148 lines
18 KiB
XML
148 lines
18 KiB
XML
<document id="BACA5A7A96A3E2999908E3A896F084AB" ID-CLB-Dataset="27706" ID-DOI="10.11646/zootaxa.4543.2.2" ID-GBIF-Dataset="7438332f-a925-43aa-b17f-059781bdfc12" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617757" ID-ZooBank="11F546CD-109F-4CBC-B87D-EA8D1AD0B96F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928261775" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos & Cerdeño, Esperanza" docDate="2019" docId="B0798797FFAA287872CBCFA3FE32FA40" docLanguage="en" docName="zootaxa.4543.2.2.pdf" docOrigin="Zootaxa 4543 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Icochilus trilineatus Ameghino 1894" docType="treatment" docVersion="8" lastPageNumber="211" masterDocId="4C40FFEFFFBA2868725CCE7FFF8EFFA2" masterDocTitle="On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina" masterLastPageNumber="220" masterPageNumber="195" pageNumber="211" updateTime="1701997211094" updateUser="plazi">
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<mods:title id="389ECAE349295EBE4D75B3F2D9DBD2B8">On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina</mods:title>
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<mods:namePart id="8CE0A4E08AC3C00B7FEFBD3A6C9BF0E6">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="B69A30570C7BA3657B2BFBADE474CBAE">Fernicola, Juan Carlos</mods:namePart>
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<mods:namePart id="52DD0A3373ECB660BCC86B042392B23D">Cerdeño, Esperanza</mods:namePart>
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<treatment id="B0798797FFAA287872CBCFA3FE32FA40" ID-DOI="http://doi.org/10.5281/zenodo.5931299" ID-GBIF-Taxon="154529770" ID-Zenodo-Dep="5931299" LSID="urn:lsid:plazi:treatment:B0798797FFAA287872CBCFA3FE32FA40" httpUri="http://treatment.plazi.org/id/B0798797FFAA287872CBCFA3FE32FA40" lastPageNumber="211" pageId="16" pageNumber="211">
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<subSubSection id="70CA650AFFAA287872CBCFA3FDAAFDFD" pageId="16" pageNumber="211" type="nomenclature">
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<paragraph id="386F3681FFAA287872CBCFA3FDE4FE54" blockId="16.[151,618,476,502]" box="[151,618,476,502]" pageId="16" pageNumber="211">
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<heading id="632781EDFFAA287872CBCFA3FDE4FE54" bold="true" box="[151,618,476,502]" fontSize="11" level="1" pageId="16" pageNumber="211" reason="1">
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<taxonomicName id="FFD04D02FFAA287872CBCFA3FDE4FE54" authority="Ameghino, 1894 a" authorityName="Ameghino" authorityYear="1894" box="[151,618,476,502]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="211" phylum="Chordata" rank="species" species="trilineatus">
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<emphasis id="0AA4EA93FFAA287872CBCFA3FDE4FE54" bold="true" box="[151,618,476,502]" pageId="16" pageNumber="211">
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<emphasis id="0AA4EA93FFAA287872CBCFA3FE04FE54" bold="true" box="[151,394,476,502]" italics="true" pageId="16" pageNumber="211">Icochilus trilineatus</emphasis>
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<bibRefCitation id="5C414B70FFAA287873CECFA3FDE4FE54" author="Ameghino, F." box="[402,618,476,502]" pageId="16" pageNumber="211" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino, 1894a</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="386F3681FFAA287872CBCC5DFDAAFDFD" blockId="16.[151,1437,546,1507]" pageId="16" pageNumber="211">
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<materialsCitation id="88B83CDCFFAA287872CBCC5DFB8DFD99" ID-GBIF-Occurrence="2147525035" box="[151,1027,546,571]" collectionCode="MACN-A" pageId="16" pageNumber="211" specimenCode="MACN-A 3633" specimenCount="1" typeStatus="syntype">
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<emphasis id="0AA4EA93FFAA287872CBCC5DFE87FD99" bold="true" box="[151,265,546,571]" pageId="16" pageNumber="211">
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<typeStatus id="E76B8823FFAA287872CBCC5DFE8BFD99" box="[151,261,546,571]" pageId="16" pageNumber="211" type="syntype">Syntypes</typeStatus>
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.
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</emphasis>
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<specimenCode id="68769EFAFFAA2878734ECC5CFE4BFD99" box="[274,453,546,571]" collectionCode="MACN-A" pageId="16" pageNumber="211">MACN-A 3633</specimenCode>
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(
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<figureCitation id="A0EB2A04FFAA2878738BCC5DFDB5FD99" box="[471,571,546,571]" captionStart="FIGURE 5" captionStartId="17.[151,249,1387,1409]" captionTargetBox="[159,1432,215,1349]" captionTargetId="figure@17.[151,1437,193,1366]" captionTargetPageId="17" captionText="FIGURE 5. Icochilus lamellosus, holotype MACN-A 3631, incomplete skull, in ventral view (A). Icochilus trilineatus, MACN-A 3633, left M2, in occlusal view (B); MACN-A 3651 to MACN-A 3657 in occlusal view (C–H): left P4 or M1/2 (C), left P4 or M1/2 (D), left M3 (E), right P4 or M1/2 (F), right M3 (G), right m3 (H). Icochilus anomalus, detail of photograph of the holotype MACN-A 3634 from Scott’s catalogue (I). Icochilus truncus, holotype MACN-A 3635, right fragmented mandible with alveoli of i1–dp1 and complete p2–m1, in occlusal (J) and lateral (K) views. Icochilus crassiramis, holotype MACN-A 3636, right mandibular fragment with i2–m3, in occlusal (L) and lateral (M) views. Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617767/files/figure.png" pageId="16" pageNumber="211">Fig. 5 B</figureCitation>
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), fragmented left maxilla with M2; and
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</materialsCitation>
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<materialsCitation id="88B83CDCFFAA28787651CC5CFB6EFD98" ID-GBIF-Occurrence="2147525034" box="[1037,1248,546,571]" collectionCode="MACN-A" pageId="16" pageNumber="211" specimenCode="MACN-A 3651" specimenCount="1" typeStatus="syntype">
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<specimenCode id="68769EFAFFAA28787651CC5CFB4EFD99" box="[1037,1216,546,571]" collectionCode="MACN-A" pageId="16" pageNumber="211">MACN-A 3651</specimenCode>
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to
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</materialsCitation>
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<materialsCitation id="88B83CDCFFAA287876B5CC5CFDAEFDFD" ID-GBIF-Occurrence="2147525031" collectionCode="MACN-A" pageId="16" pageNumber="211" specimenCode="MACN-A 3657" specimenCount="1" typeStatus="syntype">
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<specimenCode id="68769EFAFFAA287876B5CC5CFA12FD99" box="[1257,1436,546,571]" collectionCode="MACN-A" pageId="16" pageNumber="211">MACN-A 3657</specimenCode>
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(
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<figureCitation id="A0EB2A04FFAA287872C3CC39FF62FDFD" box="[159,236,582,607]" captionStart="FIGURE 5" captionStartId="17.[151,249,1387,1409]" captionTargetBox="[159,1432,215,1349]" captionTargetId="figure@17.[151,1437,193,1366]" captionTargetPageId="17" captionText="FIGURE 5. Icochilus lamellosus, holotype MACN-A 3631, incomplete skull, in ventral view (A). Icochilus trilineatus, MACN-A 3633, left M2, in occlusal view (B); MACN-A 3651 to MACN-A 3657 in occlusal view (C–H): left P4 or M1/2 (C), left P4 or M1/2 (D), left M3 (E), right P4 or M1/2 (F), right M3 (G), right m3 (H). Icochilus anomalus, detail of photograph of the holotype MACN-A 3634 from Scott’s catalogue (I). Icochilus truncus, holotype MACN-A 3635, right fragmented mandible with alveoli of i1–dp1 and complete p2–m1, in occlusal (J) and lateral (K) views. Icochilus crassiramis, holotype MACN-A 3636, right mandibular fragment with i2–m3, in occlusal (L) and lateral (M) views. Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617767/files/figure.png" pageId="16" pageNumber="211">Figs. 5</figureCitation>
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C–H), isolated cheek teeth
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="70CA650AFFAA2878729BCC15FB4FFB90" pageId="16" pageNumber="211" type="materials_examined">
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<paragraph id="386F3681FFAA2878729BCC15FEAFFD05" blockId="16.[151,1437,546,1507]" pageId="16" pageNumber="211">
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<emphasis id="0AA4EA93FFAA2878729BCC15FD5DFD21" bold="true" box="[199,723,618,643]" pageId="16" pageNumber="211">
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Age and provenance of the
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<materialsCitation id="88B83CDCFFAA2878707BCC15FD5DFD21" ID-GBIF-Occurrence="2147525029" box="[551,723,618,643]" collectionCode="SALMA" country="Argentina" location="Miocene" pageId="16" pageNumber="211" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
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<typeStatus id="E76B8823FFAA2878707BCC15FDD4FD21" box="[551,602,618,643]" pageId="16" pageNumber="211">type</typeStatus>
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material.
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</materialsCitation>
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</emphasis>
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Santacrucian SALMA, early
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<location id="3D0F605AFFAA2878766ACC15FB15FD21" LSID="urn:lsid:plazi:treatment:B0798797FFAA287872CBCFA3FE32FA40:3D0F605AFFAA2878766ACC15FB15FD21" box="[1078,1179,618,643]" country="Argentina" name="Miocene" pageId="16" pageNumber="211" stateProvince="Santa Cruz">Miocene</location>
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,
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<collectingRegion id="FA14F863FFAA287876F7CC14FA12FD21" box="[1195,1436,618,643]" country="Argentina" name="Santa Cruz" pageId="16" pageNumber="211">Santa Cruz Province</collectingRegion>
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(
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<collectingCountry id="40C77611FFAA287872C2CCF1FE9AFD05" box="[158,276,654,679]" name="Argentina" pageId="16" pageNumber="211">Argentina</collectingCountry>
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).
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</paragraph>
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<paragraph id="386F3681FFAA2878729BCCCDFE5EFCDD" blockId="16.[151,1437,546,1507]" pageId="16" pageNumber="211">
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<emphasis id="0AA4EA93FFAA2878729BCCCDFE36FD69" bold="true" box="[199,440,690,715]" pageId="16" pageNumber="211">Original description</emphasis>
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(
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<bibRefCitation id="5C414B70FFAA2878739ACCCDFD38FD69" author="Ameghino, F." box="[454,694,690,715]" pageId="16" pageNumber="211" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino 1894a: 16</bibRefCitation>
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). “
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<emphasis id="0AA4EA93FFAA2878708ACCCCFB1DFD69" box="[726,1171,690,715]" italics="true" pageId="16" pageNumber="211">Cette espèce est de la même taille que l’</emphasis>
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<taxonomicName id="FFD04D02FFAA287876C5CCCCFA9EFD68" authorityName="Ameghino" authorityYear="1889" box="[1177,1296,691,714]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="211" phylum="Chordata" rank="species" species="extensus">I. extensus</taxonomicName>
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,
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<emphasis id="0AA4EA93FFAA28787741CCCDFE32FCDC" italics="true" pageId="16" pageNumber="211">dont elle se distingue facilement aussi bien que des autres espèces, par la forme de ses molaires et prémolaires supérieures. Chacune de ces dents porte sur sa face externe un sillon large et profond qui la divise en deux lobules, et sur chaque lobule il y a un sillon étroit et profond qui le divise en deux colonnettes verticales; on voit ainsi sur la face externe de chaque dent, quatre colonnettes séparées par trois sillons, dont celui du milieu beaucoup plus large et les autres deux très étroits</emphasis>
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”.
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</paragraph>
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<paragraph id="386F3681FFAA2878729BCDF5FB4FFB90" blockId="16.[151,1437,546,1507]" pageId="16" pageNumber="211">
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English translation. This species has the same size as
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<taxonomicName id="FFD04D02FFAA28787140CDF3FC10FC01" authorityName="Ameghino" authorityYear="1889" box="[796,926,908,931]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="211" phylum="Chordata" rank="species" species="extensus">
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<emphasis id="0AA4EA93FFAA28787140CDF3FC10FC01" box="[796,926,908,931]" italics="true" pageId="16" pageNumber="211">Ic. extensus</emphasis>
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</taxonomicName>
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, from which it is easily distinguished, as well as from the other species, by the morphology of its upper molars and premolars. Each tooth presents a broad and deep groove on its external face, which divides it into two lobules, and on each lobule there is a narrow and deep groove that divides it into two vertical columns; as a result, the external face of each tooth exhibits four columns separated by three grooves, the second being the widest whereas the other two are very narrow.
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</paragraph>
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</subSubSection>
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<subSubSection id="70CA650AFFAA2878729BCA41FE32FA40" pageId="16" pageNumber="211" type="discussion">
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<paragraph id="386F3681FFAA2878729BCA41FE32FA40" blockId="16.[151,1437,546,1507]" pageId="16" pageNumber="211">
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<emphasis id="0AA4EA93FFAA2878729BCA41FEC1FBF4" bold="true" box="[199,335,1086,1111]" pageId="16" pageNumber="211">Comments.</emphasis>
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<bibRefCitation id="5C414B70FFAA2878730ACA41FDA5FBF5" author="Ameghino, F." box="[342,555,1086,1111]" pageId="16" pageNumber="211" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
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did not illustrate this species. According to Ameghino’s catalogue and
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<bibRefCitation id="5C414B70FFAA2878770CCA40FF6DFBD9" author="Mones A." pageId="16" pageNumber="211" pagination="1 - 625" refId="ref17293" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article" year="1986">Mones (1986)</bibRefCitation>
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, MACN-A 3633, a small maxillary fragment with M2, is the type specimen of the species. The ectoloph of this M2 exhibits three perpendicular grooves, one in the middle and one at each lateral lobe, resulting in an ectoloph with four columns as indicated by
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<bibRefCitation id="5C414B70FFAA287870EACAD5FC1AFB61" author="Ameghino, F." box="[694,916,1194,1219]" pageId="16" pageNumber="211" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
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. Nonetheless, when describing the species,
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<bibRefCitation id="5C414B70FFAA287872CBCAB1FE19FB45" author="Ameghino, F." box="[151,407,1230,1255]" pageId="16" pageNumber="211" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a: 16)</bibRefCitation>
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alluded to the form of its upper molars and premolars, which implies that he had more than one tooth at hand, and therefore MACN-A 3633 cannot be the only type specimen. In this context, in the Ameghino Collection, the lot MACN-A 3651 to MACN-A 3657 contains six isolated teeth (instead of seven as corresponding inventory numbers), five of which (two M3, and three upper molariforms, probably P4 and/or M1/2) show the characteristics mentioned by
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<bibRefCitation id="5C414B70FFAA287873AECB21FD5EFAD5" author="Ameghino, F." box="[498,720,1374,1399]" pageId="16" pageNumber="211" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
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; thus, we consider them as part of the type series. As the specimens in the lot lack individual labels, it is not possible to identify which is the missing seventh tooth. Therefore, MACN-A 3633 and MACN-A 3651 to MACN-A 3657 (one currently missing) are considered the
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<typeStatus id="E76B8823FFAA287872CBCBB4FF77FA41" box="[151,249,1483,1507]" pageId="16" pageNumber="211" type="syntype">syntypes</typeStatus>
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of
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<taxonomicName id="FFD04D02FFAA28787342CBB3FE3BFA41" authorityName="Ameghino" authorityYear="1894" box="[286,437,1482,1507]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="211" phylum="Chordata" rank="species" species="trilineatus">
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<emphasis id="0AA4EA93FFAA28787342CBB3FE3BFA41" box="[286,437,1482,1507]" italics="true" pageId="16" pageNumber="211">Ic. trilineatus</emphasis>
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</taxonomicName>
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.
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</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |