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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFA4FFE35AB6FA0B426BC464" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus fordii Hance 1885" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571664" ID-Zenodo-Dep="7571664" LSID="urn:lsid:plazi:treatment:43284262FFA4FFE35AB6FA0B426BC464" httpUri="http://treatment.plazi.org/id/43284262FFA4FFE35AB6FA0B426BC464" lastPageId="61" pageId="59">
<subSubSection box="[136,950,1454,1481]" pageId="59" type="nomenclature">
<paragraph blockId="59.[136,1452,1454,1660]" box="[136,950,1454,1481]" pageId="59">
<emphasis bold="true" box="[136,171,1454,1480]" pageId="59">23.</emphasis>
<taxonomicName ID-CoL="3SCL8" ID-ENA="1406743" authority="Hance (1885: 324)" authorityName="Hance" authorityPageNumber="324" authorityYear="1885" box="[178,595,1454,1481]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="59" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis bold="true" box="[178,378,1454,1481]" italics="true" pageId="59">Lasianthus fordii</emphasis>
Hance (1885: 324)
</taxonomicName>
<emphasis bold="true" box="[595,623,1454,1480]" pageId="59"></emphasis>
<figureCitation box="[623,702,1454,1480]" captionStart="FIGURE 44" captionStartId="61.[136,229,1750,1772]" captionTargetBox="[170,1412,194,1723]" captionTargetId="figure-17@61.[166,1418,193,1726]" captionTargetPageId="61" captionText="FIGURE 44. Lasianthus fordii Hance (from Dang 235, VNM and the herbarium of Iriomote Station (Japan)): A. Branch with flowers and fruits, B. Stipules, C. Abaxial leaf surfaces, DF. Inflorescence showing side view buds and flowers, G. Flower, young and mature fruits, H. Mature fruits, I. Cross section of fruit with pyrenes." figureDoi="http://doi.org/10.5281/zenodo.7571684" httpUri="https://zenodo.org/record/7571684/files/figure.png" pageId="59">Fig. 44</figureCitation>
(plate),
<figureCitation box="[796,876,1454,1480]" captionStart="FIGURE 42" captionStartId="58.[136,229,1789,1811]" captionTargetBox="[173,1409,203,1757]" captionTargetId="figure-25@58.[165,1419,193,1765]" captionTargetPageId="58" captionText="FIGURE 42. Distribution maps of Lasianthus fansipanensis, L. foetidissimus, L. fordii and L. formosensis." figureDoi="http://doi.org/10.5281/zenodo.7571678" httpUri="https://zenodo.org/record/7571678/files/figure.png" pageId="59">Fig. 42</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="59" type="materials_examined">
<paragraph blockId="59.[136,1452,1454,1660]" pageId="59">
<emphasis bold="true" box="[136,194,1494,1515]" pageId="59">Type:</emphasis>
<materialsCitation collectingDate="1883-08" collectionCode="BM" collectorName="S. Coll." country="China" elevation="244" location="Lo Fau Shan" pageId="59" specimenCode="BM000945611" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
<collectingCountry box="[219,297,1494,1515]" name="China" pageId="59">CHINA</collectingCountry>
.
<collectingRegion box="[312,426,1494,1516]" country="China" name="Guangdong" pageId="59">Guangdong</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE35AB6FA0B426BC464:CE5EA5AFFFA4FFE55B84FA734264C9A4" box="[442,578,1494,1516]" country="China" name="Lo Fau Shan" pageId="59" stateProvince="Guangdong">Lo Fau Shan</location>
,
<quantity box="[594,661,1494,1515]" metricMagnitude="2" metricUnit="m" metricValue="2.44" pageId="59" unit="m" value="244.0">
<elevation box="[594,661,1494,1515]" metricMagnitude="2" metricUnit="m" metricValue="2.44" pageId="59" unit="m" value="244.0">244 m</elevation>
</quantity>
elev.,
<date box="[733,864,1494,1515]" pageId="59" value="1883-08">
<collectingDate box="[733,864,1494,1515]" pageId="59" value="1883-08">August 1883</collectingDate>
</date>
,
<emphasis box="[880,993,1494,1515]" italics="true" pageId="59">
<collectorName box="[880,958,1494,1515]" pageId="59">S. Coll.</collectorName>
66
</emphasis>
(
<typeStatus box="[1011,1097,1494,1516]" pageId="59">holotype</typeStatus>
BM! [
<specimenCode box="[1171,1324,1494,1516]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="59">BM000945611</specimenCode>
]; isotype K! [K000763821])
</materialsCitation>
.
