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<document id="446F2E13B4600D5EAD6022F3E72E5194" ID-CLB-Dataset="27706" ID-DOI="10.11646/zootaxa.4543.2.2" ID-GBIF-Dataset="7438332f-a925-43aa-b17f-059781bdfc12" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617757" ID-ZooBank="11F546CD-109F-4CBC-B87D-EA8D1AD0B96F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928261775" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos &amp; Cerdeño, Esperanza" docDate="2019" docId="B0798797FFBC286E72CBCAA9FE4BF8DA" docLanguage="en" docName="zootaxa.4543.2.2.pdf" docOrigin="Zootaxa 4543 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Interatherium anguliferum Ameghino 1894" docType="treatment" docVersion="8" lastPageNumber="201" masterDocId="4C40FFEFFFBA2868725CCE7FFF8EFFA2" masterDocTitle="On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina" masterLastPageNumber="220" masterPageNumber="195" pageNumber="201" updateTime="1701997211094" updateUser="plazi">
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<mods:title id="AEC2E4E21A182AA84B4D83D96846DE61">On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina</mods:title>
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<mods:namePart id="8E92479EAF368D4ADE9BC4AE56D9835F">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="02AEAFC550D06657ACD0A26D31159998">Fernicola, Juan Carlos</mods:namePart>
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<mods:namePart id="95DDF4496CF60D1B1F9E8CB96ADDB564">Cerdeño, Esperanza</mods:namePart>
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<treatment id="B0798797FFBC286E72CBCAA9FE4BF8DA" ID-DOI="http://doi.org/10.5281/zenodo.5931273" ID-GBIF-Taxon="154529753" ID-Zenodo-Dep="5931273" LSID="urn:lsid:plazi:treatment:B0798797FFBC286E72CBCAA9FE4BF8DA" httpUri="http://treatment.plazi.org/id/B0798797FFBC286E72CBCAA9FE4BF8DA" lastPageNumber="201" pageId="6" pageNumber="201">
<subSubSection id="70CA650AFFBC286E72CBCAA9FD4EFB53" box="[151,704,1238,1265]" pageId="6" pageNumber="201" type="nomenclature">
<paragraph id="386F3681FFBC286E72CBCAA9FD4EFB53" blockId="6.[151,704,1238,1265]" box="[151,704,1238,1265]" pageId="6" pageNumber="201">
<heading id="632781EDFFBC286E72CBCAA9FD4EFB53" bold="true" box="[151,704,1238,1265]" fontSize="11" level="1" pageId="6" pageNumber="201" reason="1">
<taxonomicName id="FFD04D02FFBC286E72CBCAA9FD4EFB53" authority="Ameghino, 1894 a" authorityName="Ameghino" authorityYear="1894" box="[151,704,1238,1265]" class="Mammalia" family="Interatheriidae" genus="Interatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="201" phylum="Chordata" rank="species" species="anguliferum">
<emphasis id="0AA4EA93FFBC286E72CBCAA9FD4EFB53" bold="true" box="[151,704,1238,1265]" pageId="6" pageNumber="201">
<emphasis id="0AA4EA93FFBC286E72CBCAA9FE6EFB52" bold="true" box="[151,480,1238,1264]" italics="true" pageId="6" pageNumber="201">Interatherium anguliferum</emphasis>
<bibRefCitation id="5C414B70FFBC286E73B4CAA8FD4EFB53" author="Ameghino, F." box="[488,704,1239,1265]" pageId="6" pageNumber="201" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino, 1894a</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="70CA650AFFBC286E72CBCB62FB32F966" pageId="6" pageNumber="201" type="materials_examined">
<paragraph id="386F3681FFBC286E72CBCB62FB16FA94" blockId="6.[151,1436,1309,1913]" box="[151,1176,1309,1334]" pageId="6" pageNumber="201">
<materialsCitation id="88B83CDCFFBC286E72CBCB62FB16FA94" ID-GBIF-Occurrence="2147525043" box="[151,1176,1309,1334]" collectionCode="MACN-A" pageId="6" pageNumber="201" specimenCode="MACN-A 3444" specimenCount="1" typeStatus="holotype">
<emphasis id="0AA4EA93FFBC286E72CBCB62FE85FA94" bold="true" box="[151,267,1309,1334]" pageId="6" pageNumber="201">
<typeStatus id="E76B8823FFBC286E72CBCB62FE88FA94" box="[151,262,1309,1334]" pageId="6" pageNumber="201" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="68769EFAFFBC286E734ECB61FE4CFA94" box="[274,450,1310,1334]" collectionCode="MACN-A" pageId="6" pageNumber="201">MACN-A 3444</specimenCode>
(
<figureCitation id="A0EB2A04FFBC286E738ECB62FDBEFA94" box="[466,560,1309,1334]" captionStart="FIGURE 1" captionStartId="4.[151,249,1523,1545]" captionTargetBox="[163,1427,492,1485]" captionTargetId="figure@4.[150,1436,484,1502]" captionTargetPageId="4" captionText="FIGURE 1. Interatherium rodens, holotype MLP 12-2826, right maxilla with alveolus of dP1 and complete dP2 and P3M3, in occlusal view (A); reproduction of figure 26 (B) from Ameghino (1889, plate 15). Interatherium anguliferum, holotype MACN-A 3444, right maxilla with C and P2M1, in occlusal view (C). Interatherium interruptum, holotype MACN-A 3445, left maxilla with the alveolus of dP1 and complete dP23 and P4M3, in occlusal view (D). Interatherium dentatum, holotype MACN-A 9863, skull and mandible articulated, left lateral view (E). Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617759/files/figure.png" pageId="6" pageNumber="201">Fig. 1 C</figureCitation>
), a fragmented right maxilla with C and P2M1 series.
</materialsCitation>
</paragraph>
<paragraph id="386F3681FFBC286E729BCB3EFEAFFADC" blockId="6.[151,1436,1309,1913]" pageId="6" pageNumber="201">
<emphasis id="0AA4EA93FFBC286E729BCB3EFD5DFAF8" bold="true" box="[199,723,1345,1370]" pageId="6" pageNumber="201">
Age and provenance of the
<materialsCitation id="88B83CDCFFBC286E707BCB3EFD5DFAF8" ID-GBIF-Occurrence="2147524941" box="[551,723,1345,1370]" collectionCode="SALMA" country="Argentina" location="Miocene" pageId="6" pageNumber="201" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<typeStatus id="E76B8823FFBC286E707BCB3EFDD4FAF8" box="[551,602,1345,1370]" pageId="6" pageNumber="201">type</typeStatus>
material.
</materialsCitation>
</emphasis>
Santacrucian SALMA, early
<location id="3D0F605AFFBC286E766ACB3EFB15FAF8" LSID="urn:lsid:plazi:treatment:B0798797FFBC286E72CBCAA9FE4BF8DA:3D0F605AFFBC286E766ACB3EFB15FAF8" box="[1078,1179,1345,1370]" country="Argentina" name="Miocene" pageId="6" pageNumber="201" stateProvince="Santa Cruz">Miocene</location>
,
<collectingRegion id="FA14F863FFBC286E76F7CB3DFA12FAF8" box="[1195,1436,1345,1370]" country="Argentina" name="Santa Cruz" pageId="6" pageNumber="201">Santa Cruz Province</collectingRegion>
(
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).
</paragraph>
<paragraph id="386F3681FFBC286E729BCBF6FD96F997" blockId="6.[151,1436,1309,1913]" pageId="6" pageNumber="201">
<emphasis id="0AA4EA93FFBC286E729BCBF6FE36FA00" bold="true" box="[199,440,1417,1442]" pageId="6" pageNumber="201">Original description</emphasis>
(
<bibRefCitation id="5C414B70FFBC286E739BCBF6FD36FA00" author="Ameghino, F." box="[455,696,1417,1442]" pageId="6" pageNumber="201" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino 1894a: 18</bibRefCitation>
). “
<emphasis id="0AA4EA93FFBC286E7085CBF4FB1DFA00" box="[729,1171,1417,1442]" italics="true" pageId="6" pageNumber="201">De taille intermédiaire entre celle de 1</emphasis>
<taxonomicName id="FFD04D02FFBC286E76CFCBF5FB7BFA00" authorityName="Moreno" authorityYear="1882" box="[1171,1269,1417,1442]" class="Mammalia" family="Interatheriidae" genus="Interatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="201" phylum="Chordata" rank="species" species="rodens">I. rodens</taxonomicName>
<emphasis id="0AA4EA93FFBC286E76A0CBF4FA05FA00" box="[1276,1419,1417,1442]" italics="true" pageId="6" pageNumber="201">et celle de 1</emphasis>
<taxonomicName id="FFD04D02FFBC286E77D0CBF5FE8BFA67" authorityName="Ameghino" authorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Interatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="201" phylum="Chordata" rank="species" species="supernum">I. supernum</taxonomicName>
<emphasis id="0AA4EA93FFBC286E7350CBD2FC2CFA4F" italics="true" pageId="6" pageNumber="201">dont elle se distingue par l'absence de la première prémolaire supérieure. La canine supérieure est bien développée et isolée par des diastèmesen avant et en arrière. Les p.</emphasis>
<emphasis id="0AA4EA93FFBC286E71FECBAEFC4FFA4E" box="[930,961,1489,1516]" italics="true" pageId="6" pageNumber="201">
<superScript id="CFA59BC9FFBC286E71FECBAEFC24FA7D" attach="left" box="[930,938,1489,1503]" fontSize="6" pageId="6" pageNumber="201">2</superScript>
à
</emphasis>
<emphasis id="0AA4EA93FFBC286E7196CBAEFD8AF997" italics="true" pageId="6" pageNumber="201">
<superScript id="CFA59BC9FFBC286E7196CBAEFC5CFA7D" attach="none" box="[970,978,1489,1503]" fontSize="6" pageId="6" pageNumber="201">4</superScript>
ont les deux arêtes perpendiculaires de l'angle antérieur externe très fortes. Longueur du bord antérieur de la canine au bord postérieur de la première vraie molaire supérieure,
<quantity id="FF289B64FFBC286E73E5C862FD8AF997" box="[441,516,1565,1589]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="201" unit="mm" value="17.0">17 mm</quantity>
</emphasis>
”.
</paragraph>
<paragraph id="386F3681FFBC286E729BC83FFB32F966" blockId="6.[151,1436,1309,1913]" pageId="6" pageNumber="201">
English translation. Size intermediate between
<taxonomicName id="FFD04D02FFBC286E7089C83EFCB6F9FB" authorityName="Moreno" authorityYear="1882" box="[725,824,1600,1625]" class="Mammalia" family="Interatheriidae" genus="Interatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="201" phylum="Chordata" rank="species" species="rodens">
<emphasis id="0AA4EA93FFBC286E7089C83EFCB6F9FB" box="[725,824,1600,1625]" italics="true" pageId="6" pageNumber="201">I. rodens</emphasis>
</taxonomicName>
and
<taxonomicName id="FFD04D02FFBC286E7132C83EFC7DF9FA" authorityName="Ameghino" authorityYear="1887" box="[878,1011,1601,1624]" class="Mammalia" family="Interatheriidae" genus="Interatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="201" phylum="Chordata" rank="species" species="supernum">
<emphasis id="0AA4EA93FFBC286E7132C83EFC7DF9FA" box="[878,1011,1601,1624]" italics="true" pageId="6" pageNumber="201">I. supernum</emphasis>
</taxonomicName>
, from which it is distinguished by the absence of the first upper premolar. The upper canine is well developed and anteriorly and posteriorly isolated by diastemata. P24 have the two perpendicular columns of the antero-external angle very strong. Length from the anterior region of the upper canine to the posterior border of the first true upper molar,
<quantity id="FF289B64FFBC286E7639C8D2FB39F966" box="[1125,1207,1709,1733]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="201" unit="mm" value="17.0">17 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="70CA650AFFBC286E729BC8AFFE4BF8DA" pageId="6" pageNumber="201" type="discussion">
<paragraph id="386F3681FFBC286E729BC8AFFE4BF8DA" blockId="6.[151,1436,1309,1913]" pageId="6" pageNumber="201">
<emphasis id="0AA4EA93FFBC286E729BC8AFFEC1F94A" bold="true" box="[199,335,1744,1769]" pageId="6" pageNumber="201">Comments.</emphasis>
<bibRefCitation id="5C414B70FFBC286E7307C8AFFDBAF94B" author="Ameghino, F." box="[347,564,1744,1769]" pageId="6" pageNumber="201" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
did not illustrate any specimen of this species. According to Ameghinos catalogue and
<bibRefCitation id="5C414B70FFBC286E731FC88AFE67F8AF" author="Mones A." box="[323,489,1780,1805]" pageId="6" pageNumber="201" pagination="1 - 625" refId="ref17293" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article" year="1986">Mones (1986)</bibRefCitation>
, the maxilla MACN-A 3444 is the type specimen of the species. This material matches the original description provided by
<bibRefCitation id="5C414B70FFBC286E70F8C967FCF5F893" author="Ameghino, F." box="[676,891,1816,1841]" pageId="6" pageNumber="201" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
because its dP1 is absent, there are diastemata anterior and posterior to C, and the series CM1 is
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long. Therefore, MACN-A 3444 is confirmed as the
<typeStatus id="E76B8823FFBC286E72CBC91FFF77F8DB" box="[151,249,1888,1913]" pageId="6" pageNumber="201" type="holotype">holotype</typeStatus>
of
<taxonomicName id="FFD04D02FFBC286E7342C91EFE30F8DB" authorityName="Ameghino" authorityYear="1894" box="[286,446,1888,1913]" class="Mammalia" family="Interatheriidae" genus="Interatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="201" phylum="Chordata" rank="species" species="anguliferum">
<emphasis id="0AA4EA93FFBC286E7342C91EFE30F8DB" box="[286,446,1888,1913]" italics="true" pageId="6" pageNumber="201">I. anguliferum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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