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<document id="EAB11344217CAAABD2B4769FC9924F58" ID-CLB-Dataset="27706" ID-DOI="10.11646/zootaxa.4543.2.2" ID-GBIF-Dataset="7438332f-a925-43aa-b17f-059781bdfc12" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617757" ID-ZooBank="11F546CD-109F-4CBC-B87D-EA8D1AD0B96F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928261775" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos &amp; Cerdeño, Esperanza" docDate="2019" docId="B0798797FFB4286672CBCAE0FAEBF86C" docLanguage="en" docName="zootaxa.4543.2.2.pdf" docOrigin="Zootaxa 4543 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Icochilus senilis Ameghino 1894" docType="treatment" docVersion="8" lastPageNumber="209" masterDocId="4C40FFEFFFBA2868725CCE7FFF8EFFA2" masterDocTitle="On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina" masterLastPageNumber="220" masterPageNumber="195" pageNumber="209" updateTime="1701997211094" updateUser="plazi">
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<mods:title id="159B3CC1EAC80C0E7546668230351BCA">On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina</mods:title>
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<mods:namePart id="2D5E2E374B13B5078E06603892BF2E58">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="2D76965DC2688F5480B54CEC13D72202">Fernicola, Juan Carlos</mods:namePart>
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<treatment id="B0798797FFB4286672CBCAE0FAEBF86C" ID-DOI="http://doi.org/10.5281/zenodo.5931295" ID-GBIF-Taxon="154529761" ID-Zenodo-Dep="5931295" LSID="urn:lsid:plazi:treatment:B0798797FFB4286672CBCAE0FAEBF86C" httpUri="http://treatment.plazi.org/id/B0798797FFB4286672CBCAE0FAEBF86C" lastPageNumber="209" pageId="14" pageNumber="209">
<subSubSection id="70CA650AFFB4286672CBCAE0FDB5FB18" box="[151,571,1183,1210]" pageId="14" pageNumber="209" type="nomenclature">
<paragraph id="386F3681FFB4286672CBCAE0FDB5FB18" blockId="14.[151,571,1183,1210]" box="[151,571,1183,1210]" pageId="14" pageNumber="209">
<heading id="632781EDFFB4286672CBCAE0FDB5FB18" bold="true" box="[151,571,1183,1210]" fontSize="11" level="1" pageId="14" pageNumber="209" reason="1">
<taxonomicName id="FFD04D02FFB4286672CBCAE0FDB5FB18" authority="Ameghino, 1894 a" authorityName="Ameghino" authorityYear="1894" box="[151,571,1183,1210]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="senilis">
<emphasis id="0AA4EA93FFB4286672CBCAE0FDB5FB18" bold="true" box="[151,571,1183,1210]" pageId="14" pageNumber="209">
<emphasis id="0AA4EA93FFB4286672CBCAE0FED5FB1B" bold="true" box="[151,347,1183,1209]" italics="true" pageId="14" pageNumber="209">Icochilus senilis</emphasis>
<bibRefCitation id="5C414B70FFB42866733ECADFFDB5FB18" author="Ameghino, F." box="[354,571,1184,1210]" pageId="14" pageNumber="209" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino, 1894a</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="70CA650AFFB4286672CBCA99FB1BF954" pageId="14" pageNumber="209" type="materials_examined">
<paragraph id="386F3681FFB4286672CBCA99FB30FA81" blockId="14.[151,1437,1254,1999]" pageId="14" pageNumber="209">
<materialsCitation id="88B83CDCFFB4286672CBCA99FB30FA81" ID-GBIF-Occurrence="2147524937" collectionCode="MACN-A" pageId="14" pageNumber="209" specimenCode="MACN-A 3596, MACN-A 3597, MACN-A 3598, MACN-A 3630" specimenCount="1" typeStatus="holotype">
<emphasis id="0AA4EA93FFB4286672CBCA99FE85FB5D" bold="true" box="[151,267,1254,1279]" pageId="14" pageNumber="209">
<typeStatus id="E76B8823FFB4286672CBCA99FE88FB5D" box="[151,262,1254,1279]" pageId="14" pageNumber="209" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="68769EFAFFB42866734BCA98FE43FB5D" box="[279,461,1254,1279]" collectionCode="MACN-A" pageId="14" pageNumber="209">MACN-A 3596</specimenCode>
(
<figureCitation id="A0EB2A04FFB4286673BDCA99FDC4FB5D" box="[481,586,1254,1279]" captionStart="FIGURE 4" captionStartId="15.[151,249,1189,1211]" captionTargetBox="[151,1431,217,1130]" captionTargetId="figure@15.[151,1436,193,1167]" captionTargetPageId="15" captionText="FIGURE 4. Icochilus robustus, holotype MACN-A 3454, skull, in dorsal view (A); reproduction of figure 97 (B) from Ameghino (1891). Icochilus senilis, holotype MACN-A 3596, incomplete skull, in ventral view (C); holotype MACN-A 3597, incomplete mandible, in dorsal view (D). Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617765/files/figure.png" pageId="14" pageNumber="209">Fig. 4 C</figureCitation>
), incomplete skull,
<specimenCode id="68769EFAFFB42866716BCA98FC62FB5D" box="[823,1004,1254,1279]" collectionCode="MACN-A" pageId="14" pageNumber="209">MACN-A 3597</specimenCode>
(
<figureCitation id="A0EB2A04FFB42866765DCA99FBE2FB5D" box="[1025,1132,1254,1279]" captionStart="FIGURE 4" captionStartId="15.[151,249,1189,1211]" captionTargetBox="[151,1431,217,1130]" captionTargetId="figure@15.[151,1436,193,1167]" captionTargetPageId="15" captionText="FIGURE 4. Icochilus robustus, holotype MACN-A 3454, skull, in dorsal view (A); reproduction of figure 97 (B) from Ameghino (1891). Icochilus senilis, holotype MACN-A 3596, incomplete skull, in ventral view (C); holotype MACN-A 3597, incomplete mandible, in dorsal view (D). Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617765/files/figure.png" pageId="14" pageNumber="209">Fig. 4 D</figureCitation>
), complete mandible, and
<specimenCode id="68769EFAFFB4286672CBCB74FEC9FA81" box="[151,327,1290,1315]" collectionCode="MACN-A" pageId="14" pageNumber="209">MACN-A 3598</specimenCode>
to
<specimenCode id="68769EFAFFB428667337CB74FD91FA81" box="[363,543,1290,1315]" collectionCode="MACN-A" pageId="14" pageNumber="209">MACN-A 3630</specimenCode>
, postcranial elements, all of them from the same individual.
</materialsCitation>
</paragraph>
<paragraph id="386F3681FFB42866729BCB51FEAFFAC9" blockId="14.[151,1437,1254,1999]" pageId="14" pageNumber="209">
<emphasis id="0AA4EA93FFB42866729BCB51FD5DFAE5" bold="true" box="[199,723,1326,1351]" pageId="14" pageNumber="209">
Age and provenance of the
<materialsCitation id="88B83CDCFFB42866707BCB51FD5DFAE5" ID-GBIF-Occurrence="2147525041" box="[551,723,1326,1351]" collectionCode="SALMA" country="Argentina" location="Miocene" pageId="14" pageNumber="209" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<typeStatus id="E76B8823FFB42866707BCB51FDD4FAE5" box="[551,602,1326,1351]" pageId="14" pageNumber="209">type</typeStatus>
material.
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</emphasis>
Santacrucian SALMA, early
<location id="3D0F605AFFB42866766ACB51FB15FAE5" LSID="urn:lsid:plazi:treatment:B0798797FFB4286672CBCAE0FAEBF86C:3D0F605AFFB42866766ACB51FB15FAE5" box="[1078,1179,1326,1351]" country="Argentina" name="Miocene" pageId="14" pageNumber="209" stateProvince="Santa Cruz">Miocene</location>
,
<collectingRegion id="FA14F863FFB4286676F7CB50FA12FAE5" box="[1195,1436,1326,1351]" country="Argentina" name="Santa Cruz" pageId="14" pageNumber="209">Santa Cruz Province</collectingRegion>
(
<collectingCountry id="40C77611FFB4286672C2CB2DFE9AFAC9" box="[158,276,1362,1387]" name="Argentina" pageId="14" pageNumber="209">Argentina</collectingCountry>
).
</paragraph>
<paragraph id="386F3681FFB42866729BCB09FBD6F9E1" blockId="14.[151,1437,1254,1999]" pageId="14" pageNumber="209">
<emphasis id="0AA4EA93FFB42866729BCB09FE37FA2D" bold="true" box="[199,441,1398,1423]" pageId="14" pageNumber="209">Original description</emphasis>
(
<bibRefCitation id="5C414B70FFB428667396CB09FD33FA2D" author="Ameghino, F." box="[458,701,1398,1423]" pageId="14" pageNumber="209" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino 1894a: 15</bibRefCitation>
). “
<emphasis id="0AA4EA93FFB428667083CB08FBDDFA2D" box="[735,1107,1398,1423]" italics="true" pageId="14" pageNumber="209">Cette espèce est de la taille de l</emphasis>
<taxonomicName id="FFD04D02FFB42866760FCB08FB42FA2C" authorityName="Ameghino" authorityYear="1889" box="[1107,1228,1399,1422]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="extensus">I. extensus</taxonomicName>
,
<emphasis id="0AA4EA93FFB428667687CB08FBCAF9E1" italics="true" pageId="14" pageNumber="209">
mais un peu plus robuste; elle se distingue facilement par la deuxième prémolaire inférieure qui n'est pas bilobée, mais de contour elliptique et par conséquent sans sillon vertical, ni sur la face interne ni sur l'externe. La troisième et quatrième prémolaire inférieure ainsi que les vraies molaires, sont plus larges que dans les autres espèces. La deuxième prémolaire supérieure est aussi de contour elliptique et sans sillon. Il y a un diastème assez long entre la canine inférieure et la première prémolaire. Longueur des sept molaires inférieures,
<quantity id="FF289B64FFB4286671A5C854FBCAF9E1" box="[1017,1092,1579,1603]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="14" pageNumber="209" unit="mm" value="32.0">32 mm</quantity>
</emphasis>
”.
</paragraph>
<paragraph id="386F3681FFB42866729BC831FB1BF954" blockId="14.[151,1437,1254,1999]" pageId="14" pageNumber="209">
English translation. This species is of the size of
<taxonomicName id="FFD04D02FFB428667141C830FC28F9C4" authorityName="Ameghino" authorityYear="1889" box="[797,934,1615,1638]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="extensus">
<emphasis id="0AA4EA93FFB428667141C830FC28F9C4" box="[797,934,1615,1638]" italics="true" pageId="14" pageNumber="209">Ic. extensus</emphasis>
</taxonomicName>
, but it is a little more robust; it is easily distinguished by the second lower premolar that is not bilobated but elliptical and, subsequently, without internal and external vertical grooves. The third and fourth lower premolars, as well as the true molars, are wider than in the other species. The second upper premolar is also elliptical and without groove. There is a rather long diastema between the lower canine and the first premolar. Length of the seven lower molars,
<quantity id="FF289B64FFB428667663C8A0FB1FF954" box="[1087,1169,1759,1783]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="14" pageNumber="209" unit="mm" value="32.0">32 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="70CA650AFFB42866729BC97DFAEBF86C" pageId="14" pageNumber="209" type="discussion">
<paragraph id="386F3681FFB42866729BC97DFAEBF86C" blockId="14.[151,1437,1254,1999]" pageId="14" pageNumber="209">
<emphasis id="0AA4EA93FFB42866729BC97DFEC1F8B8" bold="true" box="[199,335,1794,1819]" pageId="14" pageNumber="209">Comments.</emphasis>
<bibRefCitation id="5C414B70FFB428667307C97DFDBAF8B9" author="Ameghino, F." box="[347,564,1794,1819]" pageId="14" pageNumber="209" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
did not illustrate any specimen of this species. According to Ameghinos catalogue and
<bibRefCitation id="5C414B70FFB42866736BC958FE59F89D" author="Mones A." box="[311,471,1830,1855]" pageId="14" pageNumber="209" pagination="1 - 625" refId="ref17293" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article" year="1986">Mones (1986)</bibRefCitation>
, MACN-A 3596 to MACN-A 3630, an almost complete individual, constitute the type specimen of this species. The mandible MACN-A 3597 and the skull MACN-A 3596 present the respective p2 and P2 elliptical in cross section with very shallow lingual and labial grooves. In addition, the dimensions of MACN-A 3597 (length of the diastema cdp1 =
<quantity id="FF289B64FFB42866700EC9ECFD16F808" box="[594,664,1939,1963]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="209" unit="mm" value="2.0">2 mm</quantity>
; length of the lower tooth row =
<quantity id="FF289B64FFB428667641C9ECFBFEF809" box="[1053,1136,1939,1963]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="14" pageNumber="209" unit="mm" value="32.0">32 mm</quantity>
) match those provided by
<bibRefCitation id="5C414B70FFB4286672CBC9C9FEE0F86D" author="Ameghino, F." box="[151,366,1974,1999]" pageId="14" pageNumber="209" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
. Therefore, MACN-A 3596 to MACN-A 3630 are confirmed as the
<typeStatus id="E76B8823FFB428667630C9C9FB40F86D" box="[1132,1230,1974,1999]" pageId="14" pageNumber="209" type="holotype">holotype</typeStatus>
of
<taxonomicName id="FFD04D02FFB4286676AFC9C8FAD0F86D" authorityName="Ameghino" authorityYear="1894" box="[1267,1374,1974,1999]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="senilis">
<emphasis id="0AA4EA93FFB4286676AFC9C8FAD0F86D" box="[1267,1374,1974,1999]" italics="true" pageId="14" pageNumber="209">Ic. senilis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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