</paragraph>
<paragraph blockId="59.[136,1452,1454,1660]" pageId="59">
<taxonomicName authority="Elmer (1934: 3209)" authorityName="Elmer" authorityPageNumber="3209" authorityYear="1934" box="[136,582,1566,1588]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="59" phylum="Tracheophyta" rank="species" species="zambalensis">
<emphasis box="[136,369,1566,1587]" italics="true" pageId="59">Lasianthus zambalensis</emphasis>
<materialsCitation box="[380,582,1566,1588]" collectingDate="1927-05" county="Camp Stotsenburg" location="Luzon" municipality="Pinatubo mountain" pageId="59" specimenCount="1" stateProvince="Pampanga">Elmer (1934: 3209)</materialsCitation>
</taxonomicName>
.
<typeStatus box="[596,645,1566,1587]" pageId="59">Type</typeStatus>
:—PHILLIPPINES.
<collectingCounty box="[847,1032,1566,1588]" pageId="59">Camp Stotsenburg</collectingCounty>
,
<collectingMunicipality box="[1046,1235,1566,1588]" pageId="59">Pinatubo mountain</collectingMunicipality>
,
<collectingRegion box="[1249,1448,1566,1588]" country="Philippines" name="Pampanga" pageId="59">Pampanga Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE35AB6FA0B426BC464:CE5EA5AFFFA4FFE55A83F9E740DBCA1F" box="[189,253,1602,1623]" county="Camp Stotsenburg" municipality="Pinatubo mountain" name="Luzon" pageId="59" stateProvince="Pampanga">Luzon</location>
,
<date box="[263,363,1602,1623]" pageId="59" value="1927-05">
<collectingDate box="[263,363,1602,1623]" pageId="59" value="1927-05">May 1927</collectingDate>
</date>
,
<emphasis box="[373,497,1602,1623]" italics="true" pageId="59">Elmer 22045</emphasis>
(isotype K! [K000777092]).
</paragraph>
<paragraph blockId="59.[136,1452,1454,1660]" box="[136,1444,1638,1660]" pageId="59">
<taxonomicName authority="Pierre ex Pit. (1924: 392)" authorityName="Pit." authorityPageNumber="392" authorityYear="1924" box="[136,618,1638,1660]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="59" phylum="Tracheophyta" rank="species" species="kamputensis">
<emphasis box="[136,366,1638,1659]" italics="true" pageId="59">Lasianthus kamputensis</emphasis>
<materialsCitation box="[372,618,1638,1660]" collectingDate="1913-04-03" country="Vietnam" location="Type" pageId="59" specimenCount="1" stateProvince="Quang Tri" typeStatus="syntype">Pierre ex Pit. (1924: 392)</materialsCitation>
</taxonomicName>
.
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE35AB6FA0B426BC464:CE5EA5AFFFA4FFE5584DF9C34283CA33" box="[627,677,1638,1659]" country="Vietnam" name="Type" pageId="59" stateProvince="Quang Tri">Type</location>
:—
<collectingCountry box="[705,819,1638,1659]" name="Vietnam" pageId="59">VIETNAM</collectingCountry>
,
<collectingRegion box="[828,1020,1638,1660]" country="Vietnam" name="Quang Tri" pageId="59">Quang Tri Province</collectingRegion>
,
<date box="[1029,1153,1638,1660]" pageId="59" value="1913-04-03">
<collectingDate box="[1029,1153,1638,1660]" pageId="59" value="1913-04-03">3 April 1913</collectingDate>
</date>
,
<emphasis box="[1163,1314,1638,1659]" italics="true" pageId="59">Eberhardt 2050</emphasis>
(
<typeStatus box="[1326,1402,1638,1659]" pageId="59">syntype</typeStatus>
P!).
</paragraph>
<paragraph blockId="59.[136,1452,1706,2093]" pageId="59">
<emphasis bold="true" box="[136,393,1706,1732]" pageId="59">
<collectingRegion box="[136,262,1706,1732]" pageId="59">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="2018-11-13" collectionCode="VNM" collectorName="La Hien Commune, P &amp; Dang V. S. &amp; Naiki A. &amp; Dang &amp; Japan" country="Vietnam" county="Vo Nhai District" elevation="925" location="Iriomote Station" municipality="Tam Dao National Park" pageId="59" specimenCount="1" stateProvince="Thai Nguyen">
<collectingCountry box="[422,554,1706,1732]" name="Vietnam" pageId="59">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[563,685,1706,1732]" pageId="59">Northern:</emphasis>
<collectingRegion box="[691,946,1706,1733]" country="Vietnam" name="Thai Nguyen" pageId="59">Thai Nguyen Province</collectingRegion>
,
<collectingCounty box="[956,1141,1706,1733]" pageId="59">Vo Nhai District</collectingCounty>
,
<collectorName box="[1151,1395,1706,1733]" pageId="59">
La Hien Commune,
<emphasis box="[1378,1395,1706,1732]" italics="true" pageId="59">P</emphasis>
</collectorName>
3886 (
<collectionCode box="[144,215,1742,1768]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="59" type="Herbarium">VNM</collectionCode>
);
<collectingRegion box="[235,463,1742,1768]" country="Vietnam" name="Vinh Phuc" pageId="59">Vinh Phuc Province</collectingRegion>
,
<collectingMunicipality box="[474,744,1742,1769]" pageId="59">Tam Dao National Park</collectingMunicipality>
,
<quantity box="[756,828,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="59" unit="m" value="925.0">
<elevation box="[756,828,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="59" unit="m" value="925.0">925 m</elevation>
</quantity>
elev.,
<date box="[902,1122,1742,1768]" pageId="59" value="2018-11-13">
<collectingDate box="[902,1122,1742,1768]" pageId="59" value="2018-11-13">13 November 2018</collectingDate>
</date>
,
<emphasis italics="true" pageId="59">
<collectorName box="[1134,1246,1743,1769]" pageId="59">Dang V.S.</collectorName>
&amp;
<collectorName box="[1282,1376,1742,1769]" pageId="59">Naiki A.</collectorName>
,
<collectorName box="[1389,1451,1743,1768]" pageId="59">Dang</collectorName>
235
</emphasis>
(
<collectionCode box="[193,263,1778,1804]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="59" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE35AB6FA0B426BC464:CE5EA5AFFFA4FFE55BE8F95742ABCB44" box="[470,653,1778,1804]" country="Vietnam" county="Vo Nhai District" municipality="Tam Dao National Park" name="Iriomote Station" pageId="59" stateProvince="Thai Nguyen">Iriomote Station</location>
(
<collectorName box="[669,738,1778,1804]" pageId="59">Japan</collectorName>
))
</materialsCitation>
.
<materialsCitation collectingDate="2015-07-26" collectingDateMax="2018-04-19" collectingDateMin="2015-07-26" collectionCode="DLU, FU, VNM" collectorName="Yahara T. &amp; Toyama H. &amp; Tagane S. &amp; Nguyen N. &amp; Yang C. J. &amp; Nguyen H. V &amp; Dang V. S. &amp; Hoang N. T. B. N. &amp; Dang" country="Vietnam" county="Vu Quang National Park" elevation="914" location="Hon Ba Nature Reserve" municipality="Iriomote Station" pageId="59" specimenCount="1" stateProvince="Ha Tinh">
<emphasis bold="true" box="[767,868,1778,1804]" pageId="59">Central:</emphasis>
<collectingRegion box="[875,1077,1778,1805]" country="Vietnam" name="Ha Tinh" pageId="59">Ha Tinh Province</collectingRegion>
,
<collectingCounty box="[1088,1368,1778,1805]" pageId="59">Vu Quang National Park</collectingCounty>
,
<quantity box="[1380,1452,1778,1805]" metricMagnitude="2" metricUnit="m" metricValue="9.14" pageId="59" unit="m" value="914.0">
<elevation box="[1380,1452,1778,1805]" metricMagnitude="2" metricUnit="m" metricValue="9.14" pageId="59" unit="m" value="914.0">914 m</elevation>
</quantity>
elev.,
<date box="[200,344,1814,1841]" pageId="59" value="2015-07-26">
<collectingDate box="[200,344,1814,1841]" pageId="59" value="2015-07-26">26 July 2015</collectingDate>
</date>
,
<emphasis box="[353,1165,1814,1841]" italics="true" pageId="59">
<collectorName box="[353,461,1814,1840]" pageId="59">Yahara T.</collectorName>
,
<collectorName box="[471,594,1815,1840]" pageId="59">Toyama H.</collectorName>
,
<collectorName box="[603,712,1815,1841]" pageId="59">Tagane S.</collectorName>
,
<collectorName box="[723,842,1815,1840]" pageId="59">Nguyen N.</collectorName>
,
<collectorName box="[851,959,1815,1841]" pageId="59">Yang C.J.</collectorName>
,
<collectorName box="[969,1109,1814,1840]" pageId="59">Nguyen H. V</collectorName>
3778
</emphasis>
(
<collectionCode box="[1178,1240,1814,1840]" pageId="59">DLU</collectionCode>
,
<collectionCode box="[1249,1288,1815,1841]" country="China" name="Fudan University, Department of Biology" pageId="59">FU</collectionCode>
, the herbarium of
<collectingMunicipality box="[167,349,1850,1876]" pageId="59">Iriomote Station</collectingMunicipality>
(Japan));
<collectingRegion box="[464,702,1850,1877]" country="Vietnam" name="Khanh Hoa" pageId="59">Khanh Hoa Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE35AB6FA0B426BC464:CE5EA5AFFFA4FFE558F4F89E43FECB1D" box="[714,984,1851,1877]" country="Vietnam" county="Vu Quang National Park" municipality="Iriomote Station" name="Hon Ba Nature Reserve" pageId="59" stateProvince="Ha Tinh">Hon Ba Nature Reserve</location>
,
<quantity box="[996,1067,1850,1877]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="59" unit="m" value="900.0">
<elevation box="[996,1067,1850,1877]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="59" unit="m" value="900.0">900 m</elevation>
</quantity>
elev.,
<date box="[1141,1300,1850,1876]" pageId="59" value="2018-04-19">
<collectingDate box="[1141,1300,1850,1876]" pageId="59" value="2018-04-19">19 April 2018</collectingDate>
</date>
,
<emphasis italics="true" pageId="59">
<collectorName box="[1311,1423,1851,1877]" pageId="59">Dang V.S.</collectorName>
&amp;
<collectorName box="[136,318,1887,1912]" pageId="59">Hoang N.T.B.N.</collectorName>
,
<collectorName box="[329,391,1887,1912]" pageId="59">Dang</collectorName>
183
</emphasis>
(
<collectionCode box="[455,526,1886,1912]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="59" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation box="[545,1328,1886,1913]" collectingDate="1909-06" country="Vietnam" location="Bien Hoa District" pageId="59" specimenCount="1" stateProvince="Dong Nai">
<emphasis bold="true" box="[545,665,1886,1912]" pageId="59">Southern:</emphasis>
<collectingRegion box="[672,891,1886,1913]" country="Vietnam" name="Dong Nai" pageId="59">Dong Nai Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA4FFE35AB6FA0B426BC464:CE5EA5AFFFA4FFE559B8F8FB4469CB30" box="[902,1103,1886,1913]" country="Vietnam" name="Bien Hoa District" pageId="59" stateProvince="Dong Nai">Bien Hoa District</location>
,
<date box="[1114,1231,1886,1913]" pageId="59" value="1909-06">
<collectingDate box="[1114,1231,1886,1913]" pageId="59" value="1909-06">June 1909</collectingDate>
</date>
,
<taxonomicName box="[1242,1284,1887,1913]" class="Insecta" family="Zygaenidae" genus="Sno" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="59" phylum="Arthropoda" rank="genus">
<emphasis box="[1242,1284,1887,1913]" italics="true" pageId="59">Sno</emphasis>
</taxonomicName>
(P)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="61" pageId="59" type="distribution">
<paragraph blockId="59.[136,1452,1706,2093]" pageId="59">
<emphasis bold="true" box="[189,345,1922,1948]" pageId="59">Distribution:</emphasis>
—Widespread in tropical and subtropical regions in Asia (
<collectingCountry box="[981,1101,1922,1948]" name="Cambodia" pageId="59">Cambodia</collectingCountry>
,
<collectingCountry box="[1111,1181,1922,1948]" name="China" pageId="59">China</collectingCountry>
,
<collectingCountry box="[1190,1250,1922,1948]" name="India" pageId="59">India</collectingCountry>
,
<collectingCountry box="[1259,1371,1922,1948]" name="Indonesia" pageId="59">Indonesia</collectingCountry>
,
<collectingCountry box="[1381,1448,1923,1949]" name="Japan" pageId="59">Japan</collectingCountry>
,
<collectingCountry box="[136,357,1958,1985]" name="Papua New Guinea" pageId="59">Papua New Guinea</collectingCountry>
,
<collectingCountry box="[368,497,1958,1984]" name="Philippines" pageId="59">Philippines</collectingCountry>
,
<collectingCountry box="[508,613,1958,1984]" name="Thailand" pageId="59">Thailand</collectingCountry>
). In
<collectingCountry box="[662,760,1958,1984]" name="Vietnam" pageId="59">Vietnam</collectingCountry>
,
<taxonomicName box="[772,963,1958,1984]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="59" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis box="[772,963,1958,1984]" italics="true" pageId="59">Lasianthus fordii</emphasis>
</taxonomicName>
occurs in
<collectingRegion box="[1081,1181,1959,1985]" country="Vietnam" name="Bac Kan" pageId="59">Bac Kan</collectingRegion>
,
<collectingRegion box="[1194,1307,1958,1985]" country="Vietnam" name="Dong Nai" pageId="59">Dong Nai</collectingRegion>
,
<collectingRegion box="[1319,1407,1958,1984]" country="Vietnam" name="Gia Lai" pageId="59">Gia Lai</collectingRegion>
,
<collectingRegion country="Vietnam" name="Ha Tinh" pageId="59">Ha Tinh</collectingRegion>
,
<collectingRegion box="[202,333,1994,2021]" country="Vietnam" name="Khanh Hoa" pageId="59">Khanh Hoa</collectingRegion>
,
<collectingRegion box="[344,468,1995,2021]" country="Vietnam" name="Lam Dong" pageId="59">Lam Dong</collectingRegion>
,
<collectingRegion box="[479,571,1994,2021]" country="Vietnam" name="Lao Cai" pageId="59">Lao Cai</collectingRegion>
,
<collectingRegion box="[582,699,1994,2020]" country="Vietnam" name="Quang Tri" pageId="59">Quang Tri</collectingRegion>
,
<collectingRegion box="[710,818,1994,2021]" country="Vietnam" name="Tay Ninh" pageId="59">Tay Ninh</collectingRegion>
,
<collectingRegion box="[828,977,1994,2021]" country="Vietnam" name="Thai Nguyen" pageId="59">Thai Nguyen</collectingRegion>
, Thua Thien Hue and
<collectingRegion box="[1226,1347,1994,2020]" country="Vietnam" name="Vinh Phuc" pageId="59">Vinh Phuc</collectingRegion>
.
