172 lines
18 KiB
XML
172 lines
18 KiB
XML
<document id="73FCA0656F13FA77ED808204E1C6E442" ID-CLB-Dataset="27706" ID-DOI="10.11646/zootaxa.4543.2.2" ID-GBIF-Dataset="7438332f-a925-43aa-b17f-059781bdfc12" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617757" ID-ZooBank="11F546CD-109F-4CBC-B87D-EA8D1AD0B96F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928261775" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos & Cerdeño, Esperanza" docDate="2019" docId="B0798797FFB5287872CBCB3CFEB7FE25" docLanguage="en" docName="zootaxa.4543.2.2.pdf" docOrigin="Zootaxa 4543 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Icochilus lamellosus Ameghino 1894" docType="treatment" docVersion="8" lastPageNumber="210" masterDocId="4C40FFEFFFBA2868725CCE7FFF8EFFA2" masterDocTitle="On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina" masterLastPageNumber="220" masterPageNumber="195" pageNumber="210" updateTime="1701997211094" updateUser="plazi">
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<mods:title id="AD01174F994B0BC1F022CF30199DCA1D">On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina</mods:title>
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<mods:namePart id="39F81BCAEB7A8173CE7C8A26BA7E5514">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="4625ECCECE1E35F7858356A9254CFDAD">Fernicola, Juan Carlos</mods:namePart>
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<mods:namePart id="13FE505B08E09B5E3B82F89A41CEF24C">Cerdeño, Esperanza</mods:namePart>
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<mods:date id="F180E88B649F857F292524D4681CBCE8">2019</mods:date>
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<treatment id="B0798797FFB5287872CBCB3CFEB7FE25" ID-DOI="http://doi.org/10.5281/zenodo.5931297" ID-GBIF-Taxon="154529759" ID-Zenodo-Dep="5931297" LSID="urn:lsid:plazi:treatment:B0798797FFB5287872CBCB3CFEB7FE25" httpUri="http://treatment.plazi.org/id/B0798797FFB5287872CBCB3CFEB7FE25" lastPageId="16" lastPageNumber="210" pageId="15" pageNumber="210">
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<subSubSection id="70CA650AFFB5286772CBCB3CFDE1FAFC" box="[151,623,1347,1374]" pageId="15" pageNumber="210" type="nomenclature">
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<paragraph id="386F3681FFB5286772CBCB3CFDE1FAFC" blockId="15.[151,623,1347,1374]" box="[151,623,1347,1374]" pageId="15" pageNumber="210">
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<heading id="632781EDFFB5286772CBCB3CFDE1FAFC" bold="true" box="[151,623,1347,1374]" fontSize="11" level="1" pageId="15" pageNumber="210" reason="1">
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<taxonomicName id="FFD04D02FFB5286772CBCB3CFDE1FAFC" authority="Ameghino, 1894 a" authorityName="Ameghino" authorityYear="1894" box="[151,623,1347,1374]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="lamellosus">
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<emphasis id="0AA4EA93FFB5286772CBCB3CFDE1FAFC" bold="true" box="[151,623,1347,1374]" pageId="15" pageNumber="210">
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<emphasis id="0AA4EA93FFB5286772CBCB3CFE01FAFF" bold="true" box="[151,399,1347,1373]" italics="true" pageId="15" pageNumber="210">Icochilus lamellosus</emphasis>
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<bibRefCitation id="5C414B70FFB5286773CACB3BFDE1FAFC" author="Ameghino, F." box="[406,623,1348,1374]" pageId="15" pageNumber="210" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino, 1894a</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="70CA650AFFB5286772CBCBF5FE76F802" pageId="15" pageNumber="210" type="materials_examined">
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<paragraph id="386F3681FFB5286772CBCBF5FB97FA01" blockId="15.[151,1437,1418,2025]" box="[151,1049,1418,1443]" pageId="15" pageNumber="210">
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<materialsCitation id="88B83CDCFFB5286772CBCBF5FB97FA01" ID-GBIF-Occurrence="2147525205" box="[151,1049,1418,1443]" collectionCode="MACN-A" pageId="15" pageNumber="210" specimenCode="MACN-A 3631" specimenCount="1" typeStatus="holotype">
