418 lines
55 KiB
XML
418 lines
55 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1168.100798" ID-Pensoft-Pub="1313-2970-1168-295" ID-Pensoft-UUID="F19054F21F3D516FA0292C896F566182" ID-ZooBank="745FFCD5E3E84DC1B76CC82AD1AC5C64" ModsDocID="1313-2970-1168-295" checkinTime="1688439520051" checkinUser="pensoft" docAuthor="Hou, Yanmeng, Zhao, Lingchen & Zhang, Feng" docDate="2023" docId="11FBD2656E4F5192BB03F0DCB46C4619" docLanguage="en" docName="ZooKeys 1168: 295-327" docOrigin="ZooKeys 1168" docPubDate="2023-07-03" docSource="http://dx.doi.org/10.3897/zookeys.1168.100798" docTitle="Catatemnus tengchongensis Hou & Zhao & Zhang 2023, sp. nov." docType="treatment" docUuid="344BB985-9DD6-4CBC-96A8-86B315B3F1AE" docUuidSource="ZooBank" docVersion="1" id="F19054F21F3D516FA0292C896F566182" lastPageNumber="295" masterDocId="F19054F21F3D516FA0292C896F566182" masterDocTitle="First record of the genus Catatemnus Beier, 1932 from China, with the description of six new species (Pseudoscorpiones, Atemnidae)" masterLastPageNumber="327" masterPageNumber="295" pageNumber="295" updateTime="1688439520051" updateUser="pensoft">
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<mods:titleInfo id="03B507D7C4F3E198C179B0CA33488525">
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<mods:title id="C8A2D589664F04A1BD7696BB4DFABDFC">First record of the genus Catatemnus Beier, 1932 from China, with the description of six new species (Pseudoscorpiones, Atemnidae)</mods:title>
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<mods:name id="20E20630AC9EAC618D2839D2BDF451A5" type="personal">
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<mods:namePart id="ACBB5B63AEC812EC68E6F7C55A99757C">Hou, Yanmeng</mods:namePart>
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<mods:affiliation id="4A94F4B773DA538425E86CCF66D2C268">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="D5E1D05DFC01913A4F5CC05A95386E4E">Zhao, Lingchen</mods:namePart>
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<mods:nameIdentifier id="A037A91D68E9E19C41EBE515FAC868B2" type="ORCID">https://orcid.org/0000-0001-8477-836X</mods:nameIdentifier>
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<mods:affiliation id="453E8900854C8144312D4725483540A1">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China & Changzhi Tunliu District No. 6 Middle School, Changzhi, Shanxi 046100, China</mods:affiliation>
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<mods:namePart id="FE232ED34B07336342C4F1D481AAEBFF">Zhang, Feng</mods:namePart>
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<mods:nameIdentifier id="30952AF93A0AA3A6716D03EC883EEFB7" type="ORCID">https://orcid.org/0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation id="9D7B53FEFE6F96905469FAD6DF58FAF1">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:nameIdentifier id="BDC8C89AACD85ADBFF4BEEB6B29A540D" type="email">dudu06042001@163.com</mods:nameIdentifier>
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<mods:title id="8766F7E104722C32F886E9AC38AEDAEF">ZooKeys</mods:title>
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<mods:date id="6402B077EBE7206AEC36F533F6F8E1D8">2023</mods:date>
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<mods:number id="25D7517363F91F715F6F3EDA0214F3AE">2023-07-03</mods:number>
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<mods:number id="41B21576F17B3470D3152236DF7BBBDF">1168</mods:number>
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<mods:identifier id="BE0033E69C56BDD49C933E178EB64E5C" type="DOI">http://dx.doi.org/10.3897/zookeys.1168.100798</mods:identifier>
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<treatment id="11FBD2656E4F5192BB03F0DCB46C4619" LSID="urn:lsid:zoobank.org:act:344BB985-9DD6-4CBC-96A8-86B315B3F1AE" httpUri="http://treatment.plazi.org/id/11FBD2656E4F5192BB03F0DCB46C4619" lastPageNumber="295" pageId="0" pageNumber="295">
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<subSubSection id="9E3472AFF1690EC299EF01774028774A" pageId="0" pageNumber="295" type="nomenclature">
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<paragraph id="4DBA05D3382A6F9AA37DF41B499F9CAC" pageId="0" pageNumber="295">
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<taxonomicName id="3D93ED07142D3AA53E421D3544812913" LSID="https://zoobank.org/344BB985-9DD6-4CBC-96A8-86B315B3F1AE" authority="Hou & Zhao & Zhang, 2023" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus tengchongensis" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="tengchongensis" status="sp. nov.">Catatemnus tengchongensis</taxonomicName>
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<taxonomicNameLabel id="00EB3BEDF40FA30A393C2CFFF1A8C4D1" pageId="0" pageNumber="295">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="A4AB5952E22F93DA6082D2090EAA4157" pageId="0" pageNumber="295" type="description">
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<paragraph id="1F0778A6ED93BF11FCD423839397C0C6" pageId="0" pageNumber="295">
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<figureCitation id="89A6D25844E5B689287F6C3371AD31AC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Catatemnus tengchongensis sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 2.00 mm (B); 1.00 mm (A)." figureDoi="10.3897/zookeys.1168.100798.figure13" httpUri="https://binary.pensoft.net/fig/872312" pageId="0" pageNumber="295">Figs 13</figureCitation>
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<figureCitation id="99EB5C3F8A63CF4D748440E0EF0E7976" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Catatemnus tengchongensis sp. nov., holotype male (A-D, F-I), paratype female (E) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E female genital area (ventral view) F male genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure14" httpUri="https://binary.pensoft.net/fig/872313" pageId="0" pageNumber="295">, 14</figureCitation>
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<figureCitation id="944210427C7BE89F3027C6706508C27F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">, 15 (腾冲猫伪蝎)</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="12AE753C7CB8FA5A2E2FEFEDFC3F41DE" pageId="0" pageNumber="295" type="type material">
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<paragraph id="86BE7B4845300B2183292AA2F9B5982C" pageId="0" pageNumber="295">Type material.</paragraph>
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<paragraph id="2383CC0FBA0734DCDC8493082C52A933" pageId="0" pageNumber="295">
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<emphasis id="30F6A1D915EE766ADD3E69AF62569CDB" bold="true" italics="true" pageId="0" pageNumber="295">Holotype</emphasis>
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: China • ♂; Yunnan Province, Tengchong City, Houqiao Town, Yangjiatian Village;
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<geoCoordinate id="54294765508EB2CE2E1934410DC47E29" degrees="25" direction="north" minutes="21.413" orientation="latitude" precision="1" value="25.356884">25°21.413'N</geoCoordinate>
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,
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<geoCoordinate id="5B66E4DEE90541FCC8C386939FCF9647" degrees="98" direction="east" minutes="15.308" orientation="longitude" precision="1" value="98.255135">98°15.308'E</geoCoordinate>
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; 1824 m a.s.l.; 07 Jun. 2017; Chi Jin, Beibei Zhou and Xiao Zang leg.; Ps.-MHBU-YNTC171201.
