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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FF87FFC75AB6FC834386CD24" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus bachmaensis V. S. Dang &amp; Naiki 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571564" ID-Zenodo-Dep="7571564" LSID="urn:lsid:plazi:treatment:43284262FF87FFC75AB6FC834386CD24" httpUri="http://treatment.plazi.org/id/43284262FF87FFC75AB6FC834386CD24" lastPageId="25" pageId="24">
<subSubSection box="[136,1114,806,833]" pageId="24" type="nomenclature">
<paragraph blockId="24.[136,1452,806,940]" box="[136,1114,806,833]" pageId="24">
<emphasis bold="true" box="[136,157,806,832]" pageId="24">3.</emphasis>
<taxonomicName authority="V.S.Dang &amp; Naiki" authorityName="V. S. Dang &amp; Naiki" authorityYear="2023" box="[164,664,806,833]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="24" phylum="Tracheophyta" rank="species" species="bachmaensis" status="sp. nov.">
<emphasis bold="true" box="[164,449,806,833]" italics="true" pageId="24">Lasianthus bachmaensis</emphasis>
V.S.Dang &amp; Naiki
</taxonomicName>
,
<emphasis box="[675,760,807,832]" italics="true" pageId="24">
<taxonomicNameLabel box="[675,760,807,832]" pageId="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<figureCitation box="[790,867,806,832]" captionStart="FIGURE 17" captionStartId="25.[136,229,1973,1995]" captionTargetBox="[169,1415,393,1945]" captionTargetId="figure-108@25.[166,1418,392,1948]" captionTargetPageId="25" captionText="FIGURE 17. Lasianthus bachmaensis V.S.Dang &amp; Naiki (from the holotype, VNM): A. Branch with flower buds, B. Close-up of young branch and leaves, C. Stipules, D. Abaxial leaf surfaces, EG. Inflorescence showing lateral view flower buds, H. Young fruit and buds, I. Mature fruits, J. Seeds." figureDoi="http://doi.org/10.5281/zenodo.7571600" httpUri="https://zenodo.org/record/7571600/files/figure.png" pageId="24">Fig. 17</figureCitation>
(plate),
<figureCitation box="[961,1040,806,833]" captionStart="FIGURE 15" captionStartId="22.[136,229,1778,1800]" captionTargetBox="[173,1409,201,1747]" captionTargetId="figure-24@22.[165,1418,193,1754]" captionTargetPageId="22" captionText="FIGURE 15. Distribution maps of Lasianthus annamicus, L. attenuatus, L. bachmaensis and L. bidoupensis." figureDoi="http://doi.org/10.5281/zenodo.7571596" httpUri="https://zenodo.org/record/7571596/files/figure.png" pageId="24">Fig. 15</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="24" type="materials_examined">
<paragraph blockId="24.[136,1452,806,940]" pageId="24">
<emphasis bold="true" box="[136,194,846,867]" pageId="24">Type:</emphasis>
<materialsCitation collectingDate="2019-07-26" collectionCode="VNM" collectorName="Dang V. S. &amp; Naiki A. &amp; Dang" country="Vietnam" county="Iriomote Station" elevation="1183" location="Thua Thien Hue Province" municipality="Bach Ma National Park" pageId="24" specimenCode="VNM00042867, VNM00042868, VNM00042869" specimenCount="1" typeStatus="holotype">
<collectingCountry box="[219,332,846,867]" name="Vietnam" pageId="24">VIETNAM</collectingCountry>
.
<location LSID="urn:lsid:plazi:treatment:43284262FF87FFC75AB6FC834386CD24:CE5EA5AFFF87FFC65B68FCEB4270CF2C" box="[342,598,846,868]" country="Vietnam" county="Iriomote Station" municipality="Bach Ma National Park" name="Thua Thien Hue Province" pageId="24">Thua Thien Hue Province</location>
,
<collectingMunicipality box="[609,842,846,868]" pageId="24">Bach Ma National Park</collectingMunicipality>
,
<quantity box="[854,927,846,867]" metricMagnitude="3" metricUnit="m" metricValue="1.183" pageId="24" unit="m" value="1183.0">
<elevation box="[854,927,846,867]" metricMagnitude="3" metricUnit="m" metricValue="1.183" pageId="24" unit="m" value="1183.0">1183 m</elevation>
</quantity>
elev., 16°1176.03N, 107°5084.70E, in evergreen forest,
<date box="[261,396,882,904]" pageId="24" value="2019-07-26">
<collectingDate box="[261,396,882,904]" pageId="24" value="2019-07-26">26 July 2019</collectingDate>
</date>
,
<emphasis box="[410,749,882,904]" italics="true" pageId="24">
<collectorName box="[410,511,882,904]" pageId="24">Dang V.S.</collectorName>
&amp;
<collectorName box="[550,635,882,903]" pageId="24">Naiki A.</collectorName>
,
<collectorName box="[649,702,883,904]" pageId="24">Dang</collectorName>
406
</emphasis>
(
<typeStatus box="[766,852,882,904]" pageId="24">holotype</typeStatus>
<collectionCode box="[862,923,882,903]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="24" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[942,1102,882,904]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="24" type="Herbarium">VNM00042867</specimenCode>
]; isotypes
<collectionCode box="[1211,1272,882,903]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="24" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[1291,1447,882,904]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="24" type="Herbarium">VNM00042868</specimenCode>
,
<specimenCode box="[189,346,918,940]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="24" type="Herbarium">VNM00042869</specimenCode>
], the herbarium of
<collectingCounty box="[527,684,918,940]" pageId="24">Iriomote Station</collectingCounty>
(Japan))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="24" type="diagnosis">
<paragraph blockId="24.[136,1452,986,2093]" pageId="24">
<emphasis bold="true" box="[136,261,986,1012]" pageId="24">Diagnosis:</emphasis>
—This new species is morphologically similar to
<taxonomicName authority="Naiki, Tagane &amp; Yahara (2015: 173)" authorityName="Naiki, Tagane &amp; Yahara" authorityPageNumber="173" authorityYear="2015" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="24" phylum="Tracheophyta" rank="species" species="stephanocalycinus">
<emphasis box="[830,1169,986,1012]" italics="true" pageId="24">Lasianthus stephanocalycinus</emphasis>
<bibRefCitation author="Naiki, A. &amp; Tagane, S. &amp; Chhang, P. &amp; Toyama, H. &amp; Zhu, H. &amp; Dang, V. S. &amp; Yahara, T." pageId="24" pagination="153 - 179" refId="ref54656" refString="Naiki, A., Tagane, S., Chhang, P., Toyama, H., Zhu, H., Dang, V. S. &amp; Yahara, T. (2015) Flora of Bokor National Park, Cambodia II: four new species and nine new records of Lasianthus (Rubiaceae) from Cambodia. Acta Phytotaxonomica et Geobotanica 66 (3): 153 - 179. https: // doi. org / 10.18942 / apg. KJ 00010115702" type="journal article" year="2015">Naiki, Tagane &amp; Yahara (2015: 173)</bibRefCitation>
</taxonomicName>
based on leaf shape and fruit color, but differs from the latter in having smaller leaves (1012 ×
<quantity box="[1347,1451,1022,1049]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="24" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">22.5 cm</quantity>
vs. 89.5 ×
<quantity box="[267,374,1058,1085]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="24" unit="cm" value="3.25" valueMax="3.5" valueMin="3.0">33.5 cm</quantity>
), shorter stipules (
<quantity box="[579,693,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="24" unit="mm" value="1.4" valueMax="1.8" valueMin="1.0">11.8 mm</quantity>
long vs.
<quantity box="[795,887,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long) and smaller calyx (
<quantity box="[1171,1286,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="24" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
long vs.
<quantity box="[1387,1452,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="24" unit="mm" value="2.0">2 mm</quantity>
long for tubes and
<quantity box="[346,481,1094,1121]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="24" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.52.5 mm</quantity>
long vs.
<quantity box="[583,697,1094,1121]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="24" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.53 mm</quantity>
long for lobes).
</paragraph>
</subSubSection>
<subSubSection pageId="24" type="description">
<paragraph blockId="24.[136,1452,986,2093]" pageId="24">
<emphasis bold="true" box="[189,337,1130,1156]" pageId="24">Description:</emphasis>
—Shrubs, evergreen,
<quantity box="[593,691,1130,1157]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="24" unit="m" value="1.75" valueMax="2.5" valueMin="1.0">12.5 m</quantity>
tall; branches and branchlets terete,
<quantity box="[1137,1236,1130,1157]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="24" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
diam., appressedpubescent or strigose, internodes
<quantity box="[503,585,1166,1193]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="24" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">13 cm</quantity>
long. Leaves opposite, blades elliptic or elliptic-lanceolate, 1012 ×
<quantity box="[1342,1447,1166,1193]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="24" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">22.5 cm</quantity>
, coriaceous, yellowish brown when dry, adaxially glabrous, abaxially sparsely strigose on nerves, apex narrowly acute to acuminate, aristate
<quantity box="[352,445,1238,1265]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="24" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long, base cuneate or acute, margin entire; midrib slightly prominent adaxially, prominent abaxially; secondary veins 45 on each side of midrib, ascending at an angle of 4060° from the midrib, curved to the margin, slightly distinct on both surfaces; tertiary veins parallel; petioles
<quantity box="[953,1046,1310,1337]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="24" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
long, appressed-pubescent. Stipules triangular,
<quantity box="[258,373,1346,1373]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="24" unit="mm" value="1.4" valueMax="1.8" valueMin="1.0">11.8 mm</quantity>
long, persistent, densely pubescent. Inflorescences cymose, sessile or subsessile, 13-flowered; bracts small,
<quantity box="[283,396,1382,1409]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="24" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
long, densely pubescent. Flowers fascicular or solitary, pedicel
<quantity box="[1103,1167,1382,1409]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="24" unit="mm" value="1.0">1 mm</quantity>
long; calyx campanulate,
<quantity box="[136,229,1418,1445]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="24" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
long, bluish purple, pubescent outside, calyx tube
<quantity box="[794,908,1418,1445]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="24" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
long, calyx lobes 5, narrowly triangular,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="24" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.52.5 mm</quantity>
long, tips acute or acuminate; corolla tube cylindrical,
<quantity box="[817,913,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="24" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">58 mm</quantity>
long, white purple, corolla tube
<quantity box="[1291,1386,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="24" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
long, pubescent outside, corolla lobes 5, triangular-lanceolate,
<quantity box="[770,862,1490,1517]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, long-villose outside, floccose inside, the throat hairy; stamens 5, filaments short, anthers narrowly oblong,
<quantity box="[813,949,1526,1553]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="24" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
long; ovary with 5 locules, style linear,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="24" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
long. Fruits drupaceous, hemispheric,
<quantity box="[629,725,1562,1589]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="24" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long,
<quantity box="[801,897,1562,1589]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="24" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
in diam., pubescent, crowned by 5 calyx lobes, orange when ripe; pyrenes 5. Seeds ovoid,
<quantity box="[617,710,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, irregular sulcate on the abaxial face, brown when dry.
</paragraph>
<paragraph blockId="24.[136,1452,986,2093]" pageId="24">
<emphasis bold="true" box="[189,525,1634,1660]" pageId="24">Other specimens examined:</emphasis>
<materialsCitation collectingDate="2019-07-26" collectionCode="KYO, VNM" collectorName="Dang V. S. &amp; Naiki A. &amp; Dang &amp; Japan" country="Vietnam" county="Iriomote Station" elevation="1190" location="Thua Thien Hue Province" municipality="Bach Ma National Park" pageId="24" specimenCount="1">
<collectingCountry box="[554,686,1634,1660]" name="Vietnam" pageId="24">VIETNAM</collectingCountry>
.
<location LSID="urn:lsid:plazi:treatment:43284262FF87FFC75AB6FC834386CD24:CE5EA5AFFF87FFC65884F9C743C3CA34" box="[698,997,1634,1661]" country="Vietnam" county="Iriomote Station" municipality="Bach Ma National Park" name="Thua Thien Hue Province" pageId="24">Thua Thien Hue Province</location>
,
<collectingMunicipality box="[1010,1284,1634,1661]" pageId="24">Bach Ma National Park</collectingMunicipality>
,
<quantity box="[1297,1383,1634,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="24" unit="m" value="1190.0">
<elevation box="[1297,1383,1634,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="24" unit="m" value="1190.0">1190 m</elevation>
</quantity>
elev., 16°1176.13N, 107°5084.75E, in evergreen forest,
<date box="[745,891,1670,1696]" pageId="24" value="2019-07-26">
<collectingDate box="[745,891,1670,1696]" pageId="24" value="2019-07-26">26 July 2019</collectingDate>
</date>
,
<emphasis box="[901,1272,1670,1697]" italics="true" pageId="24">
<collectorName box="[901,1011,1671,1697]" pageId="24">Dang V.S.</collectorName>
&amp;
<collectorName box="[1044,1137,1670,1697]" pageId="24">Naiki A.</collectorName>
,
<collectorName box="[1148,1210,1671,1696]" pageId="24">Dang</collectorName>
406a
</emphasis>
(
<collectionCode box="[1286,1356,1670,1696]" country="Japan" lsid="urn:lsid:biocol.org:col:15604" name="Kyoto University" pageId="24" type="Herbarium">KYO</collectionCode>
!,
<collectionCode box="[1370,1444,1670,1696]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="24" type="Herbarium">VNM</collectionCode>
!, the herbarium of
<collectingCounty box="[330,512,1706,1732]" pageId="24">Iriomote Station</collectingCounty>
(
<collectorName box="[527,596,1706,1732]" pageId="24">Japan</collectorName>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="24" type="distribution">
<paragraph blockId="24.[136,1452,986,2093]" box="[189,1310,1742,1769]" pageId="24">
<emphasis bold="true" box="[189,345,1742,1768]" pageId="24">Distribution:</emphasis>
<collectingCountry box="[373,468,1742,1768]" name="Vietnam" pageId="24">Vietnam</collectingCountry>
(known only from the Bach Ma National Park, Thua Thien Hue Province).
</paragraph>
<paragraph blockId="24.[136,1452,986,2093]" pageId="24">
<emphasis bold="true" box="[189,420,1778,1804]" pageId="24">Habitat &amp; Ecology:</emphasis>
<materialsCitation country="Vietnam" elevation="1100" location="Bach Ma National Park" municipality="Thua Thien Hue Province" pageId="24" specimenCount="1">
This new species is currently known only from the
<typeStatus box="[1013,1061,1779,1805]" pageId="24">type</typeStatus>
locality in
<location LSID="urn:lsid:plazi:treatment:43284262FF87FFC75AB6FC834386CD24:CE5EA5AFFF87FFC65E9EF9574581CB44" box="[1184,1447,1778,1805]" country="Vietnam" municipality="Thua Thien Hue Province" name="Bach Ma National Park" pageId="24">Bach Ma National Park</location>
,
<collectingMunicipality box="[136,427,1814,1841]" pageId="24">Thua Thien Hue Province</collectingMunicipality>
, central
<collectingCountry box="[521,616,1814,1840]" name="Vietnam" pageId="24">Vietnam</collectingCountry>
where it grows well under the shade of the evergreen forest at an elevation between
<quantity box="[236,394,1850,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="24" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">
<elevation box="[236,394,1850,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="24" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">10001200 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph blockId="24.[136,1452,986,2093]" box="[189,1169,1886,1913]" pageId="24">
<emphasis bold="true" box="[189,323,1886,1912]" pageId="24">Phenology:</emphasis>
—Flowering from March to August, fruiting from September to November.
</paragraph>
</subSubSection>
<subSubSection lastPageId="25" pageId="24" type="etymology">
<paragraph blockId="24.[136,1452,986,2093]" pageId="24">
<emphasis bold="true" box="[189,328,1922,1948]" pageId="24">Etymology:</emphasis>
—The specific epithet refers to the
<typeStatus box="[743,791,1923,1949]" pageId="24">type</typeStatus>
locality in the Bach Ma National Park, Thua Thien Hue Province, Central
<collectingCountry box="[339,438,1958,1984]" name="Vietnam" pageId="24">Vietnam</collectingCountry>
.
</paragraph>
<paragraph blockId="24.[136,1452,986,2093]" box="[189,655,1994,2023]" pageId="24">
<emphasis bold="true" box="[189,405,1994,2020]" pageId="24">Vernacular name:</emphasis>
—Xú hưƠng bạch mã.
</paragraph>
<paragraph blockId="24.[136,1452,986,2093]" lastBlockId="25.[136,1452,158,365]" lastPageId="25" pageId="24">
<emphasis bold="true" box="[189,647,2030,2056]" pageId="24">Preliminary conservation assessment:</emphasis>
<taxonomicName box="[675,953,2030,2056]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="24" phylum="Tracheophyta" rank="species" species="bachmaensis">
<emphasis box="[675,953,2030,2056]" italics="true" pageId="24">Lasianthus bachmaensis</emphasis>
</taxonomicName>
is only known in Bach Ma National Park, Thua Thien Hue Province. The Extent of Occurrence (EOO) is less than
<quantity box="[958,1044,2066,2092]" metricMagnitude="5" metricUnit="m" metricValue="1.7" pageId="24" unit="km" value="170.0">170 km</quantity>
<superScript attach="left" box="[1044,1052,2066,2080]" fontSize="6" pageId="24">2</superScript>
and the known Area of Occupancy (AOO) is less than
<quantity box="[367,441,158,185]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="25" unit="km" value="25.0">25 km</quantity>
<superScript attach="left" box="[441,449,158,172]" fontSize="6" pageId="25">2</superScript>
. About 1000 individuals were observed along the roadside, streams, and edges of the evergreen forest which are easily affected by human tourism activities. Thus, we propose a status of Endangered (EN) according to the IUCN Red List Categories (IUCN 2019).
</paragraph>
<paragraph blockId="25.[136,1452,158,365]" pageId="25">
<emphasis bold="true" box="[189,265,266,292]" pageId="25">Notes:</emphasis>
—This new species resembles
<taxonomicName authority="Naiki, Tagane &amp; Yahara (2015: 173)" authorityName="Naiki, Tagane &amp; Yahara" authorityPageNumber="173" authorityYear="2015" box="[607,1352,266,293]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="25" phylum="Tracheophyta" rank="species" species="stephanocalycinus">
<emphasis box="[607,943,266,292]" italics="true" pageId="25">Lasianthus stephanocalycinus</emphasis>
<bibRefCitation author="Naiki, A. &amp; Tagane, S. &amp; Chhang, P. &amp; Toyama, H. &amp; Zhu, H. &amp; Dang, V. S. &amp; Yahara, T." box="[949,1352,266,293]" pageId="25" pagination="153 - 179" refId="ref54656" refString="Naiki, A., Tagane, S., Chhang, P., Toyama, H., Zhu, H., Dang, V. S. &amp; Yahara, T. (2015) Flora of Bokor National Park, Cambodia II: four new species and nine new records of Lasianthus (Rubiaceae) from Cambodia. Acta Phytotaxonomica et Geobotanica 66 (3): 153 - 179. https: // doi. org / 10.18942 / apg. KJ 00010115702" type="journal article" year="2015">Naiki, Tagane &amp; Yahara (2015: 173)</bibRefCitation>
</taxonomicName>
which is known from Bokor National Park of
<collectingCountry box="[552,667,302,328]" name="Cambodia" pageId="25">Cambodia</collectingCountry>
and Hon Ba Nature Reserve of
<collectingCountry box="[1027,1125,302,328]" name="Vietnam" pageId="25">Vietnam</collectingCountry>
, but differs from that species based on several morphological characteristics summarized in
<tableCitation box="[838,921,338,364]" captionStart="TABLE 1" captionStartId="26.[136,229,158,184]" captionText="TABLE 1. Comparison of Lasianthus bachmaensis with its morphologically similar species." pageId="25">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>