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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFA4FFE55AB6FDD74566C9C8" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus foetidissimus Pit. 1924" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571658" ID-Zenodo-Dep="7571658" LSID="urn:lsid:plazi:treatment:43284262FFA4FFE55AB6FDD74566C9C8" httpUri="http://treatment.plazi.org/id/43284262FFA4FFE55AB6FDD74566C9C8" pageId="59">
<subSubSection box="[136,1135,626,653]" pageId="59" type="nomenclature">
<paragraph blockId="59.[136,1452,626,724]" box="[136,1135,626,653]" pageId="59">
<emphasis bold="true" box="[136,171,626,652]" pageId="59">22.</emphasis>
<taxonomicName ID-CoL="3SCL6" authority="A. Chev. ex Pit. (1924: 395)" authorityName="Pit." authorityPageNumber="395" authorityYear="1924" box="[178,781,626,653]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="59" phylum="Tracheophyta" rank="species" species="foetidissimus">
<emphasis bold="true" box="[178,465,626,653]" italics="true" pageId="59">Lasianthus foetidissimus</emphasis>
A.Chev. ex Pit. (1924: 395)
</taxonomicName>
<emphasis bold="true" box="[781,809,626,652]" pageId="59"></emphasis>
<figureCitation box="[809,888,626,652]" captionStart="FIGURE 43" captionStartId="60.[136,229,1763,1785]" captionTargetBox="[172,1416,196,1736]" captionTargetId="figure-24@60.[164,1420,193,1739]" captionTargetPageId="60" captionText="FIGURE 43. Lasianthus foetidissimus A.Chev. ex Pit. (from Dang 343, VNM and the herbarium of Iriomote Station (Japan)): A. Branch with flowers and bubs, B. Stipules, C. Abaxial leaf surfaces, DE. Inflorescence showing side view buds and flower, F. Flower, G. Flower buds and fruit, H. Young fruits, I. Mature fruits." figureDoi="http://doi.org/10.5281/zenodo.7571682" httpUri="https://zenodo.org/record/7571682/files/figure.png" pageId="59">Fig. 43</figureCitation>
(plate),
<figureCitation box="[982,1061,626,652]" captionStart="FIGURE 42" captionStartId="58.[136,229,1789,1811]" captionTargetBox="[173,1409,203,1757]" captionTargetId="figure-25@58.[165,1419,193,1765]" captionTargetPageId="58" captionText="FIGURE 42. Distribution maps of Lasianthus fansipanensis, L. foetidissimus, L. fordii and L. formosensis." figureDoi="http://doi.org/10.5281/zenodo.7571678" httpUri="https://zenodo.org/record/7571678/files/figure.png" pageId="59">Fig. 42</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="59" type="materials_examined">
<paragraph blockId="59.[136,1452,626,724]" pageId="59">
<emphasis bold="true" box="[136,194,666,687]" pageId="59">Type:</emphasis>
<materialsCitation collectingDate="1918-04-03" collectorName="A. Chevalier" country="Vietnam" elevation="1350" location="Hon Ba Nature Reserve" pageId="59" specimenCount="1" stateProvince="Khanh Hoa" typeStatus="holotype">
<collectingCountry box="[219,332,666,687]" name="Vietnam" pageId="59">VIETNAM</collectingCountry>
.
<collectingRegion box="[342,547,666,688]" country="Vietnam" name="Khanh Hoa" pageId="59">Khanh Hoa Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE55AB6FDD74566C9C8:CE5EA5AFFFA4FFE55810FD3F433ECEE7" box="[558,792,666,687]" country="Vietnam" name="Hon Ba Nature Reserve" pageId="59" stateProvince="Khanh Hoa">Hon Ba Nature Reserve</location>
,
<quantity box="[802,936,666,688]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="59" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">
<elevation box="[802,936,666,688]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="59" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">12001500 m</elevation>
</quantity>
elev.,
<date box="[999,1124,666,688]" pageId="59" value="1918-04-03">
<collectingDate box="[999,1124,666,688]" pageId="59" value="1918-04-03">3 April 1918</collectingDate>
</date>
,
<emphasis box="[1135,1324,666,687]" italics="true" pageId="59">
<collectorName box="[1135,1258,666,687]" pageId="59">A. Chevalier</collectorName>
38778
</emphasis>
(
<typeStatus box="[1338,1424,666,688]" pageId="59">holotype</typeStatus>
P! [P03980391]; isotype P!)
</materialsCitation>
.
</paragraph>
<paragraph blockId="59.[136,1452,770,1409]" pageId="59">
<emphasis bold="true" box="[136,394,770,796]" pageId="59">
<collectingRegion box="[136,262,770,796]" pageId="59">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="1778-05" collectingDateMax="2019-06-15" collectingDateMin="1778-05" collectionCode="FU, VNM" collectorName="Tagane S. &amp; Kanemitsu H. &amp; Dang V. S. &amp; Tran H. &amp; Loi X. N. &amp; Thach N. D. &amp; Hieu P. N. H. &amp; Hoang N. T. B. N. &amp; L. Averyanov &amp; N. Q. Binh &amp; P. K. Loc &amp; Toyama H. &amp; Nagamasu H. &amp; Naiki A. &amp; Yang C. J." country="Vietnam" county="Hon Ba Nature Reserve" elevation="1300" location="Lac Duong District" municipality="Iriomote Station" pageId="59" specimenCount="1" stateProvince="Khanh Hoa">
<collectingCountry box="[423,555,770,796]" name="Vietnam" pageId="59">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[566,667,770,796]" pageId="59">Central:</emphasis>
<collectingRegion box="[675,913,770,797]" country="Vietnam" name="Khanh Hoa" pageId="59">Khanh Hoa Province</collectingRegion>
,
<collectingCounty box="[926,1197,771,797]" pageId="59">Hon Ba Nature Reserve</collectingCounty>
,
<quantity box="[1209,1295,770,797]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="59" unit="m" value="1300.0">
<elevation box="[1209,1295,770,797]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="59" unit="m" value="1300.0">1300 m</elevation>
</quantity>
elev.,
<date pageId="59" value="2014-07-13">
<collectingDate pageId="59" value="2014-07-13">13 July 2014</collectingDate>
</date>
,
<emphasis box="[206,1235,806,833]" italics="true" pageId="59">
<collectorName box="[206,317,807,833]" pageId="59">Tagane S.</collectorName>
,
<collectorName box="[329,485,807,832]" pageId="59">Kanemitsu H.</collectorName>
,
<collectorName box="[496,609,807,833]" pageId="59">Dang V.S.</collectorName>
,
<collectorName box="[620,708,807,832]" pageId="59">Tran H.</collectorName>
with
<collectorName box="[769,866,807,832]" pageId="59">Loi X.N.</collectorName>
,
<collectorName box="[877,1009,806,832]" pageId="59">Thach N.D.</collectorName>
,
<collectorName box="[1020,1155,807,832]" pageId="59">Hieu P.N.H.</collectorName>
<date box="[1161,1235,806,832]" pageId="59" value="1778-05">
<collectingDate box="[1161,1235,806,832]" pageId="59" value="1778-05">V1778</collectingDate>
</date>
</emphasis>
(
<collectionCode box="[1249,1289,806,832]" country="China" name="Fudan University, Department of Biology" pageId="59">FU</collectionCode>
,
<collectionCode box="[1299,1369,806,832]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="59" type="Herbarium">VNM</collectionCode>
); in the same location,
<quantity box="[303,387,842,869]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="59" unit="m" value="1000.0">
<elevation box="[303,387,842,869]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="59" unit="m" value="1000.0">1000 m</elevation>
</quantity>
elev.,
<date box="[457,603,842,868]" pageId="59" value="2018-07-22">
<collectingDate box="[457,603,842,868]" pageId="59" value="2018-07-22">22 July 2018</collectingDate>
</date>
,
<emphasis box="[613,1057,842,869]" italics="true" pageId="59">
<collectorName box="[613,723,843,869]" pageId="59">Dang V.S.</collectorName>
&amp;
<collectorName box="[756,937,843,868]" pageId="59">Hoang N.T.B.N.</collectorName>
, Dang 190
</emphasis>
(
<collectionCode box="[1071,1142,842,868]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="59" type="Herbarium">VNM</collectionCode>
); in the same location,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.502" pageId="59" unit="m" value="1502.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.502" pageId="59" unit="m" value="1502.0">1502 m</elevation>
</quantity>
elev.,
<date box="[234,389,878,905]" pageId="59" value="2019-06-15">
<collectingDate box="[234,389,878,905]" pageId="59" value="2019-06-15">15 June 2019</collectingDate>
</date>
,
<emphasis box="[402,862,878,905]" italics="true" pageId="59">
<collectorName box="[402,515,879,905]" pageId="59">Dang V.S.</collectorName>
&amp;
<collectorName box="[553,737,879,904]" pageId="59">Hoang N.T.B.N.</collectorName>
, Dang 343
</emphasis>
(
<collectionCode box="[878,948,878,904]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="59" type="Herbarium">VNM</collectionCode>
, the herbarium of
<collectingMunicipality box="[1159,1343,878,904]" pageId="59">Iriomote Station</collectingMunicipality>
(Japan));
<collectingRegion box="[136,367,914,941]" country="Vietnam" name="Lam Dong" pageId="59">Lam Dong Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE55AB6FDD74566C9C8:CE5EA5AFFFA4FFE55B45FC36427ECFE4" box="[379,600,914,941]" country="Vietnam" county="Hon Ba Nature Reserve" municipality="Iriomote Station" name="Lac Duong District" pageId="59" stateProvince="Khanh Hoa">Lac Duong District</location>
,
<quantity box="[612,697,914,941]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="59" unit="m" value="1500.0">
<elevation box="[612,697,914,941]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="59" unit="m" value="1500.0">1500 m</elevation>
</quantity>
elev.,
<date box="[771,931,914,941]" pageId="59" value="1997-04-15">
<collectingDate box="[771,931,914,941]" pageId="59" value="1997-04-15">15 April 1997</collectingDate>
</date>
,
<emphasis box="[943,1451,914,941]" italics="true" pageId="59">
<collectorName box="[943,1092,915,941]" pageId="59">L. Averyanov</collectorName>
,
<collectorName box="[1104,1228,914,941]" pageId="59">N. Q. Binh</collectorName>
,
<collectorName box="[1240,1346,915,940]" pageId="59">P. K. Loc</collectorName>
, VH3979
</emphasis>
(P); in the same location,
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE55AB6FDD74566C9C8:CE5EA5AFFFA4FFE55B90FC134327CF98" box="[430,769,950,977]" country="Vietnam" county="Hon Ba Nature Reserve" municipality="Iriomote Station" name="Bidoup Nui Ba National Park" pageId="59" stateProvince="Khanh Hoa">Bidoup Nui Ba National Park</location>
,
<quantity box="[782,854,950,977]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="59" unit="km" value="50.0">50 km</quantity>
northeast of
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE55AB6FDD74566C9C8:CE5EA5AFFFA4FFE559D3FC134445CF98" box="[1005,1123,950,976]" country="Vietnam" county="Hon Ba Nature Reserve" municipality="Iriomote Station" name="Dalat City" pageId="59" stateProvince="Khanh Hoa">Dalat City</location>
,
<quantity box="[1135,1222,950,977]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="59" unit="m" value="1490.0">
<elevation box="[1135,1222,950,977]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="59" unit="m" value="1490.0">1490 m</elevation>
</quantity>
elev.,
<date pageId="59" value="2014-11-19">
<collectingDate pageId="59" value="2014-11-19">19 November 2014</collectingDate>
</date>
,
<emphasis box="[206,1151,986,1013]" italics="true" pageId="59">
<collectorName box="[206,330,987,1012]" pageId="59">Toyama H.</collectorName>
,
<collectorName box="[342,453,987,1013]" pageId="59">Tagane S.</collectorName>
,
<collectorName box="[465,578,987,1013]" pageId="59">Dang V.S.</collectorName>
,
<collectorName box="[589,746,987,1012]" pageId="59">Nagamasu H.</collectorName>
,
<collectorName box="[757,850,986,1013]" pageId="59">Naiki A.</collectorName>
,
<collectorName box="[862,952,987,1012]" pageId="59">Tran H.</collectorName>
,
<collectorName box="[963,1071,987,1013]" pageId="59">Yang C.J.</collectorName>
<date box="[1077,1151,986,1012]" pageId="59" value="1956-05">
<collectingDate box="[1077,1151,986,1012]" pageId="59" value="1956-05">V1956</collectingDate>
</date>
</emphasis>
(
<collectionCode box="[1165,1205,986,1012]" country="China" name="Fudan University, Department of Biology" pageId="59">FU</collectionCode>
,
<collectionCode box="[1216,1284,987,1013]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="59" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE55AB6FDD74566C9C8:CE5EA5AFFFA4FFE55A98FC5B417BC850" box="[166,349,1022,1048]" country="Vietnam" county="Hon Ba Nature Reserve" municipality="Iriomote Station" name="Iriomote Station" pageId="59" stateProvince="Khanh Hoa">Iriomote Station</location>
(Japan))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="59" type="distribution">
<paragraph blockId="59.[136,1452,770,1409]" pageId="59">
<emphasis bold="true" box="[189,345,1058,1084]" pageId="59">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[512,607,1058,1084]" name="Vietnam" pageId="59">Vietnam</collectingCountry>
(currently known as Hon Ba Nature Reserve of
<collectingRegion box="[1164,1403,1058,1085]" country="Vietnam" name="Khanh Hoa" pageId="59">Khanh Hoa Province</collectingRegion>
and Bidoup Nui Ba National Park of
<collectingRegion box="[505,737,1094,1121]" country="Vietnam" name="Lam Dong" pageId="59">Lam Dong Province</collectingRegion>
).
</paragraph>
<paragraph blockId="59.[136,1452,770,1409]" box="[189,1451,1130,1157]" pageId="59">
<emphasis bold="true" box="[189,423,1130,1156]" pageId="59">Habitat &amp; Ecology:</emphasis>
—Growing in secondary and primary forests, on the edge of the forest, at
<quantity box="[1251,1392,1130,1157]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="59" unit="m" value="1100.0" valueMax="1500.0" valueMin="700.0">7001500 m</quantity>
elev.
</paragraph>
<paragraph blockId="59.[136,1452,770,1409]" box="[189,882,1166,1193]" pageId="59">
<emphasis bold="true" box="[189,323,1166,1192]" pageId="59">Phenology:</emphasis>
—Flowering and fruiting from June to December.
</paragraph>
<paragraph blockId="59.[136,1452,770,1409]" box="[189,596,1202,1231]" pageId="59">
<emphasis bold="true" box="[189,405,1202,1228]" pageId="59">Vernacular name:</emphasis>
—Xú hưƠng hôi.
</paragraph>
<paragraph blockId="59.[136,1452,770,1409]" pageId="59">
<emphasis bold="true" box="[189,265,1238,1264]" pageId="59">Notes:</emphasis>
<taxonomicName box="[294,568,1238,1264]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="59" phylum="Tracheophyta" rank="species" species="foetidissimus">
<emphasis box="[294,568,1238,1264]" italics="true" pageId="59">Lasianthus foetidissimus</emphasis>
</taxonomicName>
is similar to
<taxonomicName authority="V. S. Dang &amp; Naiki (2019: 34)" authorityName="V. S. Dang &amp; Naiki" authorityPageNumber="34" authorityYear="2019" box="[708,1202,1238,1264]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="59" phylum="Tracheophyta" rank="species" species="fasipanensis">
<emphasis box="[708,874,1238,1264]" italics="true" pageId="59">L. fasipanensis</emphasis>
V.S.
<bibRefCitation author="Dang, V. S. &amp; Naiki, A." box="[921,1202,1238,1264]" pageId="59" pagination="33 - 40" refId="ref54157" refString="Dang, V. S. &amp; Naiki, A. (2019) Two new species of Lasianthus (Rubiaceae) from Hoang Lien National Park, northern Vietnam. Phytotaxa 423 (1): 33 - 40. https: // doi. org / 10.11646 / phytotaxa. 423.1.4" type="journal article" year="2019">Dang &amp; Naiki (2019: 34)</bibRefCitation>
</taxonomicName>
which was discovered at Hoang Lien National Park of northern
<collectingCountry box="[593,688,1274,1300]" name="Vietnam" pageId="59">Vietnam</collectingCountry>
by
<bibRefCitation author="Dang, V. S. &amp; Naiki, A." box="[729,971,1274,1301]" pageId="59" pagination="33 - 40" refId="ref54157" refString="Dang, V. S. &amp; Naiki, A. (2019) Two new species of Lasianthus (Rubiaceae) from Hoang Lien National Park, northern Vietnam. Phytotaxa 423 (1): 33 - 40. https: // doi. org / 10.11646 / phytotaxa. 423.1.4" type="journal article" year="2019">Dang &amp; Naiki (2019)</bibRefCitation>
, based on its pedunculate congested cymes and flower color at anthesis, but differs in its acute or slightly rounded leaf base (vs. oblique to obtuse), longer petioles (
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long vs.
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long), smaller calyx (
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long vs.
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long for tubes and
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long vs.
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long lobes), yellow to yellowish orange fruits (vs. blue) and fruits with 5 pyrenes (vs. 4 pyrenes).
</paragraph>
</subSubSection>
</treatment>
</document>