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<emphasis id="0AA4EA93FFB5286772CBCBF5FE85FA01" bold="true" box="[151,267,1418,1443]" pageId="15" pageNumber="210">
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<typeStatus id="E76B8823FFB5286772CBCBF5FE88FA01" box="[151,262,1418,1443]" pageId="15" pageNumber="210" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="68769EFAFFB52867734ECBF4FE4CFA01" box="[274,450,1418,1443]" collectionCode="MACN-A" pageId="15" pageNumber="210">MACN-A 3631</specimenCode>
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(
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<figureCitation id="A0EB2A04FFB52867738ECBF5FDBCFA01" box="[466,562,1418,1443]" captionStart="FIGURE 5" captionStartId="17.[151,249,1387,1409]" captionTargetBox="[159,1432,215,1349]" captionTargetId="figure@17.[151,1437,193,1366]" captionTargetPageId="17" captionText="FIGURE 5. Icochilus lamellosus, holotype MACN-A 3631, incomplete skull, in ventral view (A). Icochilus trilineatus, MACN-A 3633, left M2, in occlusal view (B); MACN-A 3651 to MACN-A 3657 in occlusal view (C–H): left P4 or M1/2 (C), left P4 or M1/2 (D), left M3 (E), right P4 or M1/2 (F), right M3 (G), right m3 (H). Icochilus anomalus, detail of photograph of the holotype MACN-A 3634 from Scott’s catalogue (I). Icochilus truncus, holotype MACN-A 3635, right fragmented mandible with alveoli of i1–dp1 and complete p2–m1, in occlusal (J) and lateral (K) views. Icochilus crassiramis, holotype MACN-A 3636, right mandibular fragment with i2–m3, in occlusal (L) and lateral (M) views. Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617767/files/figure.png" pageId="15" pageNumber="210">Fig. 5 A</figureCitation>
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), incomplete skull with complete dentition.
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</materialsCitation>
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</paragraph>
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<paragraph id="386F3681FFB52867729BCBD1FEAFFA49" blockId="15.[151,1437,1418,2025]" pageId="15" pageNumber="210">
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<emphasis id="0AA4EA93FFB52867729BCBD1FD5DFA65" bold="true" box="[199,723,1454,1479]" pageId="15" pageNumber="210">
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Age and provenance of the
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<materialsCitation id="88B83CDCFFB52867707BCBD1FD5DFA65" ID-GBIF-Occurrence="2147525042" box="[551,723,1454,1479]" collectionCode="SALMA" country="Argentina" location="Miocene" pageId="15" pageNumber="210" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
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<typeStatus id="E76B8823FFB52867707BCBD1FDD4FA65" box="[551,602,1454,1479]" pageId="15" pageNumber="210">type</typeStatus>
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material.
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</materialsCitation>
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</emphasis>
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Santacrucian SALMA, early
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<location id="3D0F605AFFB52867766ACBD1FB15FA65" LSID="urn:lsid:plazi:treatment:B0798797FFB5287872CBCB3CFEB7FE25:3D0F605AFFB52867766ACBD1FB15FA65" box="[1078,1179,1454,1479]" country="Argentina" name="Miocene" pageId="15" pageNumber="210" stateProvince="Santa Cruz">Miocene</location>
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,
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<collectingRegion id="FA14F863FFB5286776F7CBD0FA12FA65" box="[1195,1436,1454,1479]" country="Argentina" name="Santa Cruz" pageId="15" pageNumber="210">Santa Cruz Province</collectingRegion>
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(
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<collectingCountry id="40C77611FFB5286772C2CBADFE9AFA49" box="[158,276,1490,1515]" name="Argentina" pageId="15" pageNumber="210">Argentina</collectingCountry>
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).
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</paragraph>
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<paragraph id="386F3681FFB52867729BCB89FB5FF96A" blockId="15.[151,1437,1418,2025]" pageId="15" pageNumber="210">
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<emphasis id="0AA4EA93FFB52867729BCB89FE30F9AD" bold="true" box="[199,446,1526,1551]" pageId="15" pageNumber="210">Original description</emphasis>
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(
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<bibRefCitation id="5C414B70FFB52867738FCB89FD74F9AD" author="Ameghino, F." box="[467,762,1526,1551]" pageId="15" pageNumber="210" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino 1894a: 15–16</bibRefCitation>
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). “
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<emphasis id="0AA4EA93FFB52867717CCB88FCA3F903" italics="true" pageId="15" pageNumber="210">Espèce de petite taille. La canine supérieure est bien développée, de la même grandeur et à peu près de la même forme que l'incisive externe, étant séparées l'une de l'autre par un petit diastème; ces deux dents sont comprimées latéralement et ont la forme de lames tranchantes. L'incisive externe ou troisième, est séparée de la deuxième par un diastème assez long; un autre diastème un peu plus long sépare la première prémolaire de la canine. La p</emphasis>
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<emphasis id="0AA4EA93FFB528677171C8F9FBA2F903" box="[813,1068,1670,1698]" italics="true" pageId="15" pageNumber="210">
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<superScript id="CFA59BC9FFB528677171C8F9FCBBF936" attach="left" box="[813,821,1670,1684]" fontSize="6" pageId="15" pageNumber="210">1</superScript>
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est placée contre la p
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</emphasis>
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<emphasis id="0AA4EA93FFB528677670C8F9FCADF96B" italics="true" pageId="15" pageNumber="210">
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<superScript id="CFA59BC9FFB528677670C8F9FBBAF936" attach="left" box="[1068,1076,1670,1684]" fontSize="6" pageId="15" pageNumber="210">2</superScript>
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et porte un sillon vertical à son angle antérieur externe. Longueur du bord antérieur de l'i
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</emphasis>
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<emphasis id="0AA4EA93FFB528677178C8D2FBD2F96A" box="[804,1116,1709,1737]" italics="true" pageId="15" pageNumber="210">
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<superScript id="CFA59BC9FFB528677178C8D2FCA2F919" attach="left" box="[804,812,1709,1723]" fontSize="6" pageId="15" pageNumber="210">1</superScript>
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au bord postérieur de la m
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</emphasis>
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<emphasis id="0AA4EA93FFB528677600C8D2FB33F96A" box="[1116,1213,1709,1736]" italics="true" pageId="15" pageNumber="210">
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<superScript id="CFA59BC9FFB528677600C8D2FBEAF919" attach="left" box="[1116,1124,1709,1723]" fontSize="6" pageId="15" pageNumber="210">3</superScript>
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, 37 mm
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</emphasis>
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”.
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</paragraph>
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<paragraph id="386F3681FFB52867729BC8ABFE76F802" blockId="15.[151,1437,1418,2025]" pageId="15" pageNumber="210">
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English translation. Species of small size. The upper canine is well developed, of the same size and nearly the same form as the external incisor, being separated from each other by a small diastema; these two teeth are laterally compressed and have the appearance of cutting blades. The external incisor, or third, is separated from the second by a rather long diastema; another slightly longer diastema separates the first premolar from the canine. The P1 is placed against P2 and has a vertical groove at its antero-external angle. Length from the anterior edge of
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<date id="4C6E1041FFB528677761C91BFADAF8DE" box="[1341,1364,1892,1916]" pageId="15" pageNumber="210">I1</date>
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to the posterior border of m3, 37 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="70CA650AFFB52878729BC9D3FEB7FE25" lastPageId="16" lastPageNumber="211" pageId="15" pageNumber="210" type="discussion">
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<paragraph id="386F3681FFB52878729BC9D3FEB7FE25" blockId="15.[151,1437,1418,2025]" lastBlockId="16.[151,1437,151,392]" lastPageId="16" lastPageNumber="211" pageId="15" pageNumber="210">
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<emphasis id="0AA4EA93FFB52867729BC9D3FEC1F861" bold="true" box="[199,335,1964,1989]" pageId="15" pageNumber="210">Comments.</emphasis>
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<bibRefCitation id="5C414B70FFB528677305C9D3FDBEF866" author="Ameghino, F." box="[345,560,1964,1989]" pageId="15" pageNumber="210" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
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did not add any illustration to the description of this species. According to Ameghino’s catalogue and
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<bibRefCitation id="5C414B70FFB52867738CC9AEFDFCF84B" author="Mones A." box="[464,626,2000,2025]" pageId="15" pageNumber="210" pagination="1 - 625" refId="ref17293" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article" year="1986">Mones (1986)</bibRefCitation>
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, the incomplete skull MACN-A 3631 is the type specimen. MACN-A 3631 exhibits a well-developed upper canine that is subequal to
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<date id="4C6E1041FFAA28787133CEE8FC0DFF12" box="[879,899,151,176]" pageId="16" pageNumber="211">I3</date>
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—being both teeth laterally compressed—, and the diastemata
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<date id="4C6E1041FFAA2878731ECEC3FEF6FF76" box="[322,376,188,212]" pageId="16" pageNumber="211">I2–3</date>
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,
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<date id="4C6E1041FFAA287873DACEC3FE15FF76" box="[390,411,188,212]" pageId="16" pageNumber="211">I3</date>
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–C and C–dP1 mentioned by
|
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<bibRefCitation id="5C414B70FFAA287870ACCEC3FC47FF76" author="Ameghino, F." box="[752,969,188,213]" pageId="16" pageNumber="211" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
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. Even though the author did not indicate the dimensions of these discontinuities, he established relative differences in size, which resemble those obtained in the material (length of diastema
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<date id="4C6E1041FFAA2878707FCF7BFDD8FEBE" box="[547,598,260,284]" pageId="16" pageNumber="211">I2–3</date>
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||
= left,
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||
<quantity id="FF289B64FFAA287870F6CF7BFC8EFEBE" box="[682,768,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="211" unit="mm" value="0.9">0.9 mm</quantity>
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||
and right,
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||
<quantity id="FF289B64FFAA28787127CF7BFC5BFEBE" box="[891,981,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="211" unit="mm" value="1.4">1.4 mm</quantity>
|
||
; length diastema
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||
<date id="4C6E1041FFAA287876C6CF7BFB3FFEBE" box="[1178,1201,260,284]" pageId="16" pageNumber="211">I3</date>
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||
–C: left,
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||
<quantity id="FF289B64FFAA2878774ACF7BFAE3FEBE" box="[1302,1389,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="211" unit="mm" value="0.4">0.4 mm</quantity>
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||
and right,
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<quantity id="FF289B64FFAA28787280CF57FEB8FE9D" box="[220,310,296,320]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="211" unit="mm" value="0.8">0.8 mm</quantity>
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||
; length of diastema C–dP1: left,
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||
<quantity id="FF289B64FFAA287870EDCF58FC87FE9D" box="[689,777,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="16" pageNumber="211" unit="mm" value="2.6">2.6 mm</quantity>
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||
and right,
|
||
<quantity id="FF289B64FFAA287871D4CF57FC6DFEE2" box="[904,995,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="16" pageNumber="211" unit="mm" value="2.2">2.2 mm</quantity>
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||
). The length of
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||
<date id="4C6E1041FFAA287876C7CF57FB20FEE2" box="[1179,1198,296,320]" pageId="16" pageNumber="211">I1</date>
|
||
–M3 series (
|
||
<quantity id="FF289B64FFAA2878771DCF57FA1AFEE2" box="[1345,1428,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="16" pageNumber="211" unit="mm" value="38.0">38 mm</quantity>
|
||
) also matches the value provided by
|
||
<bibRefCitation id="5C414B70FFAA28787075CF33FC8EFEC6" author="Ameghino, F." box="[553,768,332,357]" pageId="16" pageNumber="211" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
|
||
. Therefore, MACN-A 3631 is confirmed as the
|
||
<typeStatus id="E76B8823FFAA28787740CF33FAF0FEC7" box="[1308,1406,332,357]" pageId="16" pageNumber="211" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="FFD04D02FFAA287872CBCF0EFEBDFE2A" authorityName="Ameghino" authorityYear="1894" box="[151,307,367,392]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="211" phylum="Chordata" rank="species" species="lamellosus">
|
||
<emphasis id="0AA4EA93FFAA287872CBCF0EFEBDFE2A" box="[151,307,367,392]" italics="true" pageId="16" pageNumber="211">Ic. lamellosus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |