165 lines
21 KiB
XML
165 lines
21 KiB
XML
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<mods:title id="293AB092F4119BADD618E21D10AAC60F">Revision of the genus Exaesiopus Reichardt, 1926 (Coleoptera, Histeridae, Saprininae)</mods:title>
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<mods:namePart id="118181C07D81211E8175FB87EE92A2BD">Lackner, Tomas</mods:namePart>
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<mods:date id="A9F69353AEC48B10DDD81E93B9F62EBE">2015</mods:date>
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<mods:number id="5FA94F528080CDD849CDADF4E788B735">479</mods:number>
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<mods:classification id="3FD3FDFD4143E9590B5041DA00CFB0CD">journal article</mods:classification>
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<mods:identifier id="FF189FB11A0BFC110DF1930B367787B0" type="DOI">http://dx.doi.org/10.3897/zookeys.479.8738</mods:identifier>
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<treatment id="115FFF770BB1B0A28CD1C298C110C8AA" ID-GBIF-Taxon="152058094" LSID="urn:lsid:plazi:treatment:115FFF770BB1B0A28CD1C298C110C8AA" httpUri="http://treatment.plazi.org/id/115FFF770BB1B0A28CD1C298C110C8AA" lastPageNumber="81" pageId="16" pageNumber="81">
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<subSubSection id="A0F532514541D7FD9EEAD4F49CC347F7" pageId="16" pageNumber="81" type="multiple">
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<paragraph id="F1381CDD66DB267709F30B639AACA70D" pageId="16" pageNumber="81">Taxon classification Animalia Coleoptera Histeridae</paragraph>
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</subSubSection>
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<subSubSection id="6E98E34D83D9EC2264053F101B921821" pageId="16" pageNumber="81" type="nomenclature">
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<paragraph id="FC0C508FCE3EEBD92D70691BD92AE487" pageId="16" pageNumber="81">
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<taxonomicName id="1E46432DB4F55170CEAD911ADD00F339" ID-CoL="3DJJM" authority="Reichardt, 1926" authorityName="Reichardt" authorityYear="1926" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus torvus" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="torvus">Exaesiopus torvus Reichardt, 1926</taxonomicName>
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Figs 55, 56-63, 64-72
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</paragraph>
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</subSubSection>
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<subSubSection id="1886E33DFCA90F91B7D9753E8D0A7EED" pageId="16" pageNumber="81" type="reference_group">
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<paragraph id="88F3CB286F6C7E3FC722000F8C23CFD5" pageId="16" pageNumber="81">
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<taxonomicName id="66E521E52BF366B0B65C70FEF06AD049" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus torvus" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="torvus">Exaesiopus torvus</taxonomicName>
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<bibRefCitation id="102F5440F51D50AF32B738D367C9D307" author="Reichardt, A" journalOrPublisher="Entomologische Blaetter" pageId="33" pageNumber="98" pagination="12 - 18" title="Ueber die mit Pachylopus verwandten Arten." volume="22" year="1926">Reichardt 1926</bibRefCitation>
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: 17;
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<bibRefCitation id="CECDB74C46BE4F8EFD1974CD01EBA586" author="Mazur, S" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="33" pageNumber="98" pagination="1 - 376" title="A world catalogue of Histeridae." volume="54" year="1984">Mazur (1984)</bibRefCitation>
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: 101;
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<bibRefCitation id="43D22D82A0A365A747184D3231DD51B1" pageId="16" pageNumber="81">(1997)</bibRefCitation>
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: 264;
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<bibRefCitation id="1BA6E999B21443D3B93665FB2EAAA0EA" author="Mazur, S" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="33" pageNumber="98" pagination="68 - 102" title="Family Histeridae." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinoidea" year="2004">(2004)</bibRefCitation>
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: 92;
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<bibRefCitation id="A914D87F42031DCC54B503257EB6AD18" author="Mazur, S" journalOrPublisher="Warsaw University of Life Sciences, SGGW Press, Warsaw" pageId="33" pageNumber="98" title="A concise catalogue of the Histeridae (Coleoptera)." year="2011">(2011)</bibRefCitation>
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: 210.
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</paragraph>
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</subSubSection>
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<subSubSection id="47535389038BE0524CDEE5AA3D1EE432" pageId="16" pageNumber="81" type="type locality">
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<paragraph id="97C8F774CD67856EFE7993C9AC805871" pageId="16" pageNumber="81">Type locality.</paragraph>
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<paragraph id="F0777D37D95933C0FA8089A95C46AA02" pageId="16" pageNumber="81">Yanidarya, Kyzyl-Ordinskij Rayon, Kazakhstan.</paragraph>
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</subSubSection>
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<subSubSection id="9031B1B1F08C15E658FB0F48655B175E" pageId="16" pageNumber="81" type="type material examined">
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<paragraph id="6D6BE91EA9DF16F5C91CB0C583518128" pageId="16" pageNumber="81">Type material examined.</paragraph>
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<paragraph id="B4D18B6F1F2FF109138795F80F096EA7" pageId="16" pageNumber="81">
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Holotype, ♀ side-mounted on a triangular point, with female genitalia extracted and glued to the subsequent label with female sign, with following labels:
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<normalizedToken id="7EB27B3666F38607A235DB3E7A55F9F1" originalValue="“♀”">"♀"</normalizedToken>
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(printed); followed by circular golden label; followed by: "Yany - Darya / perovsk u / Kyzyl-Kum / 24.iv.[1]911" (hand-written); followed by: "Type /
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<taxonomicName id="4C89966DBE8F3AE603CE36611FC05FD4" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="genus">Exaesiopus</taxonomicName>
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/
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<taxonomicName id="C174B867EBDABB9E4655FB102097DBD5" lsidName="torvus" pageId="16" pageNumber="81" rank="species" species="torvus">torvus</taxonomicName>
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m. / A. Reichardt det." (written-printed label); followed by:
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<normalizedToken id="253B3B63AE3F799D2D686EB310C450EC" originalValue="“Holotypus”">"Holotypus"</normalizedToken>
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(red label, printed) (ZIN).
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</paragraph>
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</subSubSection>
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<subSubSection id="7F6422B98619B83A96A1C22EDE8207D1" pageId="16" pageNumber="81" type="additional material examined">
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<paragraph id="F001F296C002E16536DB9DFDFE08D2FE" pageId="16" pageNumber="81">Additional material examined.</paragraph>
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<paragraph id="8070EAD813776FE20B58E47D6104757F" pageId="16" pageNumber="81">Kazakhstan: 1 ♀, left bank of the river Ural, Saraichikovsk, 8.vi.1932, Lukyanovich leg.; 1 ♀ + 2 specs., left bank of Ural river, opposite of Saraychik, 8.vi.1932, Lukyanovich leg.; 1 ♂, western bank of Aral Sea, Komsomolsk na Ustyurte, 31.v.1978, G. Medvedev; 1 spec., W Kazakhstan obl. [=reg.], Bilj-Agach, 4.vii.1952, L. Arnoldi (in leaf litter). (all exs. ZIN); 1 spec., 1933-102, left bank of the Ural River, Saraychik, 8.vi.[19]32, Lyukanovich (BMNH). Uzbekistan: 1 ♀, Karakum, Khiva, 3.v.1978, leg. Olexa; 1 ♂, ibid, but 1.-5.v.1979 (both exs. TLAN); RUSSIA: 1 spec., Astrakhan Region, Krasniy Yar district, near Dosang vill., 8.v.2009, A. Kovalyov leg. (CAS); 1 spec., ibid, but 14 km NE Dosang vill., barkhan [=sand dune] Tuvayak, 23-24.iv.2008, M. Smirnov leg. (CAS).</paragraph>
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</subSubSection>
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<subSubSection id="49BBD1BE245A9D4BD51E3E7DA97453F6" pageId="16" pageNumber="81" type="diagnostic description">
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<paragraph id="39D8BE563ADA63C07F6657BB809CDCE1" pageId="16" pageNumber="81">Diagnostic description.</paragraph>
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<paragraph id="F243581551472C3CFAF00AC973315DDF" pageId="16" pageNumber="81">
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Body length: PEL: 2.15-2.575 mm; APW: 0.575-0.875 mm; PPW: 1.625-1.925 mm; EW: 1.75-2.125; EL: 1.375-1.625 mm. Body (Fig. 55) strongly convex, cuticle as in
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<taxonomicName id="BD4C6E406FCDFE01D2838B4065F2E87F" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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, but without metallic tinge; legs, mouthparts and antennae dark yellow to light brown. Antennae as in
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<taxonomicName id="285E649B9F60CF7718C2345888D17A84" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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; sensory structures of the antennal club not examined. Mouthparts: labrum obscurely variolate, shallowly depressed medially; mentum as in
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<taxonomicName id="86AE2186A76BBE2DC2B585679BC2909E" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus henoni" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="henoni">Exaesiopus henoni</taxonomicName>
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; rest of the mouthparts similar to those of
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<taxonomicName id="9B4E6E504BA4DE64980D49C2A42D54EC" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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; terminal labial and maxillary palpomeres truncate. Clypeus (Fig. 56) as in
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<taxonomicName id="BF1DA5CB1AE28C785C5447A45A622180" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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, but rugulose-lacunose; frontal, supraorbital and postorbital striae (Fig. 57) as in
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<taxonomicName id="0ACC40EC6BB1EC24D802D585281F1648" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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; frons rugose, with several irregularly shaped carinate transverse rugae or chevrons intermingled with sparse microscopic punctures. Pronotum: pronotal disc, except for irregularly-shaped impunctate (or weakly punctate) median part entirely covered with round dense punctation, forming transverse rugae and confluent laterally, punctation reaches lateral margins; rest of pronotum as in
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<taxonomicName id="C3787A26B0C3B5C062A928B6256A4767" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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.
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</paragraph>
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<caption id="30E0066DBC254072F52F3C238ABB5307" pageId="16" pageNumber="81">
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<paragraph id="21A5A46449B44B0D561AD593EE00A2FD" pageId="16" pageNumber="81">
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Figure 55.
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<taxonomicName id="638E7C0045DC6B61BAF67106B6477F50" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus torvus" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="torvus">Exaesiopus torvus</taxonomicName>
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Reichardt, 1926 habitus.
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</paragraph>
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</caption>
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<caption id="C9E630CA2A7DD512D1B509803530C1C9" pageId="16" pageNumber="81">
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<paragraph id="B61876B15E47AF82DA293581EBE93380" pageId="16" pageNumber="81">
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Figures 56-63.
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<taxonomicName id="6A3FA77607AC75F2B55027FDC22C4133" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus torvus" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="torvus">Exaesiopus torvus</taxonomicName>
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Reichardt, 1926 56 clypeus 57 head, dorsal view 58 propygidium + pygidium 59 prosternum 60 mesoventrite 61 protibia, dorsal view 62 mesotibia, dorsal view 63 metatibia, dorsal view.
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</paragraph>
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</caption>
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<paragraph id="47F24193B38D551E868C07D3C0C7F877" pageId="16" pageNumber="81">
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Elytral humeri not particularly enlarged; inner subhumeral stria present only as a row of several punctures; elytral punctation variable, in most specimens reaching elytral base along fourth elytral interval, punctures often present in all elytral intervals, elytral flanks impunctate; punctures regular and deep, separated by about half to several times their own diameter. Propygidium (Fig. 58) and pygidium as in
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<taxonomicName id="101CCDE8623E7DD58AC157CD7793ADEE" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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, but covered with denser punctation. Prosternum: prosternal foveae (Fig. 59) weakly impressed; prosternal process otherwise similar to that of
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<taxonomicName id="ED424963FA257F643F767A0B7D39574A" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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. Disc of mesoventrite (Fig. 60) with scattered shallow punctures; intercoxal disc of metaventrite, lateral disc of metaventrite and metepisternum generally similar to those of
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<taxonomicName id="BF61F820B320A06E0EC73B68A6AA9DE1" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus henoni" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="henoni">Exaesiopus henoni</taxonomicName>
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. Intercoxal disc of first abdominal sternite as in
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<taxonomicName id="FE89B143472141149C58165E2EF1FEEF" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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. Protibia (Fig. 61) more dilated than that of
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<taxonomicName id="887D486BDFC4EA2CAAD09083EFCBC17E" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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; on outer margin with two widely-spaced low teeth, topped by large triangular denticle followed by two low rounded denticles imbedded in outer protibial margin; protibial spur inconspicuous (absent?) protibia otherwise similar to that of
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<taxonomicName id="163D11D79664BC04606D3769D3ABEBD1" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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. Mesotibia (Fig. 62) generally similar to that of
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<taxonomicName id="94211D3AEC0ACB22ACB1F6B0E6177138" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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. Metatibia (Fig. 63) perhaps most triangularly dilated and thickened of all congeners; outer margin with approximately three widely-spaced tiny denticles; inner margin with a dense row of minuscule rounded denticles; no rows of denticles present between the two rows, surface rugulose-lacunose.
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</paragraph>
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<paragraph id="83892647D02959F4D61DA64F79506761" pageId="16" pageNumber="81">
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Male genitalia. Eighth (Figs 64-65) sternite apically asetose, with tiny vela; eighth tergite apically faintly inwardly arcuate medially; eighth sternite and tergite fused laterally (Fig. 66). Ninth tergite (Figs 67-68) apically almost straight; tenth tergite apically outwardly arcuate, basally only faintly inwardly arcuate. Spiculum gastrale (Figs 69-70) generally similar to that of
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<taxonomicName id="4913AF146CA68A752CC86CACF931FD7A" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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. Aedeagus (Figs 71-72) almost parallel-sided, apex with pseudopores, parameres fused along their apical half (approximately); aedeagus slightly curved ventrad from the lateral view (Fig. 72).
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</paragraph>
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<caption id="A6E80CDADA33FB72DCDB159A326FCEA8" pageId="16" pageNumber="81">
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<paragraph id="8D32C074B96C80596F7FBA063707FB4D" pageId="16" pageNumber="81">
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Figures 64-72.
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<taxonomicName id="AAC14F3A31F802BE707BEB393C41D492" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus torvus" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="torvus">Exaesiopus torvus</taxonomicName>
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Reichardt, 1926 8th sternite + tergite, 64 ventral view 65 ditto, dorsal view 66 ditto, lateral view 67 9th + 10th tergites, dorsal view 68 ditto, lateral view 69 spiculum gastrale, ventral view 70 ditto, lateral view 71 aedeagus, dorsal view 72 ditto, lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="8FC253274EF30FA39A370D7CFBDD56EA" pageId="16" pageNumber="81" type="differential diagnosis">
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<paragraph id="F97072C6CD01D2F17EC4EB406A919CFB" pageId="16" pageNumber="81">Differential diagnosis.</paragraph>
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<paragraph id="8DD3D453141DC014D2583A1645396536" pageId="16" pageNumber="81">
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Generally the most punctate species of
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<taxonomicName id="371965C61E45702236BD50022CB56F6E" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="genus">Exaesiopus</taxonomicName>
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, which can be confused only with densely punctate specimens of
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<taxonomicName id="C1016B75A0A92B66BC2B6AA1646178AA" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="16" pageNumber="81" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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from N Africa. It clearly differs from them by the punctation of pronotum as well as male genitalia (see also Key to species for details).
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</paragraph>
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</subSubSection>
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<subSubSection id="EF2B14DA7A7BE28ADF69C061BC44B33A" pageId="16" pageNumber="81" type="biology">
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<paragraph id="F16A3B59E10CFE9374C12FE9C91529A1" pageId="16" pageNumber="81">Biology.</paragraph>
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<paragraph id="52CB8691B5AE20DA310780F6852594D0" pageId="16" pageNumber="81">Similar to that of other congeners - beetles are found in sand.</paragraph>
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</subSubSection>
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<subSubSection id="071AAD376F804543F56794B3D34F5103" pageId="16" pageNumber="81" type="distribution">
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<paragraph id="382FCA5465501657CA94A5B500CB2CEE" pageId="16" pageNumber="81">Distribution.</paragraph>
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<paragraph id="ABF989CB69D2678BC1930ED38DC698EC" pageId="16" pageNumber="81">Kazakhstan; new to Uzbekistan and Russia.</paragraph>
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</subSubSection>
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</treatment>
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</document> |