366 lines
49 KiB
XML
366 lines
49 KiB
XML
<document id="A07A6FAA3F5B67FEFE78093290251D1F" ID-DOI="10.11646/zootaxa.5543.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14386776" ID-ZooBank="E34AF3EF-4D03-4C08-8E11-C9514D42021B" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1733919186538" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2024" docId="03C83A1CFF96C34FFF772CC6519F431C" docLanguage="en" docName="zootaxa.5543.4.1.pdf" docOrigin="Zootaxa 5543 (4)" docSource="https://doi.org/10.11646/zootaxa.5543.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceramaster palmyra Mah, 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="465" masterDocId="FFF14264FF9DC341FFE02E495341417D" masterDocTitle="New genera and species of deep-sea Goniasteridae (Asteroidea) from the North Pacific" masterLastPageNumber="500" masterPageNumber="451" pageNumber="462" updateTime="1734434064719" updateUser="guilherme" zenodo-license-document="CLOSED">
|
||
<mods:mods id="DE714B3C451CBD47037058A5020B3E1F" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="30E77D28F04A11D885D7D7354F817995">
|
||
<mods:title id="635ECEE586549F4E996D7830F282F742">New genera and species of deep-sea Goniasteridae (Asteroidea) from the North Pacific</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="23024AED0356C8104D25FEDE0E0CA37C" type="personal">
|
||
<mods:role id="EDBF93EDB79502EB892812A554FFE603">
|
||
<mods:roleTerm id="6B46C59BB91E19A0CBBC2D6425E1DE3E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D7F5CD237FB2C096BA1CE8AE13FD328A">Mah, Christopher L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="75D81E84C79676F04A9359C7006ABDA5">text</mods:typeOfResource>
|
||
<mods:relatedItem id="8C234B5E25F86FAEE5717F95D7F50A5E" type="host">
|
||
<mods:titleInfo id="59F647018FF7FDBA3AA940F304F79B18">
|
||
<mods:title id="853A12C673889DE4B22A1D00A68B6FC8">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="FAE424F6E7C743434D427A64A8C11A71">
|
||
<mods:date id="EA051BD5E85CFB7E87A575ABEBA5F757">2024</mods:date>
|
||
<mods:detail id="7F58358C861659E346DF2A2CC152CB63" type="pubDate">
|
||
<mods:number id="4B43950C74E859692F51B4112D5E206F">2024-12-05</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C879932265D08EE4C5FC957F7A4B3478" type="volume">
|
||
<mods:number id="4DB93A5B296C96259EB5BC04EBE235ED">5543</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="D14E936BAE3C9DB776951EA38F8FD49C" type="issue">
|
||
<mods:number id="4950973B8BDAD68082AFD0456E2EE3E6">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="1FC672C8F29B55786B686D60ACDBE849" unit="page">
|
||
<mods:start id="26354882856DA72964FEFF087CB5EF6E">451</mods:start>
|
||
<mods:end id="B031C1D1D9EAEDCB9606C8DC13B3FF36">500</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="00FB92E618DFE4118510FA674AE2BD81">
|
||
<mods:url id="044A8A137FD08809969BE7F1E8C1AFA9">https://doi.org/10.11646/zootaxa.5543.4.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="5EC3E335523F82A6BD1341DA108895F1">journal article</mods:classification>
|
||
<mods:identifier id="3EC05F021FA499C65C86BCF5C990F454" type="DOI">10.11646/zootaxa.5543.4.1</mods:identifier>
|
||
<mods:identifier id="FA680D3F21749BE10C46C18D142DCF79" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="7442D022DAA03DA450AB01F18BAC01C9" type="Zenodo-Dep">14386776</mods:identifier>
|
||
<mods:identifier id="73B05E7A105C53259F974CBF274151D2" type="ZooBank">E34AF3EF-4D03-4C08-8E11-C9514D42021B</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C83A1CFF96C34FFF772CC6519F431C" LSID="urn:lsid:plazi:treatment:03C83A1CFF96C34FFF772CC6519F431C" httpUri="http://treatment.plazi.org/id/03C83A1CFF96C34FFF772CC6519F431C" lastPageId="14" lastPageNumber="465" pageId="11" pageNumber="462">
|
||
<subSubSection id="C37BD881FF96C34AFF772CC6529943D4" box="[151,472,654,681]" pageId="11" pageNumber="462" type="nomenclature">
|
||
<paragraph id="8BDE8B0AFF96C34AFF772CC6529943D4" blockId="11.[151,472,654,717]" box="[151,472,654,681]" pageId="11" pageNumber="462">
|
||
<heading id="D0963C66FF96C34AFF772CC6529943D4" bold="true" box="[151,472,654,681]" fontSize="11" level="1" pageId="11" pageNumber="462" reason="1">
|
||
<emphasis id="B9155718FF96C34AFF772CC6529943D4" bold="true" box="[151,472,654,681]" pageId="11" pageNumber="462">
|
||
<taxonomicName id="4C61F089FF96C34AFF772CC652CF43D4" authorityName="Mah" authorityYear="2024" box="[151,398,654,681]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="palmyra" status="sp. nov.">
|
||
<emphasis id="B9155718FF96C34AFF772CC652CF43D4" bold="true" box="[151,398,654,681]" italics="true" pageId="11" pageNumber="462">Ceramaster palmyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A226EA63FF96C34AFE752CC6529943D4" box="[405,472,655,681]" pageId="11" pageNumber="462" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37BD881FF96C34AFF772CFA52C343B0" box="[151,386,691,717]" pageId="11" pageNumber="462" type="description">
|
||
<paragraph id="8BDE8B0AFF96C34AFF772CFA52C343B0" blockId="11.[151,472,654,717]" box="[151,386,691,717]" pageId="11" pageNumber="462">
|
||