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<emphasis id="8E5DF294A5344AC9F5109F29D9F93C81" bold="true" italics="true" pageId="0" pageNumber="295">Paratypes</emphasis>
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: • 3♀; same data as the holotype; Ps.-MSWU-YNTC171202-04.
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</paragraph>
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</subSubSection>
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<subSubSection id="CFDEF79810C9CC620A2D778A065869C0" pageId="0" pageNumber="295" type="diagnosis">
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<paragraph id="B4BBEA795A32834883BD65F4817036CF" pageId="0" pageNumber="295">Diagnosis (♂♀).</paragraph>
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<paragraph id="DE076098284A97CFB62FD19878D220F6" pageId="0" pageNumber="295">
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This new species is characterized by: the serrula exterior with 25 lamellae; movable chelal finger with 28 teeth; palpal femur (♂) 2.59
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<normalizedToken id="3FAC40EA83FCD6B406365C4B315EF273" originalValue="×">x</normalizedToken>
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, (♀) 2.42-2.52
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<normalizedToken id="6455F4C6895F6640CBE2A96E1CD12AC6" originalValue="×">x</normalizedToken>
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, chela with pedicel (♂) 2.72
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<normalizedToken id="72C4E389ACA6BB24ADDE9444ACADF0BF" originalValue="×">x</normalizedToken>
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, (♀) 2.57-2.67
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<normalizedToken id="F3402872A82EB0FDCED6630902FA7F8D" originalValue="×">x</normalizedToken>
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, chela without pedicel (♂) 2.51
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<normalizedToken id="B328517472FD45B4483BBA9043C851D3" originalValue="×">x</normalizedToken>
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, (♀) 2.37-2.42
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<normalizedToken id="C930CA04148CB321C69408CB8473F1B2" originalValue="×">x</normalizedToken>
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longer than broad; distance between est and esb farther to that of ist and isb.
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</paragraph>
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</subSubSection>
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<subSubSection id="F75B50E53F79A2576D71E4AA3FEADD4E" pageId="0" pageNumber="295" type="description">
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<paragraph id="CEA417A1BB21B7686C4EBF97BD0F6838" pageId="0" pageNumber="295">Description.</paragraph>
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<paragraph id="26ACA41E12FD940EA2AECD5E3E780C8B" pageId="0" pageNumber="295">
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<emphasis id="CEFB7229A8CF29125DC13DB943DBE170" bold="true" pageId="0" pageNumber="295">Adult male</emphasis>
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(Figs
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<figureCitation id="A01C683783A42E10CE0804B6B40D9FBB" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Catatemnus tengchongensis sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 2.00 mm (B); 1.00 mm (A)." figureDoi="10.3897/zookeys.1168.100798.figure13" httpUri="https://binary.pensoft.net/fig/872312" pageId="0" pageNumber="295">13A</figureCitation>
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,
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<figureCitation id="276A0A898426AEE7EFA14AB0C05B4CA4" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Catatemnus tengchongensis sp. nov., holotype male (A-D, F-I), paratype female (E) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E female genital area (ventral view) F male genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure14" httpUri="https://binary.pensoft.net/fig/872313" pageId="0" pageNumber="295">14A-D, F-I</figureCitation>
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,
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<figureCitation id="41E072E58BC5B6F95963EAD789633D35" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15A, B, D-H, J</figureCitation>
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).
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<emphasis id="A18E23A4C6B35733B79679A873A462C9" bold="true" italics="true" pageId="0" pageNumber="295">Color</emphasis>
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: carapace dark brown, slightly paler in tergites and sternites; pedipalps reddish brown, but paler in trochanter; legs pale yellow.
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<emphasis id="F9E08BC7E54438BF384342BFC7AE5633" bold="true" italics="true" pageId="0" pageNumber="295">Carapace</emphasis>
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(Figs
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<figureCitation id="45E633BCD6932073D67C96E7DAB58593" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Catatemnus tengchongensis sp. nov., holotype male (A-D, F-I), paratype female (E) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E female genital area (ventral view) F male genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure14" httpUri="https://binary.pensoft.net/fig/872313" pageId="0" pageNumber="295">14A</figureCitation>
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,
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<figureCitation id="EE450A2C8B2BD56E08E8181554335386" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15A</figureCitation>
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): 1.20
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<normalizedToken id="A327201CC6C19B90C3EB5137A2A4EA3C" originalValue="×">x</normalizedToken>
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longer than broad; surface smooth, but with a medial furrow; with two distinct eyespots situated near anterior margin of carapace; anterior margin with five setae, posterior margin with eight setae, each seta acicular and very slightly curved.
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<emphasis id="77DEEF0EDD152D54C0397C46DBFC2E8A" bold="true" italics="true" pageId="0" pageNumber="295">Chelicera</emphasis>
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(Fig.
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<figureCitation id="68EC1287A4F497C5BE54B8035AEC2CD3" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15B, D</figureCitation>
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): much smaller than carapace length; surface smooth; four setae (sbs absent; es shorter than others) and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand; movable finger with one galeal seta (shorter than others); bs and es dentate apically, is and ls long and acute. Fixed finger with seven unequally sized teeth, movable finger with two large teeth; galea present, shorter with two small branchlets and two dentations (Fig.
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<figureCitation id="01DDC81D5CE8D5157A3E987D45EF7FE5" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15B</figureCitation>
|
||
). Serrula interior connected to fixed finger for entire length, proximally modified to form velum, serrula exterior with 25 lamellae, the basal one longest; lamina exterior present. Rallum composed of four blades, the basal two blades shorter than others, the distal two dentated anteriorly, remainder smooth (Fig.
|
||
<figureCitation id="12D5E5D9071F212F2A03CA89E9A312FD" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15D</figureCitation>
|
||
).
|
||
<emphasis id="BE712865DE1D161F4FDB4D9437CE16AD" bold="true" italics="true" pageId="0" pageNumber="295">Pedipalp</emphasis>
|
||
(Figs
|
||
<figureCitation id="750AD72666A2ED59969994AC21376469" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Catatemnus tengchongensis sp. nov., holotype male (A-D, F-I), paratype female (E) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E female genital area (ventral view) F male genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure14" httpUri="https://binary.pensoft.net/fig/872313" pageId="0" pageNumber="295">14B-D</figureCitation>
|
||
,
|
||
<figureCitation id="4A46D1F4BC9AA394442CB0B7EB4C22C9" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15G, J</figureCitation>
|
||
): stout, trochanter 1.77
|
||
<normalizedToken id="5E80D1AA17B9A97842A164100AAAA638" originalValue="×">x</normalizedToken>
|
||
, femur 2.59
|
||
<normalizedToken id="E0FC1000AB11F407234BFC8D0717CD8D" originalValue="×">x</normalizedToken>
|
||
, patella 1.95
|
||
<normalizedToken id="A3434E07B0B4535A3055DD74CDB0DFAD" originalValue="×">x</normalizedToken>
|
||
, chela with pedicel (without pedicel) 2.72
|
||
<normalizedToken id="E3407D4725759079C291C6BE202E5ED4" originalValue="×">x</normalizedToken>
|
||
(2.51
|
||
<normalizedToken id="0091950377F3066F71F80A39015ADD09" originalValue="×">x</normalizedToken>
|
||
), hand with pedicel (without pedicel) 1.66
|
||
<normalizedToken id="19605E043A87DB259E2E9844B16CA11C" originalValue="×">x</normalizedToken>
|
||
(1.45
|
||
<normalizedToken id="9A6984CA21426FC1BAA6991EDED9671C" originalValue="×">x</normalizedToken>
|
||
) longer than broad; movable chelal finger 0.64
|
||
<normalizedToken id="591CA337A93E09C3326F0A61BAE3F506" originalValue="×">x</normalizedToken>
|
||
(0.74
|
||
<normalizedToken id="FB084A056CBEFF1F67F763A96BA0D994" originalValue="×">x</normalizedToken>
|
||
) longer than hand with pedicel (without pedicel) and 0.39
|
||
<normalizedToken id="E204117A55E4E0F7CD6AF87D273DB07E" originalValue="×">x</normalizedToken>
|
||
(0.42
|
||
<normalizedToken id="90CC604497FC39DC5267A8E91EC2B493" originalValue="×">x</normalizedToken>
|
||
) longer than chela with pedicel (without pedicel). Setae generally long and acuminate. All segments (except chelal fingers) with fine granulations; trochanter with an inconspicuous ventral tubercle and a well-developed conical dorsal tubercle. Fixed chelal finger with eight trichobothria, movable chelal finger with four trichobothria: eb and esb situated at base of fixed finger on retrolateral face, esb slightly distal to eb; ib and isb situated at base of fixed finger on prolateral face, isb slightly distal to ib; est in the middle of fixed finger; et near sub-distal of fixed finger; est midway between et and esb; it slightly distal to est and proximal to et; ist situated proximal to est and distal to isb; it closer to ist than to fingertip; distance between est and esb farther to that of ist and isb; distance between it and fingertip nearly equal to distance between ist and isb; b and sb situated at base of movable finger on retrolateral face; t in the middle of movable finger and at same level as est; sb closer to b than to st; st closer to sb than to t (Fig.
|
||
<figureCitation id="FCFEB33D0648C75C1A72F57ABF66A55A" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15G</figureCitation>
|
||
). Venom apparatus only present in fixed chelal finger, venom ducts curved and short, terminating in inflated nodus ramosus between et and est. Both chelal fingers with a row of acute teeth, spaced contiguously along the margin, slightly rounded proximally: fixed chelal finger with 24 teeth; movable chelal finger with 28 teeth (nearly as large as teeth on fixed chelal finger); without accessory teeth (Fig.
|
||
<figureCitation id="030AA8B399887193411AB2452A3F8C62" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15G</figureCitation>
|
||
). Femur without long tactile setae. Movable chelal finger slightly curved in lateral view (Figs
|
||
<figureCitation id="6BD9043D2BEDDE33511232E76DC987C5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Catatemnus tengchongensis sp. nov., holotype male (A-D, F-I), paratype female (E) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E female genital area (ventral view) F male genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure14" httpUri="https://binary.pensoft.net/fig/872313" pageId="0" pageNumber="295">14B</figureCitation>
|
||
,
|
||
<figureCitation id="9B2C2505BE2DD21D3DDFF0182DCB524E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15G</figureCitation>
|
||
).
|
||
<emphasis id="644BDE690E0DED5245A6D438043234A9" bold="true" italics="true" pageId="0" pageNumber="295">Opisthosoma</emphasis>
|
||
: typical, all setae long and acuminate; pleural membrane longitudinally striate, without setae. Tergites incompletely divided, tergal chaetotaxy I-XI: 10: 8: 5: 11: 13: 14: 15: 15: 17: 17(13 + 4T): 12(10 + 2T). Sternites X and XI undivided and other sternites divided, sternal chaetotaxy IV-XI: 5-5: 8-8: 8-8: 7-8: 7-7: 7-7: 18: 16; sternites X and XI each with two pairs of long tactile setae. Anus (tergite XII and sternite XII) without raised rim. Anterior genital operculum with five or seven setae on each side, posterior margin with eight setae.
|
||
<emphasis id="2E10B560AE08156B59F29434D5E03185" bold="true" italics="true" pageId="0" pageNumber="295">Legs</emphasis>
|
||
(Figs
|
||
<figureCitation id="DFE2327B67008D04B5AA1B38D279883D" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Catatemnus tengchongensis sp. nov., holotype male (A-D, F-I), paratype female (E) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E female genital area (ventral view) F male genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure14" httpUri="https://binary.pensoft.net/fig/872313" pageId="0" pageNumber="295">14G-I</figureCitation>
|
||
,
|
||
<figureCitation id="E12830FC59FCC33DC1F3AC20F6E7A528" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15E, F</figureCitation>
|
||
): generally typical, fairly smooth, slightly stout; junction between femora and patellae I and II oblique. Femoropatella of leg IV 2.80
|
||
<normalizedToken id="7FFBE66695A775946C2A7583409D3D74" originalValue="×">x</normalizedToken>
|
||
longer than deep; tibia 3.86
|
||
<normalizedToken id="A79DC19BA05371DECD84B6A5C5A603DB" originalValue="×">x</normalizedToken>
|
||
longer than deep; with basal tactile setae on tarsal segment: tarsus 3.55
|
||
<normalizedToken id="95C3DFAACBDA08B67C1C0A2E6D72ACB7" originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.10); subterminal tarsal setae arcuate and acute. Arolium slightly shorter than claws, not divided; claws smooth.
|
||
</paragraph>
|
||
<caption id="F473AE2567073A4F021115F589C44214" doi="10.3897/zookeys.1168.100798.figure13" httpUri="https://binary.pensoft.net/fig/872312" pageId="0" pageNumber="295" start="Figure 13" startId="F13">
|
||
<paragraph id="2467A3E33960FAF77D0CA93F09D8E205" pageId="0" pageNumber="295">
|
||
<emphasis id="6332995B440697B027C394E213D3826A" bold="true" pageId="0" pageNumber="295">Figure 13.</emphasis>
|
||
<taxonomicName id="56B583B91C1172529109C4461CCF4F6C" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus tengchongensis" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="tengchongensis">
|
||
<emphasis id="B5AE2DB0944B490FE4F16E22308C127F" italics="true" pageId="0" pageNumber="295">Catatemnus tengchongensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="251756D3D329453480816C988C1EB87C" bold="true" pageId="0" pageNumber="295">A</emphasis>
|
||
holotype male, habitus (dorsal view)
|
||
<emphasis id="EB1F17BCC56A10D877BB6803DA4367A4" bold="true" pageId="0" pageNumber="295">B</emphasis>
|
||
paratype female, habitus (dorsal view). Scale bars: 2.00 mm (
|
||
<emphasis id="7E59AD0605A5392C58FC9D4B0A31E1B2" bold="true" pageId="0" pageNumber="295">B</emphasis>
|
||
); 1.00 mm (
|
||
<emphasis id="4F6E2DACE7718C1E0472063F187FEF16" bold="true" pageId="0" pageNumber="295">A</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="265332FACE6A891BAF73D39494BDD759" doi="10.3897/zookeys.1168.100798.figure14" httpUri="https://binary.pensoft.net/fig/872313" pageId="0" pageNumber="295" start="Figure 14" startId="F14">
|
||
<paragraph id="44085F4179D0666BDDF4240361966B9B" pageId="0" pageNumber="295">
|
||
<emphasis id="00D05C52CB3628542777104B2B24DD2F" bold="true" pageId="0" pageNumber="295">Figure 14.</emphasis>
|
||
<taxonomicName id="EF7DF787C177DCF96C3FE34347516C9B" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus tengchongensis" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="tengchongensis">
|
||
<emphasis id="6FFC72C8DA392920E23C646CF0C743E8" italics="true" pageId="0" pageNumber="295">Catatemnus tengchongensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis id="29F076A1ACDC784C0D25F40A154FBBED" bold="true" pageId="0" pageNumber="295">A-D, F-I</emphasis>
|
||
), paratype female (
|
||
<emphasis id="42B171F08D9406C26309663FA8797C3A" bold="true" pageId="0" pageNumber="295">E</emphasis>
|
||
)
|
||
<emphasis id="C4841F10C5F45A6878871F2D14C0F679" bold="true" pageId="0" pageNumber="295">A</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis id="CAB450AF06D490F4FC53025672679C91" bold="true" pageId="0" pageNumber="295">B</emphasis>
|
||
left chelal fingers (lateral view)
|
||
<emphasis id="A8A8DCB561E6DDC16907E6A5BF437F02" bold="true" pageId="0" pageNumber="295">C</emphasis>
|
||
left chela (lateral view)
|
||
<emphasis id="EE15338B891D306156C00D8E346153E8" bold="true" pageId="0" pageNumber="295">D</emphasis>
|
||
left pedipalp (dorsal view)
|
||
<emphasis id="592CE89AC19256E31ED06CA90425BDBC" bold="true" pageId="0" pageNumber="295">E</emphasis>
|
||
female genital area (ventral view)
|
||
<emphasis id="0F01A561AA5F3F2B3AD34DBAE77710C9" bold="true" pageId="0" pageNumber="295">F</emphasis>
|
||
male genital area (ventral view)
|
||
<emphasis id="86981F9AC63B4BF1E0BB9C4552E6FB8E" bold="true" pageId="0" pageNumber="295">G</emphasis>
|
||
tarsus of left leg IV (lateral view)
|
||
<emphasis id="EF4472360CCCEB473DA83322A6E70389" bold="true" pageId="0" pageNumber="295">H</emphasis>
|
||
left leg I (lateral view)
|
||
<emphasis id="7F5AA852A706BDB5599C9FB93088BD22" bold="true" pageId="0" pageNumber="295">I</emphasis>
|
||
left leg IV (lateral view). Scale bars: 0.50 mm (
|
||
<emphasis id="045660546001A6561E531BC25A3F02B7" bold="true" pageId="0" pageNumber="295">A, C, D</emphasis>
|
||
); 0.20 mm (
|
||
<emphasis id="41F62CC2421C0D7E33E3F1D1FEBE33FD" bold="true" pageId="0" pageNumber="295">B, E-I</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BA39FC9A3A6F5CD70C74AF2E360A72B5" doi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295" start="Figure 15" startId="F15">
|
||
<paragraph id="29EFBFAB4E5F78D065F01BF569DDB429" pageId="0" pageNumber="295">
|
||
<emphasis id="5C1BC45FA782B581838C919ADEDD4793" bold="true" pageId="0" pageNumber="295">Figure 15.</emphasis>
|
||
<taxonomicName id="61D3E18858DCCA5FDCC106AC8F634FBA" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus tengchongensis" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="tengchongensis">
|
||
<emphasis id="F270AD4B5328841015C50C8093B15135" italics="true" pageId="0" pageNumber="295">Catatemnus tengchongensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis id="BFDE7BC62F5F5C341D13223D1624FA1A" bold="true" pageId="0" pageNumber="295">A, B, D-H, J</emphasis>
|
||
), paratype female (
|
||
<emphasis id="15AD76584DD1967BC652D749A54B8DCC" bold="true" pageId="0" pageNumber="295">C, I</emphasis>
|
||
)
|
||
<emphasis id="895D054443E23DFA7FC78DE2D5A83A8C" bold="true" pageId="0" pageNumber="295">A</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis id="630426A418F5276C1F1B96FE4853EB55" bold="true" pageId="0" pageNumber="295">B</emphasis>
|
||
male galea
|
||
<emphasis id="99F0C7011123B63304C5BF178825EFFB" bold="true" pageId="0" pageNumber="295">C</emphasis>
|
||
female galea
|
||
<emphasis id="BB24E79E6F9602E226FADFCFE180E545" bold="true" pageId="0" pageNumber="295">D</emphasis>
|
||
rallum
|
||
<emphasis id="4933813729F0AE0348A47BD612A48DD3" bold="true" pageId="0" pageNumber="295">E</emphasis>
|
||
left leg I (lateral view)
|
||
<emphasis id="F2B8BDA5BA688B711AA319FC1855F3BA" bold="true" pageId="0" pageNumber="295">F</emphasis>
|
||
left leg IV (lateral view)
|
||
<emphasis id="B0E561D3AC3CC3EBBDE707A36C887019" bold="true" pageId="0" pageNumber="295">G</emphasis>
|
||
left chelal fingers (lateral view), with details of trichobothrial pattern
|
||
<emphasis id="DBC628DE749263AA9D4145EF2760E322" bold="true" pageId="0" pageNumber="295">H</emphasis>
|
||
male genital area (ventral view)
|
||
<emphasis id="863D606AA8028E20DDF8A86C94C67C42" bold="true" pageId="0" pageNumber="295">I</emphasis>
|
||
female genital area (ventral view)
|
||
<emphasis id="FFE34D65A345B838B66D9701FB44EF7C" bold="true" pageId="0" pageNumber="295">J</emphasis>
|
||
left pedipalp (dorsal view). Scale bars: 0.50 mm (
|
||
<emphasis id="2F3BBFE14BD55BAED26C786E8306DA16" bold="true" pageId="0" pageNumber="295">F, J</emphasis>
|
||
); 0.25 mm (
|
||
<emphasis id="92F0C3ABA37A8F578CF485B166266626" bold="true" pageId="0" pageNumber="295">A, E, G-I</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis id="1D0B8AC98808D17B2F86AA03D59BF1AD" bold="true" pageId="0" pageNumber="295">B-D</emphasis>
|
||
). Abbreviations:
|
||
<emphasis id="C75686101ED01A96975C1E50487C9B40" bold="true" pageId="0" pageNumber="295">b</emphasis>
|
||
= basal;
|
||
<emphasis id="B45F1F6F198400159901A2643D681B78" bold="true" pageId="0" pageNumber="295">sb</emphasis>
|
||
= sub-basal;
|
||
<emphasis id="ECC7B0AC389347F53B32911FD0453FA1" bold="true" pageId="0" pageNumber="295">st</emphasis>
|
||
= sub-terminal;
|
||
<emphasis id="4CD620A12D3DCF566A18906CE8AAAF66" bold="true" pageId="0" pageNumber="295">t</emphasis>
|
||
= terminal;
|
||
<emphasis id="0283084C43E10F55141548E32FA1646A" bold="true" pageId="0" pageNumber="295">ib</emphasis>
|
||
= interior basal;
|
||
<emphasis id="106160250D4ADCDFB9DC64396D9F601A" bold="true" pageId="0" pageNumber="295">isb</emphasis>
|
||
= interior sub-basal;
|
||
