Station</location>
(Japan)); in the same location,
<quantity box="[817,901,1929,1956]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="58" unit="m" value="1850.0">
<elevation box="[817,901,1929,1956]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="58" unit="m" value="1850.0">1850 m</elevation>
</quantity>
elev.,
<date box="[972,1130,1929,1955]" pageId="58" value="2016-04-29">
<collectingDate box="[972,1130,1929,1955]" pageId="58" value="2016-04-29">29 April 2016</collectingDate>
</date>
,
<emphasis italics="true" pageId="58">
<collectorName box="[1140,1232,1929,1956]" pageId="58">Naiki A.</collectorName>
,
<collectorName box="[1244,1353,1929,1955]" pageId="58">Yahara T.</collectorName>
, Toyama H.,
<collectorName box="[180,293,1966,1992]" pageId="58">Tagane S.</collectorName>
,
<collectorName box="[308,467,1966,1991]" pageId="58">Nagamasu H.</collectorName>
,
<collectorName box="[481,567,1966,1991]" pageId="58">Nguyen</collectorName>
</emphasis>
</materialsCitation>
<materialsCitation collectionCode="FU, VNM" collectorName="Hoang T. B. &amp; Hoang T. S. V" country="Japan" location="Iriomote Station" pageId="58" specimenCount="1" stateProvince="Northern">
<emphasis box="[576,1108,1965,1992]" italics="true" pageId="58">
<taxonomicNameLabel box="[576,625,1966,1991]" pageId="58" rank="variety">V.N.</taxonomicNameLabel>
,
<collectorName box="[638,771,1966,1991]" pageId="58">Hoang T.B.</collectorName>
,
<collectorName box="[785,938,1965,1992]" pageId="58">Hoang T.S. V</collectorName>
4782 &amp; V4804
</emphasis>
(
<collectionCode box="[1125,1165,1965,1991]" country="China" name="Fudan University, Department of Biology" pageId="58">FU</collectionCode>
,
<collectionCode box="[1178,1246,1966,1992]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="58" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFA6FFE55AB6FF3B41C7CE0C:CE5EA5AFFFA5FFE45AB6F8744118CBA3" box="[136,318,2001,2027]" country="Japan" name="Iriomote Station" pageId="58" stateProvince="Northern">Iriomote Station</location>
(
<collectingCountry box="[334,403,2001,2027]" name="Japan" pageId="58">Japan</collectingCountry>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="59" type="distribution">
<paragraph blockId="59.[136,1452,158,581]" pageId="59">
<emphasis bold="true" box="[189,345,158,184]" pageId="59">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[523,618,158,184]" name="Vietnam" pageId="59">Vietnam</collectingCountry>
(currently known only from Hoang Lien National Park of
<collectingRegion country="Vietnam" name="Lao Cai" pageId="59">Lao Cai Province</collectingRegion>
).
</paragraph>
<paragraph blockId="59.[136,1452,158,581]" box="[189,1405,230,257]" pageId="59">
<emphasis bold="true" box="[189,423,230,256]" pageId="59">Habitat &amp; Ecology:</emphasis>
—Growing under the shade of the primary and secondary forests, at
<quantity box="[1191,1346,230,257]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="59" unit="m" value="2000.0" valueMax="2100.0" valueMin="1900.0">1900–2100 m</quantity>
elev.
</paragraph>
<paragraph blockId="59.[136,1452,158,581]" box="[189,877,266,293]" pageId="59">
<emphasis bold="true" box="[189,323,266,292]" pageId="59">Phenology:</emphasis>
—Flowering and fruiting from July to December.
</paragraph>
<paragraph blockId="59.[136,1452,158,581]" box="[189,656,302,331]" pageId="59">
<emphasis bold="true" box="[189,405,302,328]" pageId="59">Vernacular name:</emphasis>
—Xú hưƠng fansipan.
</paragraph>
<paragraph blockId="59.[136,1452,158,581]" pageId="59">
<emphasis bold="true" box="[189,265,338,364]" pageId="59">Notes:</emphasis>
—
<taxonomicName box="[294,579,338,364]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="59" phylum="Tracheophyta" rank="species" species="fansipanensis">
<emphasis box="[294,579,338,364]" italics="true" pageId="59">Lasianthus fansipanensis</emphasis>
</taxonomicName>
was described from a Vietnamese specimen (
<emphasis box="[1114,1228,338,364]" italics="true" pageId="59">Dang 233</emphasis>
) by
<bibRefCitation author="Dang, V. S. &amp; Naiki, A." pageId="59" pagination="33 - 40" refId="ref54157" refString="Dang, V. S. &amp; Naiki, A. (2019) Two new species of Lasianthus (Rubiaceae) from Hoang Lien National Park, northern Vietnam. Phytotaxa 423 (1): 33 - 40. https: // doi. org / 10.11646 / phytotaxa. 423.1.4" type="journal article" year="2019">Dang &amp; Naiki (2019)</bibRefCitation>
. It is easily recognizable in the field in having lanceolate to oblong-lanceolate leaves, tip pointed
<quantity box="[1329,1452,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="59" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
long, oblique or obtuse leaf base, peduncles
<quantity box="[644,739,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="59" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
long, flowers pinkish white to white, and fruits ellipsoid, blue when ripe.
</paragraph>
<paragraph blockId="59.[136,1452,158,581]" pageId="59">
This species is closely related to
<taxonomicName authority="A. Chev. ex Pit. (1924: 395)" authorityName="Pit." authorityPageNumber="395" authorityYear="1924" box="[565,1062,482,509]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="59" phylum="Tracheophyta" rank="species" species="foetidissimus">
<emphasis box="[565,742,482,508]" italics="true" pageId="59">L. foetidissimus</emphasis>
A.Chev. ex Pit. (1924: 395)
</taxonomicName>
which is endemic to
<collectingCountry box="[1306,1404,482,508]" name="Vietnam" pageId="59">Vietnam</collectingCountry>
, but differs in its oblique to obtuse leaf base (vs. acute or slightly rounded), blue fruits (vs. yellowish orange), and fruits with 4 pyrenes (vs. 5 pyrenes).
</paragraph>
</subSubSection>
</treatment>
</document>
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