</paragraph>
<paragraph blockId="59.[136,1452,1706,2093]" box="[189,1317,2030,2057]" pageId="59">
<emphasis bold="true" box="[189,423,2030,2056]" pageId="59">Habitat &amp; Ecology:</emphasis>
—Growing under shade or on the edge of evergreen forest, at
<quantity box="[1117,1258,2030,2057]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="59" unit="m" value="900.0" valueMax="1200.0" valueMin="600.0">6001200 m</quantity>
elev.
</paragraph>
<paragraph blockId="59.[136,1452,1706,2093]" box="[189,880,2066,2093]" pageId="59">
<emphasis bold="true" box="[189,323,2066,2092]" pageId="59">Phenology:</emphasis>
—Flowering and fruiting from July to November.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571682" ID-Zenodo-Dep="7571682" httpUri="https://zenodo.org/record/7571682/files/figure.png" pageId="60" startId="60.[136,229,1763,1785]" targetBox="[172,1416,196,1736]" targetPageId="60">
<paragraph blockId="60.[136,1451,1763,1857]" pageId="60">
<emphasis bold="true" box="[136,265,1763,1785]" pageId="60">FIGURE 43.</emphasis>
<taxonomicName authority="A. Chev. ex Pit." authorityName="Pit." box="[271,660,1763,1785]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="60" phylum="Tracheophyta" rank="species" species="foetidissimus">
<emphasis box="[271,507,1763,1784]" italics="true" pageId="60">Lasianthus foetidissimus</emphasis>
A.Chev. ex Pit.
</taxonomicName>
(from
<emphasis box="[725,820,1763,1785]" italics="true" pageId="60">Dang 343</emphasis>
, VNM and the herbarium of Iriomote Station (Japan)):
<emphasis bold="true" box="[1353,1377,1763,1784]" pageId="60">A.</emphasis>
Branch with flowers and bubs,
<emphasis bold="true" box="[358,380,1799,1820]" pageId="60">B.</emphasis>
Stipules,
<emphasis bold="true" box="[474,497,1799,1821]" pageId="60">C.</emphasis>
Abaxial leaf surfaces,
<emphasis bold="true" box="[714,765,1799,1820]" pageId="60">DE.</emphasis>
Inflorescence showing side view buds and flower,
<emphasis bold="true" box="[1251,1269,1799,1820]" pageId="60">F.</emphasis>
Flower,
<emphasis bold="true" box="[1353,1378,1799,1821]" pageId="60">G.</emphasis>
Flower buds and fruit,
<emphasis bold="true" box="[281,306,1835,1856]" pageId="60">H.</emphasis>
Young fruits,
<emphasis bold="true" box="[443,458,1835,1856]" pageId="60">I.</emphasis>
Mature fruits.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571684" ID-Zenodo-Dep="7571684" httpUri="https://zenodo.org/record/7571684/files/figure.png" pageId="61" startId="61.[136,229,1750,1772]" targetBox="[170,1412,194,1723]" targetPageId="61">
<paragraph blockId="61.[136,1452,1750,1844]" pageId="61">
<emphasis bold="true" box="[136,265,1750,1772]" pageId="61">FIGURE 44.</emphasis>
<taxonomicName authority="Hance" authorityName="Hance" box="[270,500,1750,1772]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="61" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis box="[270,433,1750,1771]" italics="true" pageId="61">Lasianthus fordii</emphasis>
Hance
</taxonomicName>
(from
<emphasis box="[565,660,1751,1772]" italics="true" pageId="61">Dang 235</emphasis>
, VNM and the herbarium of Iriomote Station (Japan)):
<emphasis bold="true" box="[1188,1212,1750,1771]" pageId="61">A.</emphasis>
Branch with flowers and fruits,
<emphasis bold="true" box="[199,221,1787,1808]" pageId="61">B.</emphasis>
Stipules,
<emphasis bold="true" box="[316,339,1786,1808]" pageId="61">C.</emphasis>
Abaxial leaf surfaces,
<emphasis bold="true" box="[558,605,1787,1808]" pageId="61">DF.</emphasis>
Inflorescence showing side view buds and flowers,
<emphasis bold="true" box="[1106,1131,1786,1808]" pageId="61">G.</emphasis>
Flower, young and mature fruits,
<emphasis bold="true" box="[136,161,1823,1844]" pageId="61">H.</emphasis>
Mature fruits,
<emphasis bold="true" box="[305,320,1823,1844]" pageId="61">I.</emphasis>
Cross section of fruit with pyrenes.
</paragraph>
</caption>
<paragraph blockId="61.[136,1452,1890,2092]" box="[189,668,1890,1919]" pageId="61">
<emphasis bold="true" box="[189,405,1890,1916]" pageId="61">Vernacular name:</emphasis>
—Xú hưƠng lưỡi vành.
</paragraph>
<paragraph blockId="61.[136,1452,1890,2092]" pageId="61">
<emphasis bold="true" box="[189,265,1925,1951]" pageId="61">Notes:</emphasis>
<taxonomicName box="[294,490,1925,1951]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="61" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis box="[294,490,1925,1951]" italics="true" pageId="61">Lasianthus fordii</emphasis>
</taxonomicName>
was clarified in the revision by
<bibRefCitation author="Zhu, H. &amp; Roos, M. C. &amp; Ridsdale, C. E." box="[889,1096,1925,1951]" pageId="61" pagination="1 - 102" refId="ref55608" refString="Zhu, H., Roos, M. C. &amp; Ridsdale, C. E. (2012) A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae). Blumea 57: 1 - 102. https: // doi. org / 10.3767 / 000651912 X 652012" type="journal article" year="2012">
Zhu
<emphasis box="[947,1008,1925,1951]" italics="true" pageId="61">et al.</emphasis>
(2012)
</bibRefCitation>
and a Vietnamese species,
<taxonomicName authority="Pierre ex Pit. (1924: 392)" authorityName="Pit." authorityPageNumber="392" authorityYear="1924" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="61" phylum="Tracheophyta" rank="species" species="kamputensis">
<emphasis italics="true" pageId="61">L. kamputensis</emphasis>
Pierre ex Pit. (1924: 392)
</taxonomicName>
was treated as the synonym of
<taxonomicName box="[918,1008,1960,1986]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="61" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis box="[918,1008,1960,1986]" italics="true" pageId="61">L. fordii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="61.[136,1452,1890,2092]" pageId="61">
<taxonomicName box="[189,378,1996,2022]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="61" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis box="[189,378,1996,2022]" italics="true" pageId="61">Lasianthus fordii</emphasis>
</taxonomicName>
is very close to
<taxonomicName authority="Matsum. (1901: 17)" authorityName="Matsum." authorityPageNumber="17" authorityYear="1901" box="[547,931,1996,2022]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="61" phylum="Tracheophyta" rank="species" species="formosensis">
<emphasis box="[547,708,1996,2022]" italics="true" pageId="61">L. formosensis</emphasis>
Matsum. (1901: 17)
</taxonomicName>
based on its shape of leaf blades, inflorescences and indumentum, but differs from the latter in having triangular calyx lobes obviously shorter than the tube (vs. lanceolate calyx lobes longer than tube).
</paragraph>
</subSubSection>
</treatment>
</document>