<emphasis id="A6257DA693F639365B870CEC6BCFB91F" bold="true" pageId="0" pageNumber="295">ist</emphasis>
|
||
= interior sub-terminal;
|
||
<emphasis id="C3B297E888A62A6354C74CD7F6190F35" bold="true" pageId="0" pageNumber="295">it</emphasis>
|
||
= interior terminal;
|
||
<emphasis id="87EB21C237A19BD22A89F392258AD15E" bold="true" pageId="0" pageNumber="295">eb</emphasis>
|
||
= exterior basal;
|
||
<emphasis id="C0A806DB069138CA849220D2ACEE87B0" bold="true" pageId="0" pageNumber="295">esb</emphasis>
|
||
= exterior sub-basal;
|
||
<emphasis id="CD8F408CB0D10B868DF22A237BD47F27" bold="true" pageId="0" pageNumber="295">est</emphasis>
|
||
= exterior sub-terminal;
|
||
<emphasis id="ABC3468976966D22BF5556BCAE4FCB53" bold="true" pageId="0" pageNumber="295">et</emphasis>
|
||
= exterior terminal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="0C9396ACF9537B92A68FD7436A1A8EB7" pageId="0" pageNumber="295">
|
||
<emphasis id="AD9EEE618E35DC6B96E672B2FC06E038" bold="true" pageId="0" pageNumber="295">Adult females</emphasis>
|
||
(Figs
|
||
<figureCitation id="C524D78265A80C2B4E039CCC42C670F6" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Catatemnus tengchongensis sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 2.00 mm (B); 1.00 mm (A)." figureDoi="10.3897/zookeys.1168.100798.figure13" httpUri="https://binary.pensoft.net/fig/872312" pageId="0" pageNumber="295">13B</figureCitation>
|
||
,
|
||
<figureCitation id="396AD9AE0530DB36817D0DC536924B64" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Catatemnus tengchongensis sp. nov., holotype male (A-D, F-I), paratype female (E) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E female genital area (ventral view) F male genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure14" httpUri="https://binary.pensoft.net/fig/872313" pageId="0" pageNumber="295">14E</figureCitation>
|
||
,
|
||
<figureCitation id="8672F85E370D02E0F9717E0E272AA180" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15I</figureCitation>
|
||
): mostly same as the male, but larger and paler.
|
||
<emphasis id="A96A254CC180EE0D549DDC641C2CA579" bold="true" italics="true" pageId="0" pageNumber="295">Chelicera</emphasis>
|
||
: galea distinct relatively short, with five distinct larger branchlets (Fig.
|
||
<figureCitation id="3A7C3A0823E7F7C38B4158B60EADF672" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Catatemnus tengchongensis sp. nov., holotype male (A, B, D-H, J), paratype female (C, I) A carapace (dorsal view) B male galea C female galea D rallum E left leg I (lateral view) F left leg IV (lateral view) G left chelal fingers (lateral view), with details of trichobothrial pattern H male genital area (ventral view) I female genital area (ventral view) J left pedipalp (dorsal view). Scale bars: 0.50 mm (F, J); 0.25 mm (A, E, G-I); 0.10 mm (B-D). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure15" httpUri="https://binary.pensoft.net/fig/872314" pageId="0" pageNumber="295">15C</figureCitation>
|
||
); serrula exterior with 25 lamellae, the basal one longest.
|
||
<emphasis id="AFE608CDAB84566142801569A8633E70" bold="true" italics="true" pageId="0" pageNumber="295">Pedipalp</emphasis>
|
||
: stout, trochanter 1.67-1.78
|
||
<normalizedToken id="C6D37A76E7429DFE50E586084925288E" originalValue="×">x</normalizedToken>
|
||
, femur 2.42-2.52
|
||
<normalizedToken id="188178DB09CE5BAC19C7614BF61CE9DE" originalValue="×">x</normalizedToken>
|
||
, patella 2.00-2.25
|
||
<normalizedToken id="9CA3169D0A0750C082DB10B9340DD282" originalValue="×">x</normalizedToken>
|
||
, chela (with pedicel) 2.57-2.67
|
||
<normalizedToken id="C55474F2A25518BDA5C2C6EBD48AFD45" originalValue="×">x</normalizedToken>
|
||
, chela (without pedicel) 2.37-2.42
|
||
<normalizedToken id="E0169C72FD4F0A5F9DF0636C1C03362A" originalValue="×">x</normalizedToken>
|
||
, hand (with pedicel) 1.61-1.69
|
||
<normalizedToken id="19E6087FE1E82DBCC8F51F91CEA8091F" originalValue="×">x</normalizedToken>
|
||
, hand (without pedicel) 1.41-1.44
|
||
<normalizedToken id="D2770B5BBE0F63286A23B2E34DCECC10" originalValue="×">x</normalizedToken>
|
||
longer than broad, movable chelal finger 0.58-0.60
|
||
<normalizedToken id="D90039935C73356C41143C4B5CBEEE9C" originalValue="×">x</normalizedToken>
|
||
(0.68
|
||
<normalizedToken id="49529267287FCB3342ECF86A5E0526E3" originalValue="×">x</normalizedToken>
|
||
) longer than hand with pedicel (without pedicel) and 0.37
|
||
<normalizedToken id="242B4A5C81C04CC40E67B0C71CB1C823" originalValue="×">x</normalizedToken>
|
||
(0.40-0.41
|
||
<normalizedToken id="69F0F424C63EDC65E301DABE7F1DAD22" originalValue="×">x</normalizedToken>
|
||
) longer than chela with pedicel (without pedicel); palpal coxa with four apical setae, sub-terminal seta larger; all segments (except chelal fingers) with fine granulations.