<figureCitation id="135A978FFF96C34AFF772CFA520943B0" box="[151,328,691,717]" captionStart="FIGURE 4" captionStartId="12.[151,250,1666,1690]" captionTargetBox="[151,1435,181,1642]" captionTargetId="figure-24@12.[151,1435,181,1642]" captionTargetPageId="12" captionText="FIGURE 4. Ceramaster palmyra n. sp. MCZ 153216. Holotype. A. Abactinal. B. Closeup abactinal plates. C. Armtip and superomarginal plates. D. Actinal. E. Adambulacral plates with furrow and subambulacral spines. Scale bar=5.0 mm: A, B; Scale bar=1.0 mm: B, E; Scale bar=3.0 mm: C." figureDoi="http://doi.org/10.5281/zenodo.14386790" httpUri="https://zenodo.org/record/14386790/files/figure.png" pageId="11" pageNumber="462">FIGURE 4A–E</figureCitation>
|
||
,
|
||
<figureCitation id="135A978FFF96C34AFEB32CFA52C343B0" box="[339,386,691,717]" captionStart="FIGURE 12" captionStartId="33.[151,250,1651,1675]" captionTargetBox="[151,1436,181,1627]" captionTargetId="figure-15@33.[151,1436,181,1627]" captionTargetPageId="33" captionText="FIGURE 12. Ahuastra and Ceramaster palmyra observed in situ. A. Ahuastra kupanaha nov. gen. nov. sp. in situ, showing apparent feeding. B. Ceramaster palmyra n. sp. in situ." figureDoi="http://doi.org/10.5281/zenodo.14386822" httpUri="https://zenodo.org/record/14386822/files/figure.png" pageId="11" pageNumber="462">12B</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37BD881FF96C34AFF772CB257384220" box="[151,1145,763,861]" pageId="11" pageNumber="462" type="etymology">
|
||
<paragraph id="8BDE8B0AFF96C34AFF772CB2525F4268" blockId="11.[151,286,763,789]" box="[151,286,763,789]" pageId="11" pageNumber="462">
|
||
<emphasis id="B9155718FF96C34AFF772CB2525F4268" bold="true" box="[151,286,763,789]" pageId="11" pageNumber="462">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF96C34AFF772D0A57384220" blockId="11.[151,1145,835,861]" box="[151,1145,835,861]" pageId="11" pageNumber="462">
|
||
Species epithet is named for the
|
||
<typeStatus id="54DA35A8FF96C34AFDE32D0A51724220" box="[515,563,835,861]" pageId="11" pageNumber="462">type</typeStatus>
|
||
locality,
|
||
<collectingCountry id="F376CB9AFF96C34AFD7A2D0A507B4220" box="[666,826,835,861]" name="United States Pacific Island Wildlife Refuges" pageId="11" pageNumber="462">Palmyra Atoll</collectingCountry>
|
||
.
|
||
<location id="8EBEDDD1FF96C34AFCA52D0A50C94220" LSID="urn:lsid:plazi:treatment:03C83A1CFF96C34FFF772CC6519F431C:8EBEDDD1FF96C34AFCA52D0A50C94220" box="[837,904,835,861]" country="United States Pacific Island Wildlife Refuges" name="Name" pageId="11" pageNumber="462">Name</location>
|
||
is held in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37BD881FF96C34AFF772DE750C94500" pageId="11" pageNumber="462" type="diagnosis">
|
||
<paragraph id="8BDE8B0AFF96C34AFF772DE7525142B5" blockId="11.[151,272,942,968]" box="[151,272,942,968]" pageId="11" pageNumber="462">
|
||
<emphasis id="B9155718FF96C34AFF772DE7525142B5" bold="true" box="[151,272,942,968]" pageId="11" pageNumber="462">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF96C34AFF772DBE50C94500" blockId="11.[151,1437,1015,1149]" pageId="11" pageNumber="462">Body stellate (R/r=2.0), stout, arms triangular with curved interradial arcs. Abactinal plates with large peripheral granules and smaller central granules, the former approximately twice the size of the latter. Superomarginal plate tumid, significantly arched, abactinal surface completely bald, save for clustered granules on lateral side. Furrow spines slender, 4 –7, subambulacrals thick in 2 rows, 3 spines each.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37BD881FF96C34FFF772A87515140F4" lastPageId="14" lastPageNumber="465" pageId="11" pageNumber="462" type="discussion">
|
||
<paragraph id="8BDE8B0AFF96C34AFF772A87525F4595" blockId="11.[151,286,1230,1256]" box="[151,286,1230,1256]" pageId="11" pageNumber="462">
|
||
<heading id="D0963C66FF96C34AFF772A87525F4595" bold="true" box="[151,286,1230,1256]" fontSize="11" level="1" pageId="11" pageNumber="462" reason="1">
|
||
<emphasis id="B9155718FF96C34AFF772A87525F4595" bold="true" box="[151,286,1230,1256]" pageId="11" pageNumber="462">Comments</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF96C34AFF772B5E57E44404" blockId="11.[151,1437,1303,2013]" pageId="11" pageNumber="462">
|
||
<taxonomicName id="4C61F089FF96C34AFF772B5E52C2444C" authorityName="Mah" authorityYear="2024" box="[151,387,1303,1329]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="palmyra" status="sp. nov.">
|
||
<emphasis id="B9155718FF96C34AFF772B5E52C2444C" box="[151,387,1303,1329]" italics="true" pageId="11" pageNumber="462">Ceramaster palmyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A226EA63FF96C34AFE6F2B5E528F444C" box="[399,462,1303,1329]" pageId="11" pageNumber="462" rank="species">n. sp.</taxonomicNameLabel>
|
||
is based on the strongly tumid superomarginal plate dorsal surface, the absence of granulation from the superomarginal surfaces, slender furrow spines and large, trapezoidal granules on the abactinal plates. Peripheral abactinal granules are approximately twice the size of the central granules.
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF96C34AFF272BCA57E1479C" blockId="11.[151,1437,1303,2013]" pageId="11" pageNumber="462">
|
||
Although a full overview of
|
||
<taxonomicName id="4C61F089FF96C34AFDEF2BCA518B44E0" box="[527,714,1411,1437]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="undetermined">
|
||
<emphasis id="B9155718FF96C34AFDEF2BCA51D244E0" box="[527,659,1411,1437]" italics="true" pageId="11" pageNumber="462">Ceramaster</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
is beyond the scope herein, comparison of this species showed general morphological similarity with
|
||
<taxonomicName id="4C61F089FF96C34AFDB22BEE507344BC" box="[594,818,1447,1473]" pageId="11" pageNumber="462">
|
||
<emphasis id="B9155718FF96C34AFDB22BEE519744BC" box="[594,726,1447,1473]" italics="true" pageId="11" pageNumber="462">Ceramaster</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
that most closely cluster with
|
||
<taxonomicName id="4C61F089FF96C34AFB7A2BEE560644BD" baseAuthorityName="Sladen" baseAuthorityYear="1889" box="[1178,1351,1447,1473]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="patagonicus">
|
||
<emphasis id="B9155718FF96C34AFB7A2BEE560644BD" box="[1178,1351,1447,1473]" italics="true" pageId="11" pageNumber="462">C. patagonicus</emphasis>
|
||
</taxonomicName>
|
||
, which displays tabulate abactinal plates, a bald area on the superomarginal plates as well as trapezoidal peripheral granules which vary in size relative to the granules on the central region of the abactinal plate. This includes, but is not limited to the recently described
|
||
<taxonomicName id="4C61F089FF96C34AFDEB285A50104750" authority="Mah 2022" authorityName="Mah" authorityYear="2022" box="[523,849,1555,1581]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="vorax">
|
||
<emphasis id="B9155718FF96C34AFDEB285A51974751" box="[523,726,1555,1581]" italics="true" pageId="11" pageNumber="462">Ceramaster vorax</emphasis>
|
||
<bibRefCitation id="EFF0F6FBFF96C34AFD3E285A50104750" author="Mah, C. L." box="[734,849,1555,1581]" pageId="11" pageNumber="462" pagination="1 - 75" refId="ref22447" refString="Mah, C. L. (2022) New Genera, Species and Occurrence of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the Tropical Pacific Ocean by the CAPSTONE Expedition. Zootaxa, 5164 (1), 1 - 75. https: // doi. org / 10.11646 / zootaxa. 5164.1.1" type="journal article" year="2022">Mah 2022</bibRefCitation>
|
||
</taxonomicName>
|
||
as well as other Pacific species such as
|
||
<taxonomicName id="4C61F089FF96C34AFAF8285A515C472C" authority="Clark & McKnight 2001" authorityName="Clark & McKnight" authorityYear="2001" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="australis">
|
||
<emphasis id="B9155718FF96C34AFAF8285A53B8472C" italics="true" pageId="11" pageNumber="462">Ceramaster australis</emphasis>
|
||
<bibRefCitation id="EFF0F6FBFF96C34AFEE2287E515C472C" author="Clark, H. E. S. & McKnight, D. G." box="[258,541,1591,1617]" pageId="11" pageNumber="462" pagination="1 - 270" refId="ref20789" refString="Clark, H. E. S. & McKnight, D. G. (2001) The marine fauna of New Zealand: Echinodermata: Asteroidea (Sea-stars), Order Valvatida. NIWA Biodiversity Memoir, 117, 1 - 270." type="journal article" year="2001">Clark & McKnight 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C61F089FF96C34AFDCA287E50F4472C" authority="(Verrill 1909)" baseAuthorityName="Verrill" baseAuthorityYear="1909" box="[554,949,1591,1617]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="arcticus">
|
||
<emphasis id="B9155718FF96C34AFDCA287E504E472D" box="[554,783,1591,1617]" italics="true" pageId="11" pageNumber="462">Ceramaster arcticus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF0F6FBFF96C34AFCFF287E50EC472C" author="Verrill, A. E." box="[799,941,1591,1617]" pageId="11" pageNumber="462" pagination="59 - 70" refId="ref23408" refString="Verrill, A. E. (1909) Description of new genera and species of starfishes from the North Pacific coast of America. American Journal of Science, 28, 59 - 70. https: // doi. org / 10.2475 / ajs. s 4 - 28.163.59" type="journal article" year="1909">Verrill 1909</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C61F089FF96C34AFC22287E562C472C" authority="(Sladen, 1889)" baseAuthorityName="Sladen" baseAuthorityYear="1889" box="[962,1389,1591,1617]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="japonicus">
|
||
<emphasis id="B9155718FF96C34AFC22287E57FD472D" box="[962,1212,1591,1617]" italics="true" pageId="11" pageNumber="462">Ceramaster japonicus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF0F6FBFF96C34AFB2C287E5624472C" author="Sladen, W. P." box="[1228,1381,1591,1617]" pageId="11" pageNumber="462" pagination="1 - 893" refId="ref23200" refString="Sladen, W. P. (1889) Asteroidea. Report of the Scientific Results of H. M. S. Challenger, 30, 1 - 893." type="journal article" year="1889">Sladen, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the Atlantic
|
||
<taxonomicName id="4C61F089FF96C34AFF1A281251EA4708" authority="(Retzius 1783)" baseAuthorityName="Retzius" baseAuthorityYear="1783" box="[250,683,1627,1653]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="granularis">
|
||
<emphasis id="B9155718FF96C34AFF1A281252BD4708" box="[250,508,1627,1653]" italics="true" pageId="11" pageNumber="462">Ceramaster granularis</emphasis>
|
||
(Retzius 1783)
|
||
</taxonomicName>
|
||
as well as
|
||
<taxonomicName id="4C61F089FF96C34AFCCB281250974709" baseAuthorityName="Sladen" baseAuthorityYear="1889" box="[811,982,1627,1653]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="patagonicus">
|
||
<emphasis id="B9155718FF96C34AFCCB281250974709" box="[811,982,1627,1653]" italics="true" pageId="11" pageNumber="462">C. patagonicus</emphasis>
|
||
</taxonomicName>
|
||
itself which is widely occurring at high latitudes. Other
|
||
<taxonomicName id="4C61F089FF96C34AFEB02836517047E4" box="[336,561,1663,1689]" pageId="11" pageNumber="462">
|
||
<emphasis id="B9155718FF96C34AFEB02836529547E4" box="[336,468,1663,1689]" italics="true" pageId="11" pageNumber="462">Ceramaster</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, such as the deep-sea North Pacific
|
||
<taxonomicName id="4C61F089FF96C34AFC342836561947E4" authority="Mah 2016" authorityName="Mah" authorityYear="2016" box="[980,1368,1663,1689]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="pointsurae">
|
||
<emphasis id="B9155718FF96C34AFC342836579B47E5" box="[980,1242,1663,1689]" italics="true" pageId="11" pageNumber="462">Ceramaster pointsurae</emphasis>
|
||
<bibRefCitation id="EFF0F6FBFF96C34AFB032836561947E4" author="Mah, C. L." box="[1251,1368,1663,1689]" pageId="11" pageNumber="462" pagination="101 - 141" refId="ref22226" refString="Mah, C. L. (2016) Deep-sea (> 1000 m) Goniasteridae (Valvatida; Asteroidea) from the North Pacific, including an overview of Sibogaster, Bathyceramaster n. gen. and three new species. Zootaxa, 4175 (2), 101 - 141. https: // doi. org / 10.11646 / zootaxa. 4175.2.1" type="journal article" year="2016">Mah 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
show strong morphological affinities with the Atlantic
|
||
<taxonomicName id="4C61F089FF96C34AFD2828EA57CD47C0" authority="(Perrier, 1881)" baseAuthorityName="Perrier" baseAuthorityYear="1881" box="[712,1164,1699,1725]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="grenadensis">
|
||
<emphasis id="B9155718FF96C34AFD2828EA509D47C0" box="[712,988,1699,1725]" italics="true" pageId="11" pageNumber="462">Ceramaster grenadensis</emphasis>
|
||
(Perrier, 1881)
|
||
</taxonomicName>
|
||
and based on character comparisons form a separate cluster from those which share characters with
|
||
<taxonomicName id="4C61F089FF96C34AFC0F288E57D8479D" baseAuthorityName="Sladen" baseAuthorityYear="1889" box="[1007,1177,1735,1761]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="patagonicus">
|
||
<emphasis id="B9155718FF96C34AFC0F288E57D8479D" box="[1007,1177,1735,1761]" italics="true" pageId="11" pageNumber="462">C. patagonicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF96C34AFF2728A252AC46A0" blockId="11.[151,1437,1303,2013]" pageId="11" pageNumber="462">
|
||
This species is distinguished from
|
||
<taxonomicName id="4C61F089FF96C34AFD8328A251894679" authorityName="Mah" authorityYear="2022" box="[611,712,1771,1797]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="vorax">
|
||
<emphasis id="B9155718FF96C34AFD8328A251894679" box="[611,712,1771,1797]" italics="true" pageId="11" pageNumber="462">C. vorax</emphasis>
|
||
</taxonomicName>
|
||
based primarily on the completely bald superomarginal plate surface, which in
|
||
<taxonomicName id="4C61F089FF96C34AFEBC294652FA4655" authorityName="Mah" authorityYear="2022" box="[348,443,1807,1833]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="vorax">
|
||
<emphasis id="B9155718FF96C34AFEBC294652FA4655" box="[348,443,1807,1833]" italics="true" pageId="11" pageNumber="462">C. vorax</emphasis>
|
||
</taxonomicName>
|
||
is limited to a discrete region as well as the presence of pedicellariae, which were present in
|
||
<taxonomicName id="4C61F089FF96C34AFF53297A52524631" authorityName="Mah" authorityYear="2022" box="[179,275,1843,1869]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="vorax">
|
||
<emphasis id="B9155718FF96C34AFF53297A52524631" box="[179,275,1843,1869]" italics="true" pageId="11" pageNumber="462">C. vorax</emphasis>
|
||
</taxonomicName>
|
||
, but not observed on
|
||
<taxonomicName id="4C61F089FF96C34AFDE1297A51A94630" authorityName="Mah" authorityYear="2024" box="[513,744,1843,1869]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="palmyra" status="sp. nov.">
|
||
<emphasis id="B9155718FF96C34AFDE1297A51A94630" box="[513,744,1843,1869]" italics="true" pageId="11" pageNumber="462">Ceramaster palmyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A226EA63FF96C34AFD0F297A506B4630" box="[751,810,1843,1869]" pageId="11" pageNumber="462" rank="species">n. sp.</taxonomicNameLabel>
|
||
Subambulacral spines in
|
||
<taxonomicName id="4C61F089FF96C34AFBA6297A57824630" authorityName="Mah" authorityYear="2024" box="[1094,1219,1843,1869]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="palmyra" status="n. sp.">
|
||
<emphasis id="B9155718FF96C34AFBA6297A57824630" box="[1094,1219,1843,1869]" italics="true" pageId="11" pageNumber="462">C. palmyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A226EA63FF96C34AFB29297A56454630" box="[1225,1284,1843,1869]" pageId="11" pageNumber="462" rank="species">n. sp.</taxonomicNameLabel>
|
||
are similar in number and orientation to
|
||
<taxonomicName id="4C61F089FF96C34AFE22291E5162460D" authorityName="Mah" authorityYear="2022" box="[450,547,1879,1905]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="vorax">
|
||
<emphasis id="B9155718FF96C34AFE22291E5162460D" box="[450,547,1879,1905]" italics="true" pageId="11" pageNumber="462">C. vorax</emphasis>
|
||
</taxonomicName>
|
||
but are smaller, more widely spaced and oriented in more regular series. Other shared characters between
|
||
<taxonomicName id="4C61F089FF96C34AFE21293251E846E8" authorityName="Mah" authorityYear="2024" box="[449,681,1915,1941]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="palmyra" status="sp. nov.">
|
||
<emphasis id="B9155718FF96C34AFE21293251E846E8" box="[449,681,1915,1941]" italics="true" pageId="11" pageNumber="462">Ceramaster palmyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A226EA63FF96C34AFD4F293251AA46E8" box="[687,747,1915,1941]" pageId="11" pageNumber="462" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C61F089FF96C34AFCFF2932503E46E9" authorityName="Mah" authorityYear="2022" box="[799,895,1915,1941]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="vorax">
|
||
<emphasis id="B9155718FF96C34AFCFF2932503E46E9" box="[799,895,1915,1941]" italics="true" pageId="11" pageNumber="462">C. vorax</emphasis>
|
||
</taxonomicName>
|
||
, including similar abactinal plate granule shapes and identical furrow spine numbers. Among the known tropical Pacific
|
||
<taxonomicName id="4C61F089FF96C34AFC4F29D6571946C5" baseAuthorityName="Sladen" baseAuthorityYear="1889" box="[943,1112,1951,1977]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="462" phylum="Echinodermata" rank="species" species="patagonicus">
|
||
<emphasis id="B9155718FF96C34AFC4F29D6571946C5" box="[943,1112,1951,1977]" italics="true" pageId="11" pageNumber="462">C. patagonicus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54DA35A8FF96C34AFBBD29D657CC46C4" box="[1117,1165,1951,1977]" pageId="11" pageNumber="462">type</typeStatus>
|
||
species, these two show close morphological affinities.
|
||
</paragraph>
|
||
<caption id="DF1EDB82FF91C34DFF7728CB5136479F" ID-DOI="http://doi.org/10.5281/zenodo.14386790" ID-Zenodo-Dep="14386790" httpUri="https://zenodo.org/record/14386790/files/figure.png" pageId="12" pageNumber="463" startId="12.[151,250,1666,1690]" targetBox="[151,1435,181,1642]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8BDE8B0AFF91C34DFF7728CB5136479F" blockId="12.[151,1437,1666,1762]" pageId="12" pageNumber="463">
|
||
<emphasis id="B9155718FF91C34DFF7728CB517A47E7" bold="true" box="[151,571,1666,1690]" pageId="12" pageNumber="463">
|
||
FIGURE 4.
|
||
<taxonomicName id="4C61F089FF91C34DFEFE28CB52B747E7" authorityName="Mah" authorityYear="2024" box="[286,502,1666,1690]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="463" phylum="Echinodermata" rank="species" species="palmyra" status="sp. nov.">
|
||
<emphasis id="B9155718FF91C34DFEFE28CB52B747E7" bold="true" box="[286,502,1666,1690]" italics="true" pageId="12" pageNumber="463">Ceramaster palmyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A226EA63FF91C34DFE1F28CB517A47E7" box="[511,571,1666,1690]" pageId="12" pageNumber="463" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
MCZ 153216. Holotype. A. Abactinal. B. Closeup abactinal plates. C. Armtip and superomarginal plates. D. Actinal. E. Adambulacral plates with furrow and subambulacral spines. Scale bar=5.0 mm: A, B; Scale bar=1.0 mm: B, E; Scale bar=3.0 mm: C.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDE8B0AFF91C34DFF27296E502F46F4" blockId="12.[151,1436,1831,2038]" pageId="12" pageNumber="463">
|
||
Also similar, is the South African
|
||
<taxonomicName id="4C61F089FF91C34DFDA4296E5753463C" authority="H. L. Clark, 1923" authorityName="H. L. Clark" authorityYear="1923" box="[580,1042,1831,1858]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="463" phylum="Echinodermata" rank="species" species="trispinosus">
|
||
<emphasis id="B9155718FF91C34DFDA4296E500B463C" box="[580,842,1831,1857]" italics="true" pageId="12" pageNumber="463">Ceramaster trispinosus</emphasis>
|
||
H.L.
|
||
<bibRefCitation id="EFF0F6FBFF91C34DFC6B296E5753463C" author="Clark, H. L." box="[907,1042,1831,1857]" pageId="12" pageNumber="463" pagination="221 - 435" refId="ref20834" refString="Clark, H. L. (1923) The Echinoderm fauna of South Africa. Annals of the South African Museum, 13 (7), 221 - 435." type="journal article" year="1923">Clark, 1923</bibRefCitation>
|
||
</taxonomicName>
|
||
, which differs in showing 3 furrow spines versus
|
||
<quantity id="4C9926EFFF91C34DFED7290252D84618" box="[311,409,1867,1894]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.778" metricValueMin="1.016" pageId="12" pageNumber="463" unit="in" value="5.5" valueMax="7.0" valueMin="4.0">4 to 7 in</quantity>
|
||
<taxonomicName id="4C61F089FF91C34DFE42290251634618" authorityName="Mah" authorityYear="2024" box="[418,546,1867,1893]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="463" phylum="Echinodermata" rank="species" species="palmyra" status="n. sp.">
|
||
<emphasis id="B9155718FF91C34DFE42290251634618" box="[418,546,1867,1893]" italics="true" pageId="12" pageNumber="463">C. palmyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A226EA63FF91C34DFDCB29055128461B" box="[555,617,1868,1894]" pageId="12" pageNumber="463" rank="species">n. sp.</taxonomicNameLabel>
|
||
and showing a complete cover of granules on the superomarginal plate surface where
|
||
<taxonomicName id="4C61F089FF91C34DFEDA292652F946F4" authorityName="Mah" authorityYear="2024" box="[314,440,1903,1929]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="463" phylum="Echinodermata" rank="species" species="palmyra" status="n. sp.">
|
||
<emphasis id="B9155718FF91C34DFEDA292652F946F4" box="[314,440,1903,1929]" italics="true" pageId="12" pageNumber="463">C. palmyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A226EA63FF91C34DFE5F293952BA46F7" box="[447,507,1904,1930]" pageId="12" pageNumber="463" rank="species">n. sp.</taxonomicNameLabel>
|
||
displays a prominent bald patch.
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF91C34DFF2729DA5279468B" blockId="12.[151,1436,1831,2038]" pageId="12" pageNumber="463">
|
||
All of the other “
|
||
<taxonomicName id="4C61F089FF91C34DFE7329DA510146D0" baseAuthorityName="Sladen" baseAuthorityYear="1889" box="[403,576,1939,1965]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="463" phylum="Echinodermata" rank="species" species="patagonicus">
|
||
<emphasis id="B9155718FF91C34DFE7329DA510146D0" box="[403,576,1939,1965]" italics="true" pageId="12" pageNumber="463">C. patagonicus</emphasis>
|
||
</taxonomicName>
|
||
” cluster species, such as
|
||
<taxonomicName id="4C61F089FF91C34DFC8D29DA575846D0" baseAuthorityName="Sladen" baseAuthorityYear="1889" box="[877,1049,1939,1965]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="463" phylum="Echinodermata" rank="species" species="patagonicus">
|
||
<emphasis id="B9155718FF91C34DFC8D29DA575846D0" box="[877,1049,1939,1965]" italics="true" pageId="12" pageNumber="463">C. patagonicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C61F089FF91C34DFBCB29DA57E646D0" baseAuthorityName="Verrill" baseAuthorityYear="1909" box="[1067,1191,1939,1965]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="463" phylum="Echinodermata" rank="species" species="arcticus">
|
||
<emphasis id="B9155718FF91C34DFBCB29DA57E646D0" box="[1067,1191,1939,1965]" italics="true" pageId="12" pageNumber="463">C. arcticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C61F089FF91C34DFB5929DA567E46D0" authorityName="Clark & McKnight" authorityYear="2001" box="[1209,1343,1939,1965]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="463" phylum="Echinodermata" rank="species" species="australis">
|
||
<emphasis id="B9155718FF91C34DFB5929DA567E46D0" box="[1209,1343,1939,1965]" italics="true" pageId="12" pageNumber="463">C. australis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C61F089FF91C34DFA6329DA524C46AC" baseAuthorityName="Retzius" baseAuthorityYear="1783" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="463" phylum="Echinodermata" rank="species" species="granularis">
|
||
<emphasis id="B9155718FF91C34DFA6329DA524C46AC" italics="true" pageId="12" pageNumber="463">C. granularis</emphasis>
|
||
</taxonomicName>
|
||
, displayed substantially thicker, fewer and more elongate furrow and subambulacral spination than
|
||
<taxonomicName id="4C61F089FF91C34DFA6329FE53B54688" authorityName="Mah" authorityYear="2024" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="463" phylum="Echinodermata" rank="species" species="palmyra" status="n. sp.">
|
||
<emphasis id="B9155718FF91C34DFA6329FE53B54688" italics="true" pageId="12" pageNumber="463">C. palmyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A226EA63FF91C34DFF1C29955279468B" box="[252,312,2012,2038]" pageId="12" pageNumber="463" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF90C34CFF272EDE51FA41A8" blockId="13.[151,1436,151,213]" pageId="13" pageNumber="464">
|
||
Given the widely occurring distribution and relatively slight character differences between species, further molecular tests of
|
||
<taxonomicName id="4C61F089FF90C34CFE852EF2515E41A8" box="[357,543,187,213]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="13" pageNumber="464" phylum="Echinodermata" rank="species" species="undetermined">
|
||
<emphasis id="B9155718FF90C34CFE852EF252A841A8" box="[357,489,187,213]" italics="true" pageId="13" pageNumber="464">Ceramaster</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
are desirable.
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF90C34CFF772F6F5268403D" blockId="13.[151,297,294,320]" box="[151,297,294,320]" pageId="13" pageNumber="464">
|
||
<heading id="D0963C66FF90C34CFF772F6F5268403D" bold="true" box="[151,297,294,320]" fontSize="11" level="1" pageId="13" pageNumber="464" reason="1">
|
||
<emphasis id="B9155718FF90C34CFF772F6F5268403D" bold="true" box="[151,297,294,320]" pageId="13" pageNumber="464">Occurrence</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF90C34CFF772F26529A40F4" blockId="13.[151,475,367,393]" box="[151,475,367,393]" pageId="13" pageNumber="464">
|
||
Near
|
||
<collectingCountry id="F376CB9AFF90C34CFF352F26523240F4" box="[213,371,367,393]" name="United States Pacific Island Wildlife Refuges" pageId="13" pageNumber="464">Palmyra Atoll</collectingCountry>
|
||
,
|
||
<quantity id="4C9926EFFF90C34CFE9F2F26529640F4" box="[383,471,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.373" pageId="13" pageNumber="464" unit="m" value="1373.0">1373 m</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF90C34CFF772F9352694089" blockId="13.[151,296,474,500]" box="[151,296,474,500]" pageId="13" pageNumber="464">
|
||
<emphasis id="B9155718FF90C34CFF772F9352694089" bold="true" box="[151,296,474,500]" pageId="13" pageNumber="464">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF90C34CFF772C6A53AD431C" blockId="13.[151,1437,547,2013]" pageId="13" pageNumber="464">
|
||
Body stout, outline stellate (
|
||
<collectionCode id="ED7013CFFF90C34CFE312C6A52A44340" box="[465,485,547,573]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="464">R</collectionCode>
|
||
/r=2.0), arms triangular with blunt tips, armtips upturned. Interradial arcs curved(
|
||
<figureCitation id="135A978FFF90C34CFA922C6A539E431C" captionStart="FIGURE 4" captionStartId="12.[151,250,1666,1690]" captionTargetBox="[151,1435,181,1642]" captionTargetId="figure-24@12.[151,1435,181,1642]" captionTargetPageId="12" captionText="FIGURE 4. Ceramaster palmyra n. sp. MCZ 153216. Holotype. A. Abactinal. B. Closeup abactinal plates. C. Armtip and superomarginal plates. D. Actinal. E. Adambulacral plates with furrow and subambulacral spines. Scale bar=5.0 mm: A, B; Scale bar=1.0 mm: B, E; Scale bar=3.0 mm: C." figureDoi="http://doi.org/10.5281/zenodo.14386790" httpUri="https://zenodo.org/record/14386790/files/figure.png" pageId="13" pageNumber="464">Fig. 4A, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF90C34CFF272C22512B42B4" blockId="13.[151,1437,547,2013]" pageId="13" pageNumber="464">
|
||
Abactinal plates mostly hexagonal in shape, variably polygonal to round distally adjacent to superomarginal contact. Individual plates tabulate with broad base. Fasciolar grooves well developed cross all of disk surface (
|
||
<figureCitation id="135A978FFF90C34CFA8F2CC653FA43B0" captionStart="FIGURE 4" captionStartId="12.[151,250,1666,1690]" captionTargetBox="[151,1435,181,1642]" captionTargetId="figure-24@12.[151,1435,181,1642]" captionTargetPageId="12" captionText="FIGURE 4. Ceramaster palmyra n. sp. MCZ 153216. Holotype. A. Abactinal. B. Closeup abactinal plates. C. Armtip and superomarginal plates. D. Actinal. E. Adambulacral plates with furrow and subambulacral spines. Scale bar=5.0 mm: A, B; Scale bar=1.0 mm: B, E; Scale bar=3.0 mm: C." figureDoi="http://doi.org/10.5281/zenodo.14386790" httpUri="https://zenodo.org/record/14386790/files/figure.png" pageId="13" pageNumber="464">Fig. 4B</figureCitation>
|
||
). Each plate with 3 to 30 granules on surface, peripheral granules on radial disk and arm plates two to three times larger than those on central plate surface (
|
||
<figureCitation id="135A978FFF90C34CFD242C9E505E438C" box="[708,799,727,753]" captionStart="FIGURE 4" captionStartId="12.[151,250,1666,1690]" captionTargetBox="[151,1435,181,1642]" captionTargetId="figure-24@12.[151,1435,181,1642]" captionTargetPageId="12" captionText="FIGURE 4. Ceramaster palmyra n. sp. MCZ 153216. Holotype. A. Abactinal. B. Closeup abactinal plates. C. Armtip and superomarginal plates. D. Actinal. E. Adambulacral plates with furrow and subambulacral spines. Scale bar=5.0 mm: A, B; Scale bar=1.0 mm: B, E; Scale bar=3.0 mm: C." figureDoi="http://doi.org/10.5281/zenodo.14386790" httpUri="https://zenodo.org/record/14386790/files/figure.png" pageId="13" pageNumber="464">Fig. 4B</figureCitation>
|
||
). Granules on interradial plates and distally adjacent to superomarginal contact more homogeneous in size. Larger papular radial plates with 35–50 (four to ten, mostly six to eight per side), large trapezoidal peripheral granules different from other coarse round granules. Abactinal plates distributed on disk and approximately midway along arm, the remainder of which occupied by abutted superomarginal plates (
|
||
<figureCitation id="135A978FFF90C34CFE462D2E514542FC" box="[422,516,871,897]" captionStart="FIGURE 4" captionStartId="12.[151,250,1666,1690]" captionTargetBox="[151,1435,181,1642]" captionTargetId="figure-24@12.[151,1435,181,1642]" captionTargetPageId="12" captionText="FIGURE 4. Ceramaster palmyra n. sp. MCZ 153216. Holotype. A. Abactinal. B. Closeup abactinal plates. C. Armtip and superomarginal plates. D. Actinal. E. Adambulacral plates with furrow and subambulacral spines. Scale bar=5.0 mm: A, B; Scale bar=1.0 mm: B, E; Scale bar=3.0 mm: C." figureDoi="http://doi.org/10.5281/zenodo.14386790" httpUri="https://zenodo.org/record/14386790/files/figure.png" pageId="13" pageNumber="464">Fig. 4C</figureCitation>
|
||
). Interradii with closely arranged plates bearing homogeneously sized, closely distributed granules. Papulae present on radial regions, absent interradially. No pedicellariae. Madreporite quadrate in shape, flanked by four abactinal plates.
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF90C34CFF272D9A577144E0" blockId="13.[151,1437,547,2013]" pageId="13" pageNumber="464">
|
||
Superomarginals and inferomarginals, 24 (armtip to armtip), both series offset along complete series forming zigzag contact between them on lateral surface. Superomarginal plates with convex surface and strongly rounded dorsolateral edge, making plate rounded in cross-section. Superomarginal plates wide, almost completely with dorsal-facing with series forming distinct boundary around abactinal surface, occupying approximately 16% of the disk radius “r” (=0.4/2.4). Distalmost five superomarginal plates completely abutted over mid radius with sixth superomarginal plate partially abutted on all arms (
|
||
<figureCitation id="135A978FFF90C34CFD352ACE506F45DC" box="[725,814,1159,1185]" captionStart="FIGURE 4" captionStartId="12.[151,250,1666,1690]" captionTargetBox="[151,1435,181,1642]" captionTargetId="figure-24@12.[151,1435,181,1642]" captionTargetPageId="12" captionText="FIGURE 4. Ceramaster palmyra n. sp. MCZ 153216. Holotype. A. Abactinal. B. Closeup abactinal plates. C. Armtip and superomarginal plates. D. Actinal. E. Adambulacral plates with furrow and subambulacral spines. Scale bar=5.0 mm: A, B; Scale bar=1.0 mm: B, E; Scale bar=3.0 mm: C." figureDoi="http://doi.org/10.5281/zenodo.14386790" httpUri="https://zenodo.org/record/14386790/files/figure.png" pageId="13" pageNumber="464">Fig. 4C</figureCitation>
|
||
). Round abactinal surface of each superomarginal plate bald, smooth, with nearly all granules absent. Lateral side of superomarginal plates with cluster of densely packed, round granules, approximately 15–50, mostly 20–25, with largest number interradially decreasing to approximately 10–15 distally becoming nearly absent near armtip. Superomarginal plates each with 15–30 closely packed quadrate granules forming periphery around each plate. Inferomarginal plates covered by a dense pavement of closely packed round, easily abraded granules, 100–800 completely covering plate surface. Distalmost three to five inferomarginal plates with irregular polygonal to triangular raised, smooth, bald surface. No pedicellariae on either marginal series observed. Terminal plate triangular, half the size of adjacent superomarginal plate.
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF90C34CFF272BEE5745479C" blockId="13.[151,1437,547,2013]" pageId="13" pageNumber="464">
|
||
Actinal plate surface with three to four rows arranged in chevron-like series (
|
||
<figureCitation id="135A978FFF90C34CFBF22BEE572D44BC" box="[1042,1132,1447,1473]" captionStart="FIGURE 4" captionStartId="12.[151,250,1666,1690]" captionTargetBox="[151,1435,181,1642]" captionTargetId="figure-24@12.[151,1435,181,1642]" captionTargetPageId="12" captionText="FIGURE 4. Ceramaster palmyra n. sp. MCZ 153216. Holotype. A. Abactinal. B. Closeup abactinal plates. C. Armtip and superomarginal plates. D. Actinal. E. Adambulacral plates with furrow and subambulacral spines. Scale bar=5.0 mm: A, B; Scale bar=1.0 mm: B, E; Scale bar=3.0 mm: C." figureDoi="http://doi.org/10.5281/zenodo.14386790" httpUri="https://zenodo.org/record/14386790/files/figure.png" pageId="13" pageNumber="464">Fig. 4D</figureCitation>
|
||
). Individual plates quadrate to rectangular in shape with those distalmost interradially adjacent to the inferomarginal plate contact smallest and most irregular in shape. Plates covered by relatively coarse granules, approximately three count along a 1.0 mm line. Individual plates covered by granules, 8– 60 per plate, with granules differing in shape for peripheral and central plate regions with peripheral granules more quadrate versus those on central areas which are more round. All granules evenly but closely spaced. Actinal plates adjacent to the adambulacral plate series rectangular and relatively elongate compared to those occurring on the remainder of the actinal intermediate region. Granular cover becomes more homogeneous distally adjacent to the inferomarginal plate contact. Actinal granules approximately 1.5–2.0 the size of those on the inferomarginal plate surface. No pedicellariae.
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF90C34FFF2728A25655403C" blockId="13.[151,1437,547,2013]" lastBlockId="14.[151,1436,151,393]" lastPageId="14" lastPageNumber="465" pageId="13" pageNumber="464">
|
||
Adambulacral plates with furrow spines, 4 to 7, with fewest furrow spines distally, two series of subambulacral thick spines, each with three spines per series and a series of granules, 2 to 4 which are similar in appearance to those on actinal intermediate plate surface (
|
||
<figureCitation id="135A978FFF90C34CFD6F297A51A84630" box="[655,745,1843,1869]" captionStart="FIGURE 4" captionStartId="12.[151,250,1666,1690]" captionTargetBox="[151,1435,181,1642]" captionTargetId="figure-24@12.[151,1435,181,1642]" captionTargetPageId="12" captionText="FIGURE 4. Ceramaster palmyra n. sp. MCZ 153216. Holotype. A. Abactinal. B. Closeup abactinal plates. C. Armtip and superomarginal plates. D. Actinal. E. Adambulacral plates with furrow and subambulacral spines. Scale bar=5.0 mm: A, B; Scale bar=1.0 mm: B, E; Scale bar=3.0 mm: C." figureDoi="http://doi.org/10.5281/zenodo.14386790" httpUri="https://zenodo.org/record/14386790/files/figure.png" pageId="13" pageNumber="464">Fig. 4E</figureCitation>
|
||
). Furrow spines with blunt, rounded tips, quadrate in cross-section, palmate arrangement with distinct space setting them apart from subambulacral spines. Subambulacral spines, three, each spine quadrate to polygonal in cross-section, approximately twice as thick as an individual furrow spine, approximately 70% of the height of the furrow spines. Third subambulacral series, three to four, short, approximately 50% of height of adjacent subambulacral spine with blunt to pointed tip. similar in height to actinal granules. Oral plates with 9 to 10 furrow spines, with two projecting into mouth per interradius. Oral spines triangular to quadrate in cross-section, with blunt or pointed tips. Oral plate surface with two or three short thick spines, triangular to polygonal in cross-section, similar in appearance to subambulacral spines on adambulacral plates. Fossae between two halves of the oral plate with short, thick spines, similar to the second and third series of subambulacral spines, polygonal to triangular in cross-section. Spines on oral plates all widely spaced.
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF93C34FFF272F02515140F4" blockId="14.[151,1436,151,393]" pageId="14" pageNumber="465">
|
||
Color of living individuals was yellow with white highlights along the disk inter radii and around the superomarginal plates (
|
||
<figureCitation id="135A978FFF93C34FFE7D2F26514240F4" box="[413,515,367,393]" captionStart="FIGURE 12" captionStartId="33.[151,250,1651,1675]" captionTargetBox="[151,1436,181,1627]" captionTargetId="figure-15@33.[151,1436,181,1627]" captionTargetPageId="33" captionText="FIGURE 12. Ahuastra and Ceramaster palmyra observed in situ. A. Ahuastra kupanaha nov. gen. nov. sp. in situ, showing apparent feeding. B. Ceramaster palmyra n. sp. in situ." figureDoi="http://doi.org/10.5281/zenodo.14386822" httpUri="https://zenodo.org/record/14386822/files/figure.png" pageId="14" pageNumber="465">Fig. 12B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37BD881FF93C34FFF772F93519F431C" pageId="14" pageNumber="465" type="materials_examined">
|
||
<paragraph id="8BDE8B0AFF93C34FFF772F9352CA4089" blockId="14.[151,395,474,500]" box="[151,395,474,500]" pageId="14" pageNumber="465">
|
||
<heading id="D0963C66FF93C34FFF772F9352CA4089" bold="true" box="[151,395,474,500]" fontSize="11" level="1" pageId="14" pageNumber="465" reason="1">
|
||
<emphasis id="B9155718FF93C34FFF772F9352CA4089" bold="true" box="[151,395,474,500]" pageId="14" pageNumber="465">Material Examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDE8B0AFF93C34FFF772C6A519F431C" blockId="14.[151,1436,546,609]" pageId="14" pageNumber="465">
|
||
<materialsCitation id="3B098157FF93C34FFF772C6A519B431C" collectingDate="2019-06-30" collectionCode="MCZ" collectorName="E / V Nautilus" latitude="5.620168" location="Palmyra Atoll, South Pacific Ocean" longitude="-161.74697" pageId="14" pageNumber="465" specimenCode="MCZ 153216" typeStatus="holotype">
|
||
<typeStatus id="54DA35A8FF93C34FFF772C6A52424340" box="[151,259,547,573]" pageId="14" pageNumber="465" type="holotype">Holotype</typeStatus>
|
||
.
|
||
<specimenCode id="DBC72371FF93C34FFEF22C6A52EF4340" box="[274,430,546,573]" pageId="14" pageNumber="465">MCZ 153216</specimenCode>
|
||
Near
|
||
<location id="8EBEDDD1FF93C34FFE1B2C6A50DE4340" LSID="urn:lsid:plazi:treatment:03C83A1CFF96C34FFF772CC6519F431C:8EBEDDD1FF93C34FFE1B2C6A50DE4340" box="[507,927,547,573]" name="Palmyra Atoll, South Pacific Ocean" pageId="14" pageNumber="465">Palmyra Atoll, South Pacific Ocean</location>
|
||
,
|
||
<geoCoordinate id="EE55EDCDFF93C34FFC4F2C6B575A4340" box="[943,1051,546,573]" orientation="latitude" pageId="14" pageNumber="465" value="5.620168">5.620168</geoCoordinate>
|
||
°
|
||
<geoCoordinate id="EE55EDCDFF93C34FFBCF2C6A57F04340" box="[1071,1201,547,573]" orientation="longitude" pageId="14" pageNumber="465" value="-161.74697">-161.74697</geoCoordinate>
|
||
°
|
||
<specimenCount id="9D674083FF93C34FFB212C6A56634340" box="[1217,1314,547,573]" pageId="14" pageNumber="465" type="male">
|
||
<quantity id="4C9926EFFF93C34FFB212C6A565F4340" box="[1217,1310,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.373" pageId="14" pageNumber="465" unit="m" value="1373.0">1373 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Coll.
|
||
<collectorName id="2694EEDCFF93C34FFA902C6A53B7431C" pageId="14" pageNumber="465">
|
||
E/V
|
||
<taxonomicName id="4C61F089FF93C34FFF772C0E53B7431C" box="[151,246,583,609]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="14" pageNumber="465" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9155718FF93C34FFF772C0E53B7431C" box="[151,246,583,609]" italics="true" pageId="14" pageNumber="465">Nautilus</emphasis>
|
||
</taxonomicName>
|
||
</collectorName>
|
||
<date id="FFDFADCAFF93C34FFF1D2C0E52D4431C" box="[253,405,583,609]" pageId="14" pageNumber="465" value="2019-06-30">
|
||
<collectingDate id="EF9B5422FF93C34FFF1D2C0E52D4431C" box="[253,405,583,609]" pageId="14" pageNumber="465" value="2019-06-30">30 June 2019</collectingDate>
|
||
</date>
|
||
, 1 wet spec. R=4.8 r=
|
||
<quantity id="4C9926EFFF93C34FFD6A2C0E519B431C" box="[650,730,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="14" pageNumber="465" unit="cm" value="2.4">2.4 cm</quantity>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |