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<document id="315D48A0593715C384F415F28D3532CA" ID-DOI="10.5281/zenodo.193406" ID-GBIF-Dataset="5e8043a5-e685-4a21-b696-0f7a706ab223" ID-ISSN="1175-5326" ID-Zenodo-Dep="193406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460138703176" checkinUser="plazi" docAuthor="Toledo, Mario, Spangler, Paul J. &amp; Balke, Michael" docDate="2010" docId="11578788FE52FFCD7DB5F93612D1DCC6" docLanguage="en" docName="zt02347p058.pdf" docOrigin="Zootaxa 2347" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laccodytes neblinae Toledo, Spangler &amp; Balke, 2010, sp.n." docType="treatment" docVersion="9" lastPageNumber="52" masterDocId="ED6EFFF0FE5CFFC27D22FFF3160ED852" masterDocTitle="Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)" masterLastPageNumber="58" masterPageNumber="37" pageNumber="51" updateTime="1698235333712" updateUser="plazi">
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<mods:title id="4EA141E9ADD2774FBE065BEACA2A17E5">Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)</mods:title>
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<mods:namePart id="C59BB949FBDFFEBD07C68F3E9C5265DE">Toledo, Mario</mods:namePart>
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<mods:namePart id="0E87D03790C00EDF0DAD4BD865FC6D36">Spangler, Paul J.</mods:namePart>
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<mods:namePart id="13D438AE6E092BD4092420CFC2D70F65">Balke, Michael</mods:namePart>
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<mods:date id="742C42A32005F44070FCC891F99B8118">2010</mods:date>
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<treatment id="11578788FE52FFCD7DB5F93612D1DCC6" ID-DOI="http://doi.org/10.5281/zenodo.6198380" ID-GBIF-Taxon="124955827" ID-Zenodo-Dep="6198380" LSID="urn:lsid:plazi:treatment:11578788FE52FFCD7DB5F93612D1DCC6" httpUri="http://treatment.plazi.org/id/11578788FE52FFCD7DB5F93612D1DCC6" lastPageId="15" lastPageNumber="52" pageId="14" pageNumber="51">
<subSubSection id="D1E46515FE52FFCC7DB5F936144CDF54" pageId="14" pageNumber="51" type="nomenclature">
<paragraph id="9941369EFE52FFCC7DB5F93617C0DE8D" blockId="14.[151,578,1733,1798]" box="[151,462,1733,1759]" pageId="14" pageNumber="51">
<heading id="C20981F2FE52FFCC7DB5F93617C0DE8D" bold="true" box="[151,462,1733,1759]" fontSize="11" level="1" pageId="14" pageNumber="51" reason="1">
<emphasis id="AB8AEA8CFE52FFCC7DB5F93617C0DE8D" bold="true" box="[151,462,1733,1759]" pageId="14" pageNumber="51">
<taxonomicName id="5EFE4D1DFE52FFCC7DB5F9361782DE8D" ID-CoL="3RLZH" box="[151,396,1733,1759]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="neblinae" status="sp. nov.">
<emphasis id="AB8AEA8CFE52FFCC7DB5F9361782DE8D" bold="true" box="[151,396,1733,1759]" italics="true" pageId="14" pageNumber="51">Laccodytes neblinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0B957F7FE52FFCC7CB1F93617C0DE8D" box="[403,462,1733,1759]" pageId="14" pageNumber="51" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9941369EFE52FFCC7DB5F91F144CDF54" blockId="14.[151,578,1733,1798]" box="[151,578,1772,1798]" pageId="14" pageNumber="51">Figs (6, 8, 11, 14, 25, 33, 49, 59, 62)</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE52FFCC7DB5F8CB17EBDFBC" pageId="14" pageNumber="51" type="materials_examined">
<paragraph id="9941369EFE52FFCC7DB5F8CB154ADF01" blockId="14.[151,1436,1848,2030]" box="[151,836,1848,1875]" pageId="14" pageNumber="51">
<emphasis id="AB8AEA8CFE52FFCC7DB5F8CB1733DF00" bold="true" box="[151,317,1848,1874]" pageId="14" pageNumber="51">
<typeStatus id="4645883CFE52FFCC7DB5F8CB16DADF00" box="[151,212,1848,1874]" pageId="14" pageNumber="51">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E1E9760EFE52FFCC7C66F8CA17CFDF01" box="[324,449,1849,1875]" name="Venezuela" pageId="14" pageNumber="51">Venezuela</collectingCountry>
: Amazonas, Cerro de la Neblina.
</paragraph>
<paragraph id="9941369EFE52FFCC7DE4F8AC17EBDFBC" blockId="14.[151,1436,1848,2030]" pageId="14" pageNumber="51">
<emphasis id="AB8AEA8CFE52FFCC7DE4F8AC1775DF2B" bold="true" box="[198,379,1887,1913]" pageId="14" pageNumber="51">
<typeStatus id="4645883CFE52FFCC7DE4F8AC170FDF2B" box="[198,257,1887,1913]" pageId="14" pageNumber="51">Type</typeStatus>
material.
</emphasis>
<typeStatus id="4645883CFE52FFCC7CA7F89317FCDF28" box="[389,498,1888,1914]" pageId="14" pageNumber="51" type="holotype">Holotype</typeStatus>
3 (
<collectionCode id="FFEFAE5BFE52FFCC7F02F8931472DF28" LSID="urn:lsid:biocol.org:col:34871" box="[544,636,1888,1914]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="51">USNM</collectionCode>
):
<collectingCountry id="E1E9760EFE52FFCC7FB3F8931502DF28" box="[657,780,1888,1914]" name="Venezuela" pageId="14" pageNumber="51">Venezuela</collectingCountry>
, TF Amazonas, Cerro de la Neblina, basecamp,
<quantity id="5E069B7BFE52FFCC786BF8931399DF28" box="[1353,1431,1888,1914]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="51" unit="m" value="140.0">140 m</quantity>
,
<geoCoordinate id="FCCA5059FE52FFCC7DB5F87516E5DFF2" box="[151,235,1926,1952]" direction="north" orientation="latitude" pageId="14" pageNumber="51" precision="925" value="0.8333333">0°50'N</geoCoordinate>
<geoCoordinate id="FCCA5059FE52FFCC7DD1F8751755DFF2" box="[243,347,1926,1952]" direction="west" orientation="longitude" pageId="14" pageNumber="51" precision="925" value="-66.166664">66°10'W</geoCoordinate>
,
<date id="ED40105EFE52FFCC7C4BF8751423DFF2" box="[361,557,1926,1952]" pageId="14" pageNumber="51" value="1985-01-27" valueMax="1985-02-27" valueMin="1985-01-27">27.I.27.II.1985</date>
, P.J. Spangler, P.M. Spangler, R. Faitoute, W.E. Steiner.
<typeStatus id="4645883CFE52FFCC79F4F8751341DFF2" box="[1238,1359,1926,1952]" pageId="14" pageNumber="51" type="paratype">Paratypes</typeStatus>
: Same data as
<typeStatus id="4645883CFE52FFCC7DCDF85E1758DF95" box="[239,342,1965,1991]" pageId="14" pageNumber="51" type="holotype">holotype</typeStatus>
(150 exs
<collectionCode id="FFEFAE5BFE52FFCC7CEBF85E1404DF95" box="[457,522,1965,1991]" pageId="14" pageNumber="51">CMT</collectionCode>
,
<collectionCode id="FFEFAE5BFE52FFCC7F34F85E1460DF95" LSID="urn:lsid:biocol.org:col:34871" box="[534,622,1965,1991]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="51">USNM</collectionCode>
,
<collectionCode id="FFEFAE5BFE52FFCC7F5BF85E14B7DF95" box="[633,697,1965,1991]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="51">ZSM</collectionCode>
,
<collectionCode id="FFEFAE5BFE52FFCC7FE7F85E151BDF95" box="[709,789,1965,1991]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="51">NMW</collectionCode>
); Puerto Ayacucho,
<quantity id="5E069B7BFE52FFCC7922F85E1242DF95" box="[1024,1100,1965,1991]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="51" unit="km" value="37.0">37 km</quantity>
SE Rio Gavilan,
<date id="ED40105EFE52FFCC7839F85E1399DF95" box="[1307,1431,1965,1991]" pageId="14" pageNumber="51" value="1986-02-22">22.II.1986</date>
, P.J. Spangler (1 ex.
<collectionCode id="FFEFAE5BFE52FFCC7C5DF82717D7DFBC" LSID="urn:lsid:biocol.org:col:34871" box="[383,473,2004,2030]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="51">USNM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE53FFCD7DE4FF64145EDBAE" pageId="15" pageNumber="52" type="diagnosis">
<paragraph id="9941369EFE53FFCD7DE4FF64130DD88A" blockId="15.[151,1437,151,1172]" pageId="15" pageNumber="52">
<emphasis id="AB8AEA8CFE53FFCD7DE4FF641748D8E3" bold="true" box="[198,326,151,177]" pageId="15" pageNumber="52">Diagnosis.</emphasis>
Habitus (
<figureCitation id="01C52A1BFE53FFCD7CE2FF6B1410D8E0" box="[448,542,152,178]" captionStart="FIGURES 28 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="15" pageNumber="52">Figs 33</figureCitation>
a, b). Body length 1.71.8 mm. Body slightly drop-shaped, tip of elytron slightly truncate.
<collectingCountry id="E1E9760EFE53FFCD7C41FF4D1790D88A" box="[355,414,190,216]" name="India" pageId="15" pageNumber="52">Hind</collectingCountry>
angle of pronotum rounded. Angle between pronotum and elytra missing.
</paragraph>
<paragraph id="9941369EFE53FFCD7DE4FF171295D9C5" blockId="15.[151,1437,151,1172]" pageId="15" pageNumber="52">
Color. Head and pronotum dark yellow; base of pronotum black. Elytron brown with obscure paler pattern of dark yellowish patches in the following positions (
<figureCitation id="01C52A1BFE53FFCD7E23FEF91554D976" box="[769,858,266,292]" captionStart="FIGURES 28 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="15" pageNumber="52">Figs 33</figureCitation>
a, b for explanation): subbasal-subsutural; mediomarginal to medio-discal; apical or subapical. These patches are not contrasting and sometimes extremely obscure. Appendages yellowish. Ventral side of head and prosternum mainly dark yellow; epipleuron, metaventrite, metacoxa and metacoxal process mainly brown; abdomen dark yellowish.
</paragraph>
<paragraph id="9941369EFE53FFCD7DE4FE5012E0DA62" blockId="15.[151,1437,151,1172]" pageId="15" pageNumber="52">
Sculpture. The entire dorsal and ventral surface of the beetle with well impressed MR of regular, polygonal meshes. Few punctures on pronotum, elytron and metaventrite visible, denser and deeper on lateral expansions of metaventrite (“metasternal wings”). Metacoxal shagreenation similar to
<taxonomicName id="5EFE4D1DFE53FFCD79B8FE03132ADA5B" box="[1178,1316,496,522]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="52" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="AB8AEA8CFE53FFCD79B8FE0312A4DA5B" box="[1178,1194,496,521]" italics="true" pageId="15" pageNumber="52">L</emphasis>
.
<emphasis id="AB8AEA8CFE53FFCD7998FE03132ADA5B" box="[1210,1316,496,521]" italics="true" pageId="15" pageNumber="52">apalodes</emphasis>
</taxonomicName>
(
<figureCitation id="01C52A1BFE53FFCD7814FE031380DA58" box="[1334,1422,496,522]" captionStart="FIGURES 17 25" captionStartId="4.[151,270,1905,1929]" captionTargetBox="[163,1426,187,1879]" captionTargetId="figure@4.[163,1428,187,1880]" captionTargetPageId="4" captionText="FIGURES 17 25. SEMs of MR in some Laccodytes species: 17) detail of elytron of L. americanus (magn. 2100 x); 18) idem, (magn. 2600 x); 19) metacoxa (MTX) and metaventrite (MTS) of L. americanus; 20) idem, detail at higher magnification; 21) idem, detail of metaventrite (MTX) at higher magnification; 22) detail of elytron of L. apalodes; 23) detail of metacoxa of L. apalodes; 24) idem, detail at higher magnification; 25) detail of metacoxa of L. neblinae." httpUri="https://zenodo.org/record/193409/files/figure.png" pageId="15" pageNumber="52">Fig. 25</figureCitation>
). A series of transverse shallow grooves is visible at about the anterior third of metacoxal plates.
</paragraph>
<paragraph id="9941369EFE53FFCD7DE4FDCE1742DB44" blockId="15.[151,1437,151,1172]" pageId="15" pageNumber="52">
Structures. Pronotum with narrow lateral bead; posterior angle rounded. Prosternum and prosternal process with small ridge; prosternal process broadly carinate and with a moderately long, needle-like, acute tip. Epipleuron broad up to level of sternite 6 or 7. Fore and middle legs very long and slender. Metatarsomeres 14 with apicolateral angle slightly lobed (
<figureCitation id="01C52A1BFE53FFCD7E7DFD5C15B6DA9B" box="[863,952,687,713]" captionStart="FIGURES 3 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="15" pageNumber="52">Fig. 14</figureCitation>
). Outer spur of metatibia broad and flat, acuminate at tip.
<collectingCountry id="E1E9760EFE53FFCD7C44FD2517ACDAA2" box="[358,418,726,752]" name="India" pageId="15" pageNumber="52">Hind</collectingCountry>
lobes of metacoxal process poorly rounded, divided by a deep and narrow, V-shaped notch (
<figureCitation id="01C52A1BFE53FFCD7DCAFD0F1730DB44" box="[232,318,764,790]" captionStart="FIGURES 3 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="15" pageNumber="52">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="9941369EFE53FFCD7DE4FCD114E0DBFD" blockId="15.[151,1437,151,1172]" pageId="15" pageNumber="52">
Male. Pro- and mesotarsi not dilated.
<collectingCountry id="E1E9760EFE53FFCD7FACFCD114C7DB6E" box="[654,713,802,828]" name="India" pageId="15" pageNumber="52">Hind</collectingCountry>
margin of sternite 7 rounded (
<figureCitation id="01C52A1BFE53FFCD791EFCD11298DB6E" box="[1084,1174,802,828]" captionStart="FIGURES 54 61" captionStartId="12.[151,273,1107,1131]" captionTargetBox="[153,1431,196,1081]" captionTargetId="figure@12.[151,1436,194,1083]" captionTargetPageId="12" captionText="FIGURES 54 61. Last abdominal segment (left male; right female) in Laccodytes species: 54) L. apalodes; 55) L. rondonia; 56) L. americanus; 57) L. phalacroides; 58) L. takutuanus; 59) L. neblinae; 60) L. bassignanii; 61) Laccodytes sp. 1 female." httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="15" pageNumber="52">Fig. 59</figureCitation>
). Aedeagus (
<figureCitation id="01C52A1BFE53FFCD7811FCD11381DB6E" box="[1331,1423,802,828]" captionStart="FIGURES 48 53" captionStartId="11.[151,270,1543,1567]" captionTargetBox="[189,1376,223,1499]" captionTargetId="figure@11.[184,1403,197,1518]" captionTargetPageId="11" captionText="FIGURES 48 53. Male genitals in Laccodytes species: 48) L. takutuanus: a) median lobe of aedeagus in dorsal view, b) in lateral view, c-d) right and left parameres; 49) L. neblinae: a) median lobe of aedeagus in dorsal view, b) in lateral view, c-d) right and left parameres; 50) L. bassignanii: a) median lobe of aedeagus in dorsal view, b-c) in dorsal-lateral and lateral view, d-e) right and left parameres; 51) L. obscuratus: a) median lobe of aedeagus in dorsal view, b) in lateral view, c-d) right and left parameres; 52) L. olibroides: a) median lobe of aedeagus in lateral view, b-c) right an left parameres; 53) L. androginus: a) median lobe of aedeagus in dorsal view, b) in lateral view, c-d) right and left parameres." httpUri="https://zenodo.org/record/193414/files/figure.png" pageId="15" pageNumber="52">Fig. 49</figureCitation>
): median lobe elongated, rounded along dorsal side, straight ventrally, with tip acuminate: gently bent down in lateral view; gently turned to the right in dorsal view. Parameres small and rounded, almost of the same size and quite similar in shape, both with an apical seta.
</paragraph>
<paragraph id="9941369EFE53FFCD7DE4FC48145EDBAE" blockId="15.[151,1437,151,1172]" pageId="15" pageNumber="52">
Female. Sternite 7 (
<figureCitation id="01C52A1BFE53FFCD7C89FC48140FDB87" box="[427,513,955,981]" captionStart="FIGURES 54 61" captionStartId="12.[151,273,1107,1131]" captionTargetBox="[153,1431,196,1081]" captionTargetId="figure@12.[151,1436,194,1083]" captionTargetPageId="12" captionText="FIGURES 54 61. Last abdominal segment (left male; right female) in Laccodytes species: 54) L. apalodes; 55) L. rondonia; 56) L. americanus; 57) L. phalacroides; 58) L. takutuanus; 59) L. neblinae; 60) L. bassignanii; 61) Laccodytes sp. 1 female." httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="15" pageNumber="52">Fig. 59</figureCitation>
) slightly concave on both sides, hind margin with a deep and narrow V-shaped emargination, almost acute on vertex.
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE53FFCD7DE4FBFB150CDC70" box="[198,770,1032,1058]" pageId="15" pageNumber="52" type="distribution">
<paragraph id="9941369EFE53FFCD7DE4FBFB150CDC70" blockId="15.[151,1437,151,1172]" box="[198,770,1032,1058]" pageId="15" pageNumber="52">
<emphasis id="AB8AEA8CFE53FFCD7DE4FBFB1751DC70" bold="true" box="[198,351,1032,1058]" pageId="15" pageNumber="52">Distribution</emphasis>
(
<figureCitation id="01C52A1BFE53FFCD7C4DFBFB17C9DC70" box="[367,455,1032,1058]" captionStart="FIGURE 62" captionStartId="18.[151,255,1941,1965]" captionText="FIGURE 62. Known distribution of Laccodytes species." pageId="15" pageNumber="52">Fig. 62</figureCitation>
). Southwestern
<collectingCountry id="E1E9760EFE53FFCD7FA0FBFB14F0DC70" box="[642,766,1032,1058]" name="Venezuela" pageId="15" pageNumber="52">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE53FFCD7DE4FBDD15CFDC3C" pageId="15" pageNumber="52" type="biology_ecology">
<paragraph id="9941369EFE53FFCD7DE4FBDD15CFDC3C" blockId="15.[151,1437,151,1172]" pageId="15" pageNumber="52">
<emphasis id="AB8AEA8CFE53FFCD7DE4FBDD1724DC1A" bold="true" box="[198,298,1070,1096]" pageId="15" pageNumber="52">Biology.</emphasis>
Specimens were seined from rocks in the rapids of Rio Baria; some 30 exemplars from Cerro de la Neblina were picked by hand from leaf packs taken from a stream.
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE53FFCD7DE4FB8912D1DCC6" box="[198,1247,1146,1172]" pageId="15" pageNumber="52" type="etymology">
<paragraph id="9941369EFE53FFCD7DE4FB8912D1DCC6" blockId="15.[151,1437,151,1172]" box="[198,1247,1146,1172]" pageId="15" pageNumber="52">
<emphasis id="AB8AEA8CFE53FFCD7DE4FB8917A3DCC6" bold="true" box="[198,429,1146,1172]" pageId="15" pageNumber="52">Derivatio nominis.</emphasis>
Name derived from Cerro de la Neblina, type-locality of this species.
</paragraph>
</subSubSection>
</treatment>
</document>