|
||
<emphasis id="02057E723A9A095060717490F96973B8" bold="true" italics="true" pageId="0" pageNumber="295">Opisthosoma</emphasis>
|
||
: tergites incompletely divided, tergal chaetotaxy I-XI: 8: 9: 10: 13: 16: 14: 15: 17: 15: 16(12 + 4T): 11(9 + 2T). Sternites divided, sternal chaetotaxy IV-XI: 7-8: 8-8: 8-8: 8-9: 8-8: 9-9: 16(12 + 4T): 14(12 + 2T). Genitalia simple, spermathecae provided with separated median cribriform plates; anterior genital operculum with five setae on each side, posterior margin with four or five setae on each side.
|
||
<emphasis id="2C27AB1A84C577B808A59D522C28C623" bold="true" italics="true" pageId="0" pageNumber="295">Legs</emphasis>
|
||
: femoropatella of leg IV 2.81-2.93
|
||
<normalizedToken id="734C4378698CF0FBDBD19055D322A5A0" originalValue="×">x</normalizedToken>
|
||
longer than deep; tibia 3.28-3.41
|
||
<normalizedToken id="7F58CCA9A4907BCBEEE2E86CBDE0434A" originalValue="×">x</normalizedToken>
|
||
longer than deep; with basal tactile setae on tarsal segment: tarsus 3.75-4.00
|
||
<normalizedToken id="F34AF2F2083BC36DBA1FB6E23B7637E5" originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.09-0.11).
|
||
</paragraph>
|
||
<paragraph id="0D19AA1390C2E777894612B31E561433" pageId="0" pageNumber="295">
|
||
<emphasis id="2F93838A32229F73880C4D69A2F77578" bold="true" pageId="0" pageNumber="295">Dimensions</emphasis>
|
||
(length/breadth or, in the case of the legs, length/depth in mm). Male (females in parentheses): body length 2.73 (4.25-4.40). Carapace 0.85/0.71 (0.93-1.01/0.80-0.84). Pedipalp: trochanter 0.46/0.26 (0.45-0.48/0.27), femur 0.75/0.29 (0.75-0.78/0.31), patella 0.72/0.37 (0.72-0.81/0.36), chela (with pedicel) 1.28/0.47 (1.39/0.52-0.54), chela (without pedicel) 1.18 (1.26-1.28), hand (with pedicel) 0.78 (0.87-0.88), hand (without pedicel) 0.68 (0.75-0.76), movable finger length 0.50 (0.51-0.52). Leg I: trochanter 0.18/0.15 (0.17-0.20/0.14-0.17), femur 0.25/0.19 (0.24-0.26/0.20-0.21), patella 0.37/0.17 (0.41/0.18), tibia 0.39/0.12 (0.42-0.44/0.12), tarsus 0.38/0.09 (0.40-0.42/0.08-0.09). Leg IV: trochanter 0.29/0.16 (0.32-0.33/0.21), femoropatella 0.70/0.25 (0.76-0.79/0.27), tibia 0.54/0.15 (0.58-0.59/0.17-0.18), tarsus 0.39/0.11 (0.44-0.45/0.11-0.12).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CCB4E12153FDA71424F5F47ACD716656" pageId="0" pageNumber="295" type="remarks">
|
||
<paragraph id="5DAB5E3BFBFB8822C2047EDDC407C714" pageId="0" pageNumber="295">Remarks.</paragraph>
|
||
<paragraph id="1ED17C28EA851A72E1D230AB5869E1F6" pageId="0" pageNumber="295">
|
||
<taxonomicName id="77CB55FB063799D55E40416F06B3E01D" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus tengchongensis" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="tengchongensis">
|
||
<emphasis id="D53C645B50B60D85556D547BD3FF3C45" italics="true" pageId="0" pageNumber="295">Catatemnus tengchongensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName id="2FD8F58E6622061F35A438C44DBCF322" class="Arachnida" kingdom="Animalia" lsidName="C. birmanicus" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis id="76FAC5459B8CEFA4B6C592E5D8D8DAAA" italics="true" pageId="0" pageNumber="295">C. birmanicus</emphasis>
|
||
</taxonomicName>
|
||
by the smaller body size (♂) (e.g., body length 2.73 mm vs 4.27 mm; palpal femur 2.59
|
||
<normalizedToken id="93D6AED383CA474C772768A9854833AB" originalValue="×">x</normalizedToken>
|
||
vs 2.44
|
||
<normalizedToken id="14B4F5033D335056879582E727C3581F" originalValue="×">x</normalizedToken>
|
||
longer than broad, length 0.75 mm vs 1.09 mm; movable chelal finger length 0.50 mm vs 0.78 mm) and the trait of eyespots (with two distinct eyespots vs with two faint eyespots); from
|
||
<taxonomicName id="BBE61114A6D103756421C03B489CEDD9" class="Arachnida" kingdom="Animalia" lsidName="C. concavus" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="concavus">
|
||
<emphasis id="68C6FFFB0C4FA4B65D5839AE755C314D" italics="true" pageId="0" pageNumber="295">C. concavus</emphasis>
|
||
</taxonomicName>
|
||
by the trait of eyespots (with two distinct eyespots vs with two faint eyespots), the stouter pedipalps (♀) (e.g., palpal femur 2.42-2.52
|
||
<normalizedToken id="B7024DBBF0DF0CA9EB9247950AB6E0C1" originalValue="×">x</normalizedToken>
|
||
vs 2.29
|
||
<normalizedToken id="536C9B776BDA355F9CC52546B1A09864" originalValue="×">x</normalizedToken>
|
||
longer than broad, length 0.75-0.78 mm vs 0.90 mm) and the traits of the three first tergites (regularly convex from the one side to the other vs with a descending lateral portion, perpendicular on the dorsal part); from
|
||
<taxonomicName id="711290A67779C05689286EF41B7FE2BF" class="Arachnida" kingdom="Animalia" lsidName="C. monitor" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="monitor">
|
||
<emphasis id="083BF4733803CA40D99FBB7296B77517" italics="true" pageId="0" pageNumber="295">C. monitor</emphasis>
|
||
</taxonomicName>
|
||
by the trait of eyespots (with two distinct eyespots vs no eyespots visible) and the stouter pedipalps (e.g., palpal femur (♂) 2.59
|
||
<normalizedToken id="195027581A0C46D437E8E2F7839C0F2A" originalValue="×">x</normalizedToken>
|
||
vs 2.25
|
||
<normalizedToken id="8057C8D4903644FA97A7EC227CD24A4B" originalValue="×">x</normalizedToken>
|
||
, (♀) 2.42-2.52
|
||
<normalizedToken id="6621655D6FB1DE9C0387BC6E4579D432" originalValue="×">x</normalizedToken>
|
||
vs 2.27
|
||
<normalizedToken id="161214280039839F5DF52DAC4CEAA8A2" originalValue="×">x</normalizedToken>
|
||
longer than broad, length (♂) 0.75 mm vs 1.01 mm, (♀) 0.75-0.78 mm vs 0.95 mm); from
|
||
<taxonomicName id="9D00B1C7203D0CD58A326A8E4F960294" class="Arachnida" kingdom="Animalia" lsidName="C. nicobarensis" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="nicobarensis">
|
||
<emphasis id="52C0E833DD8FB5BA229B16500804A429" italics="true" pageId="0" pageNumber="295">C. nicobarensis</emphasis>
|
||
</taxonomicName>
|
||
by the trait of eyespots (with two distinct eyespots vs with two indistinct eyespots) and the traits of the palpal trochanter (anterior surface granular vs smooth or almost smooth); from
|
||
<taxonomicName id="1147D2EC27E43A302349215CD2F59B91" class="Arachnida" kingdom="Animalia" lsidName="C. thorelli" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="thorelli">
|
||
<emphasis id="7CBFD298DD39A2ED2E295AD31FF4FFDD" italics="true" pageId="0" pageNumber="295">C. thorelli</emphasis>
|
||
</taxonomicName>
|
||
by the trait of furrow on the carapace (distinctly curved vs almost straight) and the traits of pedipalps (surface of all segments except chelal fingers granular vs surface of palps smooth; palpal femur 2.59
|
||
<normalizedToken id="0557E9426244ABAB4CC95612314875E6" originalValue="×">x</normalizedToken>
|
||
vs 2.40
|
||
<normalizedToken id="61A9CA954AA31BCBAFCDB7D66EBB1133" originalValue="×">x</normalizedToken>
|
||
longer than broad, length 0.75 mm vs 1.15 mm; movable chelal finger teeth length 0.50 mm vs 0.70 mm) (
|
||
<bibRefCitation id="09D69622192C174A1A50EFDBE76C0BB7" DOI="https://doi.org/10.5962/bhl.title.10400" author="Thorell, T" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="295" pagination="521 - 729" refId="B9" refString="Thorell, T, 1889. Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885-1887. Annali del Museo Civico di Storia Naturale di Genova 7 (2): 521 - 729, DOI: https://doi.org/10.5962/bhl.title.10400" title="Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885 - 1887." url="https://doi.org/10.5962/bhl.title.10400" volume="7" year="1889">Thorell 1889</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4FE8CC0602A5CF7228A5618491773DA4" author="Balzan, L" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="295" pagination="497 - 512" refId="B1" refString="Balzan, L, 1891. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones). Annales de la Societe Entomologique de France 60: 497 - 512" title="Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones)." volume="60" year="1891">Balzan 1891</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="26F3B249D052987976AC4A7C295C9B36" author="With, CJ" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="295" refId="B10" refString="With, CJ, 1906. The Danish expedition to Siam 1899-1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Oversigt over det Konigelige Danske Videnskabernes Selskabs Forhandlinger (7)3: 1-214." title="The Danish expedition to Siam 1899 - 1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Oversigt over det Konigelige Danske Videnskabernes Selskabs Forhandlinger (7) 3: 1 - 214." year="1906">With 1906</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B709DDECF6C5B540C0798F7A01B1E8DC" DOI="https://doi.org/10.1515/9783111385402" author="Beier, M" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="295" refId="B2" refString="Beier, M, 1932. Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich 58: [i-xxi,] 1-294. https://doi.org/10.1515/9783111385402" title="Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich 58: [i-xxi,] 1 - 294." url="https://doi.org/10.1515/9783111385402" year="1932">Beier 1932</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E60AA6BEBEF8FA4867961388392A9167" pageId="0" pageNumber="295" type="distribution">
|
||
<paragraph id="D8BF541049E39305F9344D31269E83AD" pageId="0" pageNumber="295">Distribution.</paragraph>
|
||
<paragraph id="70D80303A4810EC7AE7C36AEE5A47611" pageId="0" pageNumber="295">Known only from the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A66D0414373B711CBED92981BD0AEF5F" pageId="0" pageNumber="295" type="etymology">
|
||
<paragraph id="FFF59D0E7A14E2B9C45019245A6529BF" pageId="0" pageNumber="295">Etymology.</paragraph>
|
||
<paragraph id="17F2F393FCF2694FA234ABBB287601B3" pageId="0" pageNumber="295">Named after the type locality, Tengchong (Yunnan).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |