treatments-xml/data/E9/26/E6/E926E62F434C541C88253CA249C0E706.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1059.70085" ID-GBIF-Dataset="0398ea8b-fbff-4395-a8fd-7458c85d25bd" ID-PMC="PMC8448723" ID-Pensoft-Pub="1313-2970-1059-135" ID-Pensoft-UUID="B9B0ABCC937A5E868C43F703F7DDE872" ID-PubMed="34611455" ID-ZooBank="1AC2998758F74FC3BFC91C4C7C90F2A0" ModsDocID="1313-2970-1059-135" checkinTime="1631288215658" checkinUser="pensoft" docAuthor="Pohjoismaeki, Jaakko &amp; Haarto, Antti" docDate="2021" docId="E926E62F434C541C88253CA249C0E706" docLanguage="en" docName="ZooKeys 1059: 135-156" docOrigin="ZooKeys 1059" docPubDate="2021-09-10" docSource="http://dx.doi.org/10.3897/zookeys.1059.70085" docTitle="Scenopinus jerei Pohjoismäki &amp; Haarto 2021, sp. nov." docType="treatment" docUuid="7BBF06EE-AA55-42A8-B2DF-88FC4640AFA0" docUuidSource="ZooBank" docVersion="4" id="B9B0ABCC937A5E868C43F703F7DDE872" lastPageNumber="135" masterDocId="B9B0ABCC937A5E868C43F703F7DDE872" masterDocTitle="Scenopinus jerei, a new species of window fly (Diptera, Scenopinidae) from Finland" masterLastPageNumber="156" masterPageNumber="135" pageNumber="135" updateTime="1668150889829" updateUser="ExternalLinkService">
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<mods:title>Scenopinus jerei, a new species of window fly (Diptera, Scenopinidae) from Finland</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pohjoismaeki, Jaakko</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1185-3610</mods:nameIdentifier>
<mods:affiliation>University of Eastern Finland, Department of Biology, P. O. Box 111, FI- 80101 Joensuu, Finland</mods:affiliation>
<mods:nameIdentifier type="email">jaakko.pohjoismaki@uef.fi</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Haarto, Antti</mods:namePart>
<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014 Turku, Finland</mods:affiliation>
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<mods:date>2021</mods:date>
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<mods:number>2021-09-10</mods:number>
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<mods:number>1059</mods:number>
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<mods:start>135</mods:start>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1059.70085</mods:identifier>
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<treatment ID-GBIF-Taxon="187732824" LSID="urn:lsid:zoobank.org:act:7BBF06EE-AA55-42A8-B2DF-88FC4640AFA0" httpUri="http://treatment.plazi.org/id/E926E62F434C541C88253CA249C0E706" lastPageNumber="135" pageId="0" pageNumber="135">
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<paragraph pageId="0" pageNumber="135">
<taxonomicName LSID="http://zoobank.org/7BBF06EE-AA55-42A8-B2DF-88FC4640AFA0" authority="Pohjoismäki &amp; Haarto, 2021" authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei" status="sp. nov.">Scenopinus jerei</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="135">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="description">
<paragraph pageId="0" pageNumber="135">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of northern European Scenopinus fenestralis group species of Scenopinidae A holotype male of Scenopinus jerei sp. nov., Kouvola, Finland B paratype female of Scenopinus jerei sp. nov., Kouvola, Finland C male of Scenopinus fenestralis, Liperi, Finland D female of Scenopinus fenestralis, Liperi, Finland E male of Scenopinus vitripennis, Wuerttburg, Germany. Photograph by Susanne Leidenroth F female of Scenopinus vitripennis, Warszawa, Poland. Photograph by D. Schimrosczyk G female of Scenopinus glabrifrons, Ockstadt, Germany. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure1" httpUri="https://binary.pensoft.net/fig/586907" pageId="0" pageNumber="135">Figures 1A, B</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male heads of Scenopinus species A lateral view of holotype Scenopinus jerei sp. nov., male head B frontal view of holotype Scenopinus jerei sp. nov., head C lateral view of Scenopinus fenestralis male head D frontal view of Scenopinus fenestralis male head. For the illustrations of male heads of S. vitripennis and S. glabrifrons, see Krivosheina (1981). Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure2" httpUri="https://binary.pensoft.net/fig/586908" pageId="0" pageNumber="135">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Female heads of Scenopinus species A lateral view of paratype Scenopinus jerei sp. nov., female head B frontal view of paratype Scenopinus jerei sp. nov., head C lateral view of Scenopinus fenestralis female head D frontal view of Scenopinus fenestralis female head E l ateral view of Scenopinus vitripennis female head. Photograph D. Schimrosczyk F frontal view of Scenopinus vitripennis female head. Photograph D. Schimrosczyk G lateral view of Scenopinus glabrifrons female head H frontal view of Scenopinus glabrifrons female head. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure3" httpUri="https://binary.pensoft.net/fig/586909" pageId="0" pageNumber="135">, 3A, B</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scenopinus male terminalia A hypandrium of Scenopinus jerei sp. nov. paratype B epandrium of Scenopinus jerei sp. nov. paratype C hypandrium of Scenopinus fenestralis D epandrium of Scenopinus fenestralis E terminal segments of Scenopinus jerei sp. nov. paratype F aedeagus of Scenopinus jerei sp. nov. paratype G terminal segments of Scenopinus fenestralis H aedeagus of Scenopinus fenestralis. aed - aedeagus; aed lb - aedeagal lobe; distph - distiphallus; ej apod - ejaculatory apodeme; epand - epandrium; goncx apod - gonocoxal apodeme; gonst - gonostylym; hypand - hypandrium; pm sh - parameral sheath. For the illustrations of male terminalia of S. vitripennis and S. glabrifrons, see Krivosheina (1981)." figureDoi="10.3897/zookeys.1059.70085.figure4" httpUri="https://binary.pensoft.net/fig/586910" pageId="0" pageNumber="135">, 4A, B</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scenopinus male terminalia A hypandrium of Scenopinus jerei sp. nov. paratype B epandrium of Scenopinus jerei sp. nov. paratype C hypandrium of Scenopinus fenestralis D epandrium of Scenopinus fenestralis E terminal segments of Scenopinus jerei sp. nov. paratype F aedeagus of Scenopinus jerei sp. nov. paratype G terminal segments of Scenopinus fenestralis H aedeagus of Scenopinus fenestralis. aed - aedeagus; aed lb - aedeagal lobe; distph - distiphallus; ej apod - ejaculatory apodeme; epand - epandrium; goncx apod - gonocoxal apodeme; gonst - gonostylym; hypand - hypandrium; pm sh - parameral sheath. For the illustrations of male terminalia of S. vitripennis and S. glabrifrons, see Krivosheina (1981)." figureDoi="10.3897/zookeys.1059.70085.figure4" httpUri="https://binary.pensoft.net/fig/586910" pageId="0" pageNumber="135">, 4E, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="type material">
<paragraph pageId="0" pageNumber="135">Type material.</paragraph>
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Holotype</emphasis>
(1♂): Finland, Sa: Kouvola, 674-679:347-350 [60.7686-61.2184, 26.4495-27.0000] / e.l. 2018 ex 
<taxonomicName authorityName="Linnaeus" authorityYear="1758" class="Aves" family="Strigidae" genus="Strix" kingdom="Animalia" lsidName="Strix aluco" order="Strigiformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="aluco">
<emphasis italics="true" pageId="0" pageNumber="135">Strix aluco</emphasis>
</taxonomicName>
nest box. / M. Mutanen leg. // 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
&amp; Haarto 2021 / (
<taxonomicName authorityName="Latreille" authorityYear="1802" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" rank="order">Diptera</taxonomicName>
: 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
) / J. 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
det. // HOLOTYPE [red label] [MZH] 
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Paratypes</emphasis>
: 1♂ (dissected, DNA barcoded JP2020-S1), 1♀, same collection data; // 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
&amp; Haarto 2021 / (
<taxonomicName authorityName="Latreille" authorityYear="1802" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" rank="order">Diptera</taxonomicName>
: 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
) / J. 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
det. // PARATYPE [yellow label] [MZH]; 1♂: Finland, EP: 
<normalizedToken originalValue="Isokyrö">Isokyroe</normalizedToken>
/ Orisberg 6983:[3]265 [
<geoCoordinate degrees="62.8744" direction="north" orientation="latitude" precision="5" value="62.8744">62.8744</geoCoordinate>
, 
<geoCoordinate degrees="22.3795" direction="east" orientation="longitude" precision="5" value="22.3795">22.3795</geoCoordinate>
]/ 7.7.1999/ A. Haarto leg.// 
<taxonomicName class="Insecta" family="Scenopinidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="family">SCENOPINIDAE</taxonomicName>
/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
/ 
<taxonomicName authorityName="Meigen" authorityYear="1824" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus vitripennis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="vitripennis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus vitripennis</emphasis>
</taxonomicName>
Meig./ det. A. Haarto 1999// PARATYPE/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" rank="order">Diptera</taxonomicName>
: 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
/ 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
/ 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
&amp; Haarto 2021 [red label] [AHC]; 1♂: Finland, ES [
<normalizedToken originalValue="Etelä-Savo">Etelae-Savo</normalizedToken>
]: Rantasalmi/ Korhola 68720:[3]5802 [
<geoCoordinate degrees="61.9458" direction="north" orientation="latitude" precision="5" value="61.9458">61.9458</geoCoordinate>
, 
<geoCoordinate degrees="28.5278" direction="east" orientation="longitude" precision="5" value="28.5278">28.5278</geoCoordinate>
] / 27.6.2006/ A. Haarto leg.// 
<taxonomicName class="Insecta" family="Scenopinidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="family">SCENOPINIDAE</taxonomicName>
/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinus</taxonomicName>
/ sp./ det. A. Haarto 2006// PARATYPE/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" rank="order">Diptera</taxonomicName>
: 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
/ 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
/ 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
&amp; Haarto 2021 [red label] [AHC]; 2♂: Finland, Kb: Ilomantsi/ Kelovaara 70008:36826 [
<geoCoordinate degrees="63.0638" direction="north" orientation="latitude" precision="5" value="63.0638">63.0638</geoCoordinate>
, 
<geoCoordinate degrees="30.6144" direction="east" orientation="longitude" precision="5" value="30.6144">30.6144</geoCoordinate>
]/ 24.7.2021/ J. 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
leg. // PARATYPE/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" rank="order">Diptera</taxonomicName>
: 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
/ 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
/ 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
&amp; Haarto 2021 [yellow label] [JPC]; 1♀: Finland, ES: Rantasalmi/ Korhola 68720:5802 [
<geoCoordinate degrees="61.9458" direction="north" orientation="latitude" precision="5" value="61.9458">61.9458</geoCoordinate>
, 
<geoCoordinate degrees="28.5278" direction="east" orientation="longitude" precision="5" value="28.5278">28.5278</geoCoordinate>
]/ 29.6.2006/ A. Haarto leg.// 
<taxonomicName class="Insecta" family="Scenopinidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="family">SCENOPINIDAE</taxonomicName>
/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinus</taxonomicName>
/ sp./ det. A. Haarto 2006// PARATYPE/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" rank="order">Diptera</taxonomicName>
: 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
/ 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
/ 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
&amp; Haarto 2021 [red label] [AHC]; 1♀: Finland, Kb: Liperi/ 
<normalizedToken originalValue="Viinijärvi">Viinijaervi</normalizedToken>
6951:3615 [
<geoCoordinate degrees="62.6451" direction="north" orientation="latitude" precision="5" value="62.6451">62.6451</geoCoordinate>
, 
<geoCoordinate degrees="29.2425" direction="east" orientation="longitude" precision="5" value="29.2425">29.2425</geoCoordinate>
] / e larva 2013/ Ali Karhu leg.// 
<normalizedToken originalValue="linnunpönttö">linnunpoenttoe</normalizedToken>
[nest box]// 
<taxonomicName class="Insecta" family="Scenopinidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="family">SCENOPINIDAE</taxonomicName>
/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
/ sp./ det. A. Haarto 2014/ AHa14-000891// PARATYPE/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" rank="order">Diptera</taxonomicName>
: 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
/ 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
/ 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
&amp; Haarto 2021 [red label] [AHC]; 1♀: Finland, Kb: Liperi/ 
<normalizedToken originalValue="Käsämä">Kaesaemae</normalizedToken>
suo 6950:3619 [
<geoCoordinate degrees="62.6349" direction="north" orientation="latitude" precision="5" value="62.6349">62.6349</geoCoordinate>
, 
<geoCoordinate degrees="29.3197" direction="east" orientation="longitude" precision="5" value="29.3197">29.3197</geoCoordinate>
]/ 26.-28.6.2013/ Ali Karhu leg.// 
<taxonomicName class="Insecta" family="Scenopinidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="family">SCENOPINIDAE</taxonomicName>
/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
/ sp./ det. A. Haarto 2020/ AHa20-000473// PARATYPE/ 
<taxonomicName authorityName="Latreille" authorityYear="1802" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" rank="order">Diptera</taxonomicName>
: 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
/ 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
/ 
<normalizedToken originalValue="Pohjoismäki">Pohjoismaeki</normalizedToken>
&amp; Haarto 2021 [red label] [MZT].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="diagnosis">
<paragraph pageId="0" pageNumber="135">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="135">
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. belongs to the 
<taxonomicName lsidName="S. fenestralis" pageId="0" pageNumber="135" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">S. fenestralis</emphasis>
</taxonomicName>
group and is easily recognisable from the other species in this group based on the contrasting colour differences between the femora and the yellow to orange tibiae. The coxae as well as the knob of the halteres are always uniformly black or dark brown, similar to the colour of the thorax.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="description">
<paragraph pageId="0" pageNumber="135">Description.</paragraph>
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" pageId="0" pageNumber="135">Male</emphasis>
(Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of northern European Scenopinus fenestralis group species of Scenopinidae A holotype male of Scenopinus jerei sp. nov., Kouvola, Finland B paratype female of Scenopinus jerei sp. nov., Kouvola, Finland C male of Scenopinus fenestralis, Liperi, Finland D female of Scenopinus fenestralis, Liperi, Finland E male of Scenopinus vitripennis, Wuerttburg, Germany. Photograph by Susanne Leidenroth F female of Scenopinus vitripennis, Warszawa, Poland. Photograph by D. Schimrosczyk G female of Scenopinus glabrifrons, Ockstadt, Germany. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure1" httpUri="https://binary.pensoft.net/fig/586907" pageId="0" pageNumber="135">1A</figureCitation>
; 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male heads of Scenopinus species A lateral view of holotype Scenopinus jerei sp. nov., male head B frontal view of holotype Scenopinus jerei sp. nov., head C lateral view of Scenopinus fenestralis male head D frontal view of Scenopinus fenestralis male head. For the illustrations of male heads of S. vitripennis and S. glabrifrons, see Krivosheina (1981). Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure2" httpUri="https://binary.pensoft.net/fig/586908" pageId="0" pageNumber="135">2A, B</figureCitation>
; 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scenopinus male terminalia A hypandrium of Scenopinus jerei sp. nov. paratype B epandrium of Scenopinus jerei sp. nov. paratype C hypandrium of Scenopinus fenestralis D epandrium of Scenopinus fenestralis E terminal segments of Scenopinus jerei sp. nov. paratype F aedeagus of Scenopinus jerei sp. nov. paratype G terminal segments of Scenopinus fenestralis H aedeagus of Scenopinus fenestralis. aed - aedeagus; aed lb - aedeagal lobe; distph - distiphallus; ej apod - ejaculatory apodeme; epand - epandrium; goncx apod - gonocoxal apodeme; gonst - gonostylym; hypand - hypandrium; pm sh - parameral sheath. For the illustrations of male terminalia of S. vitripennis and S. glabrifrons, see Krivosheina (1981)." figureDoi="10.3897/zookeys.1059.70085.figure4" httpUri="https://binary.pensoft.net/fig/586910" pageId="0" pageNumber="135">4A, B, F, G</figureCitation>
) (characters in square brackets refer to the holotype). Body length: [4.1]-4.9 mm (n = 6) [dried specimens, fresh specimens are longer).
</paragraph>
<caption doi="10.3897/zookeys.1059.70085.figure1" httpUri="https://binary.pensoft.net/fig/586907" pageId="0" pageNumber="135" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" pageId="0" pageNumber="135">Figure 1.</emphasis>
Habitus of northern European 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
group species of 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="135">A</emphasis>
holotype male of 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov., Kouvola, Finland 
<emphasis bold="true" pageId="0" pageNumber="135">B</emphasis>
paratype female of 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov., Kouvola, Finland 
<emphasis bold="true" pageId="0" pageNumber="135">C</emphasis>
male of 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
, Liperi, Finland 
<emphasis bold="true" pageId="0" pageNumber="135">D</emphasis>
female of 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
, Liperi, Finland 
<emphasis bold="true" pageId="0" pageNumber="135">E</emphasis>
male of 
<taxonomicName authorityName="Meigen" authorityYear="1824" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus vitripennis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="vitripennis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus vitripennis</emphasis>
</taxonomicName>
, 
<normalizedToken originalValue="Württburg">Wuerttburg</normalizedToken>
, Germany. Photograph by Susanne Leidenroth 
<emphasis bold="true" pageId="0" pageNumber="135">F</emphasis>
female of 
<taxonomicName authorityName="Meigen" authorityYear="1824" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus vitripennis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="vitripennis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus vitripennis</emphasis>
</taxonomicName>
, Warszawa, Poland. Photograph by D. Schimrosczyk 
<emphasis bold="true" pageId="0" pageNumber="135">G</emphasis>
female of 
<taxonomicName authorityName="Meigen" authorityYear="1824" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus glabrifrons" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="glabrifrons">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus glabrifrons</emphasis>
</taxonomicName>
, Ockstadt, Germany. Scale bar: 500 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Head</emphasis>
(Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male heads of Scenopinus species A lateral view of holotype Scenopinus jerei sp. nov., male head B frontal view of holotype Scenopinus jerei sp. nov., head C lateral view of Scenopinus fenestralis male head D frontal view of Scenopinus fenestralis male head. For the illustrations of male heads of S. vitripennis and S. glabrifrons, see Krivosheina (1981). Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure2" httpUri="https://binary.pensoft.net/fig/586908" pageId="0" pageNumber="135">2A, B</figureCitation>
). Black with greasy-looking shine, including oral margin and occiput, apart from weak grey microtomentum around occipital foramen, mouth edge and antennal base; semi-circular, height 1.5-1.7 [1.6] (n = 6) 
<normalizedToken originalValue="×">x</normalizedToken>
its maximum width in lateral view. Antennal insertion slightly below mid eye level. Antenna dark brown with pedicel and anterior part of flagellomere paler; scape short and subrectangular; pedicel short and cylindrical, [0.8]-1.0 (n = 6) 
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, flagellomere laterally flattened, 1.8[1.9]-2.0 
<normalizedToken originalValue="×">x</normalizedToken>
as long as high, subrectangular, narrowing apically and 4.8-[5.4] (n = 6) 
<normalizedToken originalValue="×">x</normalizedToken>
as long as pedicel and with subcircular, subapical, sensory pore on outer side. Eyes large and bare; fronto-orbital plates meeting at [0.25]-0.3 (n = 6) length of frons; no frontal vitta; gena reduced to narrow strip between lower eye margin and mouth edge. Diameter of ommatidia on upper half of compound eye, above antennal base, 2-3 
<normalizedToken originalValue="×">x</normalizedToken>
diameter of ommatidia on lower half. Ocellar triangle acute, distance between posterior ocelli distinctly shorter than their distance to anterior ocellus. Frons bare but patterned with minute pits. No setae or setulae on head, apart for short brown setulae at lower posterior part of gena behind mouth edge. Mouthparts, including palpus, black.
</paragraph>
<caption doi="10.3897/zookeys.1059.70085.figure2" httpUri="https://binary.pensoft.net/fig/586908" pageId="0" pageNumber="135" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" pageId="0" pageNumber="135">Figure 2.</emphasis>
Male heads of 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
species 
<emphasis bold="true" pageId="0" pageNumber="135">A</emphasis>
lateral view of holotype 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov., male head 
<emphasis bold="true" pageId="0" pageNumber="135">B</emphasis>
frontal view of holotype 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov., head 
<emphasis bold="true" pageId="0" pageNumber="135">C</emphasis>
lateral view of 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
male head 
<emphasis bold="true" pageId="0" pageNumber="135">D</emphasis>
frontal view of 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
male head. For the illustrations of male heads of 
<taxonomicName lsidName="S. vitripennis" pageId="0" pageNumber="135" rank="species" species="vitripennis">
<emphasis italics="true" pageId="0" pageNumber="135">S. vitripennis</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="S. glabrifrons" pageId="0" pageNumber="135" rank="species" species="glabrifrons">
<emphasis italics="true" pageId="0" pageNumber="135">S. glabrifrons</emphasis>
</taxonomicName>
, see 
<bibRefCitation author="Krivosheina, NP" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="135" pagination="24 - 31" refId="B15" refString="Krivosheina, NP, 1981. Taxonomy of fenestralis group species of the genus Scenopinus Latr. (Diptera, Scenopinidae). Vestnik Zoologii 4: 24 - 31" title="Taxonomy of fenestralis group species of the genus Scenopinus Latr. (Diptera, Scenopinidae)." volume="4" year="1981">Krivosheina (1981)</bibRefCitation>
. Scale bar: 500 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Thorax</emphasis>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of northern European Scenopinus fenestralis group species of Scenopinidae A holotype male of Scenopinus jerei sp. nov., Kouvola, Finland B paratype female of Scenopinus jerei sp. nov., Kouvola, Finland C male of Scenopinus fenestralis, Liperi, Finland D female of Scenopinus fenestralis, Liperi, Finland E male of Scenopinus vitripennis, Wuerttburg, Germany. Photograph by Susanne Leidenroth F female of Scenopinus vitripennis, Warszawa, Poland. Photograph by D. Schimrosczyk G female of Scenopinus glabrifrons, Ockstadt, Germany. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure1" httpUri="https://binary.pensoft.net/fig/586907" pageId="0" pageNumber="135">1A</figureCitation>
). Dorsally and laterally black with greasy-looking shine. Scutum patterned with small rugae and minute, barely distinguishable setulae. Pleura with similar patterning but with more distinct, short, sparse, greyish to brownish setulae. Hirsuteness most developed on anepisternum, where longest setulae are approximately same length as width of flagellomere.
</paragraph>
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Legs</emphasis>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of northern European Scenopinus fenestralis group species of Scenopinidae A holotype male of Scenopinus jerei sp. nov., Kouvola, Finland B paratype female of Scenopinus jerei sp. nov., Kouvola, Finland C male of Scenopinus fenestralis, Liperi, Finland D female of Scenopinus fenestralis, Liperi, Finland E male of Scenopinus vitripennis, Wuerttburg, Germany. Photograph by Susanne Leidenroth F female of Scenopinus vitripennis, Warszawa, Poland. Photograph by D. Schimrosczyk G female of Scenopinus glabrifrons, Ockstadt, Germany. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure1" httpUri="https://binary.pensoft.net/fig/586907" pageId="0" pageNumber="135">1A</figureCitation>
). Coxae black; femora brown and apically paler. Fore and mid tibiae [pale brown] to dirty orange, clearly paler than femora. Hind tibia otherwise of similar colour as femora but paler at base and apex. Hind coxa with thin black posterior setulae, longest setulae as long as width of coxa at its base. Femora with thin posterodorsal setulae, longest being 0.5 
<normalizedToken originalValue="×">x</normalizedToken>
as long as width of femora. Fore tibia preapically with [2]-3 ventral setulae. Mid tibia with two short, ventral, preapical setae and two adjacent setulae. Hind tibia with one ventral preapical setula and thin posteroventral setulae covering proximal half, longest of which are as long as width of tibia. Apart from aforementioned setae and setulae, all legs covered in minute setulae that provide rugous texture.
</paragraph>
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Wings</emphasis>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of northern European Scenopinus fenestralis group species of Scenopinidae A holotype male of Scenopinus jerei sp. nov., Kouvola, Finland B paratype female of Scenopinus jerei sp. nov., Kouvola, Finland C male of Scenopinus fenestralis, Liperi, Finland D female of Scenopinus fenestralis, Liperi, Finland E male of Scenopinus vitripennis, Wuerttburg, Germany. Photograph by Susanne Leidenroth F female of Scenopinus vitripennis, Warszawa, Poland. Photograph by D. Schimrosczyk G female of Scenopinus glabrifrons, Ockstadt, Germany. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure1" httpUri="https://binary.pensoft.net/fig/586907" pageId="0" pageNumber="135">1A</figureCitation>
). Hyaline with greyish tinge. Tegula black, basicosta brownish black, wing veins brown. Petiole and knob of haltere uniformly black.
</paragraph>
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Abdomen</emphasis>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of northern European Scenopinus fenestralis group species of Scenopinidae A holotype male of Scenopinus jerei sp. nov., Kouvola, Finland B paratype female of Scenopinus jerei sp. nov., Kouvola, Finland C male of Scenopinus fenestralis, Liperi, Finland D female of Scenopinus fenestralis, Liperi, Finland E male of Scenopinus vitripennis, Wuerttburg, Germany. Photograph by Susanne Leidenroth F female of Scenopinus vitripennis, Warszawa, Poland. Photograph by D. Schimrosczyk G female of Scenopinus glabrifrons, Ockstadt, Germany. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure1" httpUri="https://binary.pensoft.net/fig/586907" pageId="0" pageNumber="135">1A</figureCitation>
). Elongated, dorsally flattened except for domed terminal segments, black in colour, with greasy shine and covered by irregular, minute, robust setulae. Tergites 6, 7, and 8 with black marginal setulae, longest of which ca. as long as width of hind femora.
</paragraph>
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Terminalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scenopinus male terminalia A hypandrium of Scenopinus jerei sp. nov. paratype B epandrium of Scenopinus jerei sp. nov. paratype C hypandrium of Scenopinus fenestralis D epandrium of Scenopinus fenestralis E terminal segments of Scenopinus jerei sp. nov. paratype F aedeagus of Scenopinus jerei sp. nov. paratype G terminal segments of Scenopinus fenestralis H aedeagus of Scenopinus fenestralis. aed - aedeagus; aed lb - aedeagal lobe; distph - distiphallus; ej apod - ejaculatory apodeme; epand - epandrium; goncx apod - gonocoxal apodeme; gonst - gonostylym; hypand - hypandrium; pm sh - parameral sheath. For the illustrations of male terminalia of S. vitripennis and S. glabrifrons, see Krivosheina (1981)." figureDoi="10.3897/zookeys.1059.70085.figure4" httpUri="https://binary.pensoft.net/fig/586910" pageId="0" pageNumber="135">4A, B, E, F</figureCitation>
). Hypandrium divided to two similar subrectangular halves, posterior margin with long dense setulae (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scenopinus male terminalia A hypandrium of Scenopinus jerei sp. nov. paratype B epandrium of Scenopinus jerei sp. nov. paratype C hypandrium of Scenopinus fenestralis D epandrium of Scenopinus fenestralis E terminal segments of Scenopinus jerei sp. nov. paratype F aedeagus of Scenopinus jerei sp. nov. paratype G terminal segments of Scenopinus fenestralis H aedeagus of Scenopinus fenestralis. aed - aedeagus; aed lb - aedeagal lobe; distph - distiphallus; ej apod - ejaculatory apodeme; epand - epandrium; goncx apod - gonocoxal apodeme; gonst - gonostylym; hypand - hypandrium; pm sh - parameral sheath. For the illustrations of male terminalia of S. vitripennis and S. glabrifrons, see Krivosheina (1981)." figureDoi="10.3897/zookeys.1059.70085.figure4" httpUri="https://binary.pensoft.net/fig/586910" pageId="0" pageNumber="135">4A</figureCitation>
). Epandrium similarly divided into subtriangular halves with long posteroventral marginal setulae (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scenopinus male terminalia A hypandrium of Scenopinus jerei sp. nov. paratype B epandrium of Scenopinus jerei sp. nov. paratype C hypandrium of Scenopinus fenestralis D epandrium of Scenopinus fenestralis E terminal segments of Scenopinus jerei sp. nov. paratype F aedeagus of Scenopinus jerei sp. nov. paratype G terminal segments of Scenopinus fenestralis H aedeagus of Scenopinus fenestralis. aed - aedeagus; aed lb - aedeagal lobe; distph - distiphallus; ej apod - ejaculatory apodeme; epand - epandrium; goncx apod - gonocoxal apodeme; gonst - gonostylym; hypand - hypandrium; pm sh - parameral sheath. For the illustrations of male terminalia of S. vitripennis and S. glabrifrons, see Krivosheina (1981)." figureDoi="10.3897/zookeys.1059.70085.figure4" httpUri="https://binary.pensoft.net/fig/586910" pageId="0" pageNumber="135">4B</figureCitation>
). Cercus subtriangular (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scenopinus male terminalia A hypandrium of Scenopinus jerei sp. nov. paratype B epandrium of Scenopinus jerei sp. nov. paratype C hypandrium of Scenopinus fenestralis D epandrium of Scenopinus fenestralis E terminal segments of Scenopinus jerei sp. nov. paratype F aedeagus of Scenopinus jerei sp. nov. paratype G terminal segments of Scenopinus fenestralis H aedeagus of Scenopinus fenestralis. aed - aedeagus; aed lb - aedeagal lobe; distph - distiphallus; ej apod - ejaculatory apodeme; epand - epandrium; goncx apod - gonocoxal apodeme; gonst - gonostylym; hypand - hypandrium; pm sh - parameral sheath. For the illustrations of male terminalia of S. vitripennis and S. glabrifrons, see Krivosheina (1981)." figureDoi="10.3897/zookeys.1059.70085.figure4" httpUri="https://binary.pensoft.net/fig/586910" pageId="0" pageNumber="135">4E</figureCitation>
); gonocoxal apodeme scapula-shaped and curving slightly inwards at its posterior part; gonostylym bluntly subrectangular, posteriorly concave and ending with a ventral apex. Distiphallus with long, narrow, curved apical processes, hooking outwards prior to apex (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scenopinus male terminalia A hypandrium of Scenopinus jerei sp. nov. paratype B epandrium of Scenopinus jerei sp. nov. paratype C hypandrium of Scenopinus fenestralis D epandrium of Scenopinus fenestralis E terminal segments of Scenopinus jerei sp. nov. paratype F aedeagus of Scenopinus jerei sp. nov. paratype G terminal segments of Scenopinus fenestralis H aedeagus of Scenopinus fenestralis. aed - aedeagus; aed lb - aedeagal lobe; distph - distiphallus; ej apod - ejaculatory apodeme; epand - epandrium; goncx apod - gonocoxal apodeme; gonst - gonostylym; hypand - hypandrium; pm sh - parameral sheath. For the illustrations of male terminalia of S. vitripennis and S. glabrifrons, see Krivosheina (1981)." figureDoi="10.3897/zookeys.1059.70085.figure4" httpUri="https://binary.pensoft.net/fig/586910" pageId="0" pageNumber="135">4F</figureCitation>
). Aedeagus rod-shaped, slightly bent at middle and with a short-forked apex.
</paragraph>
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" pageId="0" pageNumber="135">Female</emphasis>
(Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of northern European Scenopinus fenestralis group species of Scenopinidae A holotype male of Scenopinus jerei sp. nov., Kouvola, Finland B paratype female of Scenopinus jerei sp. nov., Kouvola, Finland C male of Scenopinus fenestralis, Liperi, Finland D female of Scenopinus fenestralis, Liperi, Finland E male of Scenopinus vitripennis, Wuerttburg, Germany. Photograph by Susanne Leidenroth F female of Scenopinus vitripennis, Warszawa, Poland. Photograph by D. Schimrosczyk G female of Scenopinus glabrifrons, Ockstadt, Germany. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure1" httpUri="https://binary.pensoft.net/fig/586907" pageId="0" pageNumber="135">1B</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Female heads of Scenopinus species A lateral view of paratype Scenopinus jerei sp. nov., female head B frontal view of paratype Scenopinus jerei sp. nov., head C lateral view of Scenopinus fenestralis female head D frontal view of Scenopinus fenestralis female head E l ateral view of Scenopinus vitripennis female head. Photograph D. Schimrosczyk F frontal view of Scenopinus vitripennis female head. Photograph D. Schimrosczyk G lateral view of Scenopinus glabrifrons female head H frontal view of Scenopinus glabrifrons female head. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure3" httpUri="https://binary.pensoft.net/fig/586909" pageId="0" pageNumber="135">3A, B</figureCitation>
). Differs from male as follows:
</paragraph>
<paragraph pageId="0" pageNumber="135">
Body length: 4.3-5.7 mm (n = 4). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Head</emphasis>
(Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Female heads of Scenopinus species A lateral view of paratype Scenopinus jerei sp. nov., female head B frontal view of paratype Scenopinus jerei sp. nov., head C lateral view of Scenopinus fenestralis female head D frontal view of Scenopinus fenestralis female head E l ateral view of Scenopinus vitripennis female head. Photograph D. Schimrosczyk F frontal view of Scenopinus vitripennis female head. Photograph D. Schimrosczyk G lateral view of Scenopinus glabrifrons female head H frontal view of Scenopinus glabrifrons female head. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure3" httpUri="https://binary.pensoft.net/fig/586909" pageId="0" pageNumber="135">3A, B</figureCitation>
). Frons broad, at its narrowest point 0.62-0.69 (n = 4) 
<normalizedToken originalValue="×">x</normalizedToken>
as wide as an eye in dorsal view. Frons shiny black with minute longitudinal rugae on frontal stripe and transverse rugae on sides. Orbital plates smooth and shiny. Ocellar triangle equilateral. No obvious size difference between ommatidia of upper and lower half of compound eye. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Thorax</emphasis>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of northern European Scenopinus fenestralis group species of Scenopinidae A holotype male of Scenopinus jerei sp. nov., Kouvola, Finland B paratype female of Scenopinus jerei sp. nov., Kouvola, Finland C male of Scenopinus fenestralis, Liperi, Finland D female of Scenopinus fenestralis, Liperi, Finland E male of Scenopinus vitripennis, Wuerttburg, Germany. Photograph by Susanne Leidenroth F female of Scenopinus vitripennis, Warszawa, Poland. Photograph by D. Schimrosczyk G female of Scenopinus glabrifrons, Ockstadt, Germany. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure1" httpUri="https://binary.pensoft.net/fig/586907" pageId="0" pageNumber="135">1B</figureCitation>
). Very weak whitish grey microtomentum at anterior parts of postpronotum and proepisternum. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Abdomen</emphasis>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of northern European Scenopinus fenestralis group species of Scenopinidae A holotype male of Scenopinus jerei sp. nov., Kouvola, Finland B paratype female of Scenopinus jerei sp. nov., Kouvola, Finland C male of Scenopinus fenestralis, Liperi, Finland D female of Scenopinus fenestralis, Liperi, Finland E male of Scenopinus vitripennis, Wuerttburg, Germany. Photograph by Susanne Leidenroth F female of Scenopinus vitripennis, Warszawa, Poland. Photograph by D. Schimrosczyk G female of Scenopinus glabrifrons, Ockstadt, Germany. Scale bar: 500 mm." figureDoi="10.3897/zookeys.1059.70085.figure1" httpUri="https://binary.pensoft.net/fig/586907" pageId="0" pageNumber="135">1B</figureCitation>
). Dorsally flattened along its entire length. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">Terminalia</emphasis>
. Last visible tergite 9 bluntly triangular at its posterior edge and not divided into hemitergites.
</paragraph>
<caption doi="10.3897/zookeys.1059.70085.figure3" httpUri="https://binary.pensoft.net/fig/586909" pageId="0" pageNumber="135" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" pageId="0" pageNumber="135">Figure 3.</emphasis>
Female heads of 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
species 
<emphasis bold="true" pageId="0" pageNumber="135">A</emphasis>
lateral view of paratype 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov., female head 
<emphasis bold="true" pageId="0" pageNumber="135">B</emphasis>
frontal view of paratype 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov., head 
<emphasis bold="true" pageId="0" pageNumber="135">C</emphasis>
lateral view of 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
female head 
<emphasis bold="true" pageId="0" pageNumber="135">D</emphasis>
frontal view of 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
female head 
<emphasis bold="true" pageId="0" pageNumber="135">E</emphasis>
l ateral view of 
<taxonomicName authorityName="Meigen" authorityYear="1824" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus vitripennis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="vitripennis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus vitripennis</emphasis>
</taxonomicName>
female head. Photograph D. Schimrosczyk 
<emphasis bold="true" pageId="0" pageNumber="135">F</emphasis>
frontal view of 
<taxonomicName authorityName="Meigen" authorityYear="1824" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus vitripennis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="vitripennis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus vitripennis</emphasis>
</taxonomicName>
female head. Photograph D. Schimrosczyk 
<emphasis bold="true" pageId="0" pageNumber="135">G</emphasis>
lateral view of 
<taxonomicName authorityName="Meigen" authorityYear="1824" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus glabrifrons" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="glabrifrons">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus glabrifrons</emphasis>
</taxonomicName>
female head 
<emphasis bold="true" pageId="0" pageNumber="135">H</emphasis>
frontal view of 
<taxonomicName authorityName="Meigen" authorityYear="1824" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus glabrifrons" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="glabrifrons">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus glabrifrons</emphasis>
</taxonomicName>
female head. Scale bar: 500 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1059.70085.figure4" httpUri="https://binary.pensoft.net/fig/586910" pageId="0" pageNumber="135" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" pageId="0" pageNumber="135">Figure 4.</emphasis>
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
male terminalia 
<emphasis bold="true" pageId="0" pageNumber="135">A</emphasis>
hypandrium of 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. paratype 
<emphasis bold="true" pageId="0" pageNumber="135">B</emphasis>
epandrium of 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. paratype 
<emphasis bold="true" pageId="0" pageNumber="135">C</emphasis>
hypandrium of 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="135">D</emphasis>
epandrium of 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="135">E</emphasis>
terminal segments of 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. paratype 
<emphasis bold="true" pageId="0" pageNumber="135">F</emphasis>
aedeagus of 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. paratype 
<emphasis bold="true" pageId="0" pageNumber="135">G</emphasis>
terminal segments of 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="135">H</emphasis>
aedeagus of 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
. 
<emphasis italics="true" pageId="0" pageNumber="135">aed</emphasis>
- aedeagus; 
<emphasis italics="true" pageId="0" pageNumber="135">aed lb</emphasis>
- aedeagal lobe; 
<emphasis italics="true" pageId="0" pageNumber="135">distph</emphasis>
- distiphallus; 
<emphasis italics="true" pageId="0" pageNumber="135">ej apod</emphasis>
- ejaculatory apodeme; 
<emphasis italics="true" pageId="0" pageNumber="135">epand</emphasis>
- epandrium; 
<emphasis italics="true" pageId="0" pageNumber="135">goncx apod</emphasis>
- gonocoxal apodeme; gonst - gonostylym; 
<emphasis italics="true" pageId="0" pageNumber="135">hypand</emphasis>
- hypandrium; 
<emphasis italics="true" pageId="0" pageNumber="135">pm sh</emphasis>
- parameral sheath. For the illustrations of male terminalia of 
<taxonomicName lsidName="S. vitripennis" pageId="0" pageNumber="135" rank="species" species="vitripennis">
<emphasis italics="true" pageId="0" pageNumber="135">S. vitripennis</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="S. glabrifrons" pageId="0" pageNumber="135" rank="species" species="glabrifrons">
<emphasis italics="true" pageId="0" pageNumber="135">S. glabrifrons</emphasis>
</taxonomicName>
, see 
<bibRefCitation author="Krivosheina, NP" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="135" pagination="24 - 31" refId="B15" refString="Krivosheina, NP, 1981. Taxonomy of fenestralis group species of the genus Scenopinus Latr. (Diptera, Scenopinidae). Vestnik Zoologii 4: 24 - 31" title="Taxonomy of fenestralis group species of the genus Scenopinus Latr. (Diptera, Scenopinidae)." volume="4" year="1981">Krivosheina (1981)</bibRefCitation>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="dna barcode divergence among scenopinus">
<paragraph pageId="0" pageNumber="135">
DNA barcode divergence among 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="135">
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
spp. are poorly covered in the DNA barcode databases, such as Barcode of Life Database (BOLD, www.boldsystems.org) or GenBank. It is noteworthy that all 
<taxonomicName lsidName="S. fenestralis" pageId="0" pageNumber="135" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">S. fenestralis</emphasis>
</taxonomicName>
specimens in the databases from Europe to North America have almost identical 
<emphasis italics="true" pageId="0" pageNumber="135">COI</emphasis>
sequences and represent the same barcode index number (BIN). The DNA barcode of 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. differs markedly from the other northern European species, its closest match being 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus fenestralis" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus fenestralis</emphasis>
</taxonomicName>
from which it is separated by 12.48% sequence difference (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Maximum likelihood tree of COI sequence similarities among Scenopinus spp. Numbers at nodes indicate posterior probabilities and scale bar the relative sequence divergence. BOLD sample ID number is given after the species name." figureDoi="10.3897/zookeys.1059.70085.figure5" httpUri="https://binary.pensoft.net/fig/586911" pageId="0" pageNumber="135">5</figureCitation>
). There are no other closer matches among the barcode sequences in the BOLD or GenBank.
</paragraph>
<caption doi="10.3897/zookeys.1059.70085.figure5" httpUri="https://binary.pensoft.net/fig/586911" pageId="0" pageNumber="135" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="135">
<emphasis bold="true" pageId="0" pageNumber="135">Figure 5.</emphasis>
Maximum likelihood tree of 
<emphasis italics="true" pageId="0" pageNumber="135">COI</emphasis>
sequence similarities among 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
spp. Numbers at nodes indicate posterior probabilities and scale bar the relative sequence divergence. BOLD sample ID number is given after the species name.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="notes">
<paragraph pageId="0" pageNumber="135">Notes</paragraph>
<paragraph pageId="0" pageNumber="135">
on the biology and distribution of 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. The larvae of 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
species are predators of other invertebrates living in dry organic substrates, such as in animal nests. In Finland, 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. has been collected inside sheds, attics and indoor storages as well as reared from nest boxes of birds. These rearings produced large numbers of tineid moths (
<taxonomicName authorityName="Latreille" authorityYear="1802" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="135" rank="order">Lepidoptera</taxonomicName>
: Tineidea), especially 
<taxonomicName class="Insecta" family="Tineidae" genus="Monopis" kingdom="Animalia" lsidName="Monopis laevigella" order="Lepidoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="laevigella">
<emphasis italics="true" pageId="0" pageNumber="135">Monopis laevigella</emphasis>
</taxonomicName>
(Denis &amp; 
<normalizedToken originalValue="Schiffermüller">Schiffermueller</normalizedToken>
), but also other 
<taxonomicName class="Insecta" family="Tineidae" genus="Monopis" kingdom="Animalia" lsidName="Monopis" order="Lepidoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Monopis</emphasis>
</taxonomicName>
spp., 
<taxonomicName baseAuthorityName="Matsumura" baseAuthorityYear="1931" class="Insecta" family="Tineidae" genus="Niditinea" kingdom="Animalia" lsidName="Niditinea striolella" order="Lepidoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="striolella">
<emphasis italics="true" pageId="0" pageNumber="135">Niditinea striolella</emphasis>
</taxonomicName>
(Matsumura), and 
<taxonomicName class="Insecta" family="Tineidae" genus="Tinea" kingdom="Animalia" lsidName="Tinea" order="Lepidoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Tinea</emphasis>
</taxonomicName>
spp. Other insects observed from the same nest boxes included 
<taxonomicName class="Insecta" family="Ceratophyllidae" genus="Ceratophyllus" kingdom="Animalia" lsidName="Ceratophyllus" order="Siphonaptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Ceratophyllus</emphasis>
</taxonomicName>
fleas, various beetles (
<taxonomicName authorityName="Gyllenhal" authorityYear="1808" class="Insecta" family="Histeridae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
, 
<taxonomicName class="Insecta" family="Dermestidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="family">Dermestidae</taxonomicName>
) and flies (
<taxonomicName authorityName="Macquart" authorityYear="1835" class="Insecta" family="Piophilidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="family">Piophilidae</taxonomicName>
, 
<taxonomicName class="Insecta" family="Fanniidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="family">Fanniidae</taxonomicName>
, 
<taxonomicName class="Insecta" family="Heleomyzidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="family">Heleomyzidae</taxonomicName>
). Apart for two male specimens found dead on a windowsill in an attic of an old house in Kelovaara on July 24 (see type specimens), most observations are from third week of June. According to the observations of Jere 
<normalizedToken originalValue="Kahanpää">Kahanpaeae</normalizedToken>
(pers. comm.), 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. hibernates as full-grown larvae and the adults emerge in a couple of weeks in room temperature rearing conditions. Based on the collection locations, it is likely that 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. is a boreal forest specialist.
</paragraph>
<paragraph pageId="0" pageNumber="135">
Like other 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus</emphasis>
</taxonomicName>
spp., 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. is not very active flier, does not visit flowers and therefore is rarely collected by active netting or traps. Judging from the few Finnish observations, the species appears widespread in the southern and central parts of the country. We are certain that 
<taxonomicName authorityName="Pohjoismäki &amp; Haarto" authorityYear="2021" class="Insecta" family="Scenopinidae" genus="Scenopinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scenopinus jerei" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="jerei">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinus jerei</emphasis>
</taxonomicName>
sp. nov. can also be found in boreal forest biotopes in the other Nordic countries and Russia but has been until now overlooked.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="etymology">
<paragraph pageId="0" pageNumber="135">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="135">
This species is named after Mr. Jere 
<normalizedToken originalValue="Kahanpää">Kahanpaeae</normalizedToken>
, Helsinki, who was to first to discover that the taxon is new to science and kindly agreed with the current arrangement for its formal description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="provisional key to the identification of european scenopinidae species">
<paragraph pageId="0" pageNumber="135">
Provisional key to the identification of European 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="135">Scenopinidae</emphasis>
</taxonomicName>
species
</paragraph>
<paragraph pageId="0" pageNumber="135">
Because the existing literature on the European species of 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
are outdated or difficult to obtain, we felt necessary to provide a key for the known European species of 
<taxonomicName class="Insecta" family="Scenopinidae" genus="Scenopinidae" kingdom="Animalia" lsidName="Scenopinidae" order="Diptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Scenopinidae</taxonomicName>
. We must emphasise that we have been only able to examine the species with specimens listed in this paper, for which the identification key should work well. For the remainder, our approach was to go through the written species descriptions and pick features which we judged, by our collective species identification experience, to be useful for determination. To us this approach was better justified than reproducing the keys given in earlier literature, which are often difficult to follow or focus on limited number of poorly defined features. The diagnostic features for the key have been obtained from the descriptions in 
<bibRefCitation author="Kroeber, O" journalOrPublisher="Zoologicheskii Zhurnal" pageId="0" pageNumber="135" refId="B17" refString="Kroeber, O, 1925. 27. Omphralidae (Scenopinidae). In: Lindner E (Ed.) Die Fliegen der Palaearktischen Region IV (Lieferung 123): 1-8." title="27. Omphralidae (Scenopinidae). In: Lindner E (Ed.) Die Fliegen der Palaearktischen Region IV (Lieferung 123): 1 - 8." year="1925">
<normalizedToken originalValue="Kröber">Kroeber</normalizedToken>
(1925)
</bibRefCitation>
, 
<bibRefCitation author="Trojan, P" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="135" pagination="147 - 156" refId="B39" refString="Trojan, P, 1956. Notes on the taxonomy of some European species of the genus Omphrale Meigen (Diptera, Omphralidae). Annales Zoologici 16: 147 - 156" title="Notes on the taxonomy of some European species of the genus Omphrale Meigen (Diptera, Omphralidae)." volume="16" year="1956">Trojan (1956)</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.16405" author="Kelsey, LP" journalOrPublisher="Bulletin of the United States National Museum" pageId="0" pageNumber="135" pagination="1 - 336" refId="B13" refString="Kelsey, LP, 1969. A revision of the Scenopinidae (Diptera) of the world. Bulletin of the United States National Museum 277: 1 - 336, DOI: https://doi.org/10.5962/bhl.title.16405" title="A revision of the Scenopinidae (Diptera) of the world." url="https://doi.org/10.5962/bhl.title.16405" volume="277" year="1969">Kelsey (1969)</bibRefCitation>
, 
<bibRefCitation author="Krivosheina, NP" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="135" pagination="24 - 31" refId="B15" refString="Krivosheina, NP, 1981. Taxonomy of fenestralis group species of the genus Scenopinus Latr. (Diptera, Scenopinidae). Vestnik Zoologii 4: 24 - 31" title="Taxonomy of fenestralis group species of the genus Scenopinus Latr. (Diptera, Scenopinidae)." volume="4" year="1981">Krivosheina (1981)</bibRefCitation>
, 
<bibRefCitation author="Narchuk, EP" editor="Bei-Bienko, GI" journalOrPublisher="Volume V. Diptera and Siphonaptera. Part I. Amerind Publishing Co. Pvt. Ltd. New Delhi" pageId="0" pageNumber="135" pagination="838 - 842" refId="B22" refString="Narchuk, EP, 1988. 42. Familiy Scenopinidae (Omphralidae). In: Bei-Bienko, GI, Ed., Keys to the insects of the European USSR. Volume V. Diptera and Siphonaptera. Part I. Amerind Publishing Co. Pvt. Ltd. New Delhi: 838 - 842" title="42. Familiy Scenopinidae (Omphralidae)." volumeTitle="Keys to the insects of the European USSR." year="1988">Narchuk (1988)</bibRefCitation>
, and 
<bibRefCitation author="Carles-Tolra, M" journalOrPublisher="Boletin de la Asociacion Espanola de Entomologia" pageId="0" pageNumber="135" pagination="35 - 41" refId="B4" refString="Carles-Tolra, M, 2001. Two new species of Scenopinus Latreille from Spain (Diptera, Scenopinidae). Boletin de la Asociacion Espanola de Entomologia 25: 35 - 41" title="Two new species of Scenopinus Latreille from Spain (Diptera, Scenopinidae)." volume="25" year="2001">
<normalizedToken originalValue="Carles-Tolrá">Carles-Tolra</normalizedToken>
(2001)
</bibRefCitation>
. Fortunately, the European species separate into three easily recognisable species groups, each with relatively few species. The species groups appear in the key in alphabetical order, enabling fast navigation when one is familiar with the groups. Although result appears satisfactory, the key might not capture all the variations seen within each species and we strongly encourage DNA barcoding of specimens for future reference. In any case, we hope that this key can form a basis for forthcoming work with this interesting family of flies.
</paragraph>
</subSubSection>
</treatment>
</document>