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<document id="21A6B4412EDD6AAC8470B14CCAE0216B" ID-CLB-Dataset="27706" ID-DOI="10.11646/zootaxa.4543.2.2" ID-GBIF-Dataset="7438332f-a925-43aa-b17f-059781bdfc12" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617757" ID-ZooBank="11F546CD-109F-4CBC-B87D-EA8D1AD0B96F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928261775" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos &amp; Cerdeño, Esperanza" docDate="2019" docId="B0798797FFA8287A72CBCF33FB04FC01" docLanguage="en" docName="zootaxa.4543.2.2.pdf" docOrigin="Zootaxa 4543 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Icochilus truncus Ameghino 1894" docType="treatment" docVersion="8" lastPageNumber="213" masterDocId="4C40FFEFFFBA2868725CCE7FFF8EFFA2" masterDocTitle="On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina" masterLastPageNumber="220" masterPageNumber="195" pageNumber="213" updateTime="1701997211094" updateUser="plazi">
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<mods:title id="B3D37EAD2E4136C759DB022D9C16FF36">On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina</mods:title>
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<mods:namePart id="C677F3BB8F887C445111235EA1A8B52E">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="938D9A5E41385DECB32F5F8BBF16873A">Fernicola, Juan Carlos</mods:namePart>
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<mods:namePart id="717C3D6F6BEC3C9BD36DB93B7C2C3AA8">Cerdeño, Esperanza</mods:namePart>
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<treatment id="B0798797FFA8287A72CBCF33FB04FC01" ID-DOI="http://doi.org/10.5281/zenodo.5931303" ID-GBIF-Taxon="154529772" ID-Zenodo-Dep="5931303" LSID="urn:lsid:plazi:treatment:B0798797FFA8287A72CBCF33FB04FC01" httpUri="http://treatment.plazi.org/id/B0798797FFA8287A72CBCF33FB04FC01" lastPageNumber="213" pageId="18" pageNumber="213">
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<paragraph id="386F3681FFA8287A72CBCF33FDC5FEC4" blockId="18.[151,587,332,358]" box="[151,587,332,358]" pageId="18" pageNumber="213">
<heading id="632781EDFFA8287A72CBCF33FDC5FEC4" bold="true" box="[151,587,332,358]" fontSize="11" level="1" pageId="18" pageNumber="213" reason="1">
<taxonomicName id="FFD04D02FFA8287A72CBCF33FDC5FEC4" authority="Ameghino, 1894 a" authorityName="Ameghino" authorityYear="1894" box="[151,587,332,358]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="18" pageNumber="213" phylum="Chordata" rank="species" species="truncus">
<emphasis id="0AA4EA93FFA8287A72CBCF33FDC5FEC4" bold="true" box="[151,587,332,358]" pageId="18" pageNumber="213">
<emphasis id="0AA4EA93FFA8287A72CBCF33FEE5FEC4" bold="true" box="[151,363,332,358]" italics="true" pageId="18" pageNumber="213">Icochilus truncus</emphasis>
<bibRefCitation id="5C414B70FFA8287A732FCF33FDC5FEC4" author="Ameghino, F." box="[371,587,332,358]" pageId="18" pageNumber="213" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino, 1894a</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="70CA650AFFA8287A72CBCFEDFC0AFD69" pageId="18" pageNumber="213" type="materials_examined">
<paragraph id="386F3681FFA8287A72CBCFEDFA1EFE09" blockId="18.[151,1437,402,931]" box="[151,1424,402,427]" pageId="18" pageNumber="213">
<materialsCitation id="88B83CDCFFA8287A72CBCFEDFA1EFE09" ID-GBIF-Occurrence="2147525036" box="[151,1424,402,427]" collectingDate="i 1" collectionCode="MACN-A" pageId="18" pageNumber="213" specimenCode="MACN-A 3635" specimenCount="1" typeStatus="holotype">
<emphasis id="0AA4EA93FFA8287A72CBCFEDFE85FE09" bold="true" box="[151,267,402,427]" pageId="18" pageNumber="213">
<typeStatus id="E76B8823FFA8287A72CBCFEDFE88FE09" box="[151,262,402,427]" pageId="18" pageNumber="213" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="68769EFAFFA8287A734ECFECFE4CFE09" box="[274,450,402,427]" collectionCode="MACN-A" pageId="18" pageNumber="213">MACN-A 3635</specimenCode>
(
<figureCitation id="A0EB2A04FFA8287A738ECFEDFD91FE09" box="[466,543,402,427]" captionStart="FIGURE 5" captionStartId="17.[151,249,1387,1409]" captionTargetBox="[159,1432,215,1349]" captionTargetId="figure@17.[151,1437,193,1366]" captionTargetPageId="17" captionText="FIGURE 5. Icochilus lamellosus, holotype MACN-A 3631, incomplete skull, in ventral view (A). Icochilus trilineatus, MACN-A 3633, left M2, in occlusal view (B); MACN-A 3651 to MACN-A 3657 in occlusal view (CH): left P4 or M1/2 (C), left P4 or M1/2 (D), left M3 (E), right P4 or M1/2 (F), right M3 (G), right m3 (H). Icochilus anomalus, detail of photograph of the holotype MACN-A 3634 from Scotts catalogue (I). Icochilus truncus, holotype MACN-A 3635, right fragmented mandible with alveoli of i1dp1 and complete p2m1, in occlusal (J) and lateral (K) views. Icochilus crassiramis, holotype MACN-A 3636, right mandibular fragment with i2m3, in occlusal (L) and lateral (M) views. Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617767/files/figure.png" pageId="18" pageNumber="213">Figs. 5</figureCitation>
JK), right fragmented mandible with alveoli of
<collectingDate id="5C2AE9A9FFA8287A761ACFEDFBD7FE09" box="[1094,1113,402,427]" pageId="18" pageNumber="213">
<date id="4C6E1041FFA8287A761ACFEDFBD7FE09" box="[1094,1113,402,427]" pageId="18" pageNumber="213">i1</date>
</collectingDate>
dp1 and the series p2m1.
</materialsCitation>
</paragraph>
<paragraph id="386F3681FFA8287A729BCFC9FEAFFE51" blockId="18.[151,1437,402,931]" pageId="18" pageNumber="213">
<emphasis id="0AA4EA93FFA8287A729BCFC9FD5DFE6D" bold="true" box="[199,723,438,463]" pageId="18" pageNumber="213">
Age and provenance of the
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<typeStatus id="E76B8823FFA8287A707BCFC9FDD4FE6D" box="[551,602,438,463]" pageId="18" pageNumber="213">type</typeStatus>
material.
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</emphasis>
Santacrucian SALMA, early
<location id="3D0F605AFFA8287A766ACFC9FB15FE6D" LSID="urn:lsid:plazi:treatment:B0798797FFA8287A72CBCF33FB04FC01:3D0F605AFFA8287A766ACFC9FB15FE6D" box="[1078,1179,438,463]" country="Argentina" name="Miocene" pageId="18" pageNumber="213" stateProvince="Santa Cruz">Miocene</location>
,
<collectingRegion id="FA14F863FFA8287A76F7CFC8FA12FE6D" box="[1195,1436,438,463]" country="Argentina" name="Santa Cruz" pageId="18" pageNumber="213">Santa Cruz Province</collectingRegion>
(
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).
</paragraph>
<paragraph id="386F3681FFA8287A729BCF81FB73FDFD" blockId="18.[151,1437,402,931]" pageId="18" pageNumber="213">
<emphasis id="0AA4EA93FFA8287A729BCF81FE36FDB5" bold="true" box="[199,440,510,535]" pageId="18" pageNumber="213">Original description</emphasis>
(
<bibRefCitation id="5C414B70FFA8287A739BCF81FD36FDB5" author="Ameghino, F." box="[455,696,510,535]" pageId="18" pageNumber="213" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino 1894a: 16</bibRefCitation>
). “
<emphasis id="0AA4EA93FFA8287A7084CF80FBD0FDB5" box="[728,1118,510,535]" italics="true" pageId="18" pageNumber="213">A peu près de la même taille que l</emphasis>
<taxonomicName id="FFD04D02FFA8287A7601CF80FB5BFDB4" authorityName="Ameghino" authorityYear="1889" box="[1117,1237,511,534]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="18" pageNumber="213" phylum="Chordata" rank="species" species="extensus">I. extensus</taxonomicName>
.
<emphasis id="0AA4EA93FFA8287A76BFCF81FB67FDFD" italics="true" pageId="18" pageNumber="213">Elle se distingue par la présence de la première prémolaire inférieure très petite, et par l'absence de la canine inférieure. A la place de la canine il y a un diastème assez étendu qui sépare l'incisive externe de la première prémolaire</emphasis>
”.
</paragraph>
<paragraph id="386F3681FFA8287A729BCC15FC0AFD69" blockId="18.[151,1437,402,931]" pageId="18" pageNumber="213">
English translation. Size almost the same as
<taxonomicName id="FFD04D02FFA8287A70E8CC13FCB8FD21" authorityName="Ameghino" authorityYear="1889" box="[692,822,620,643]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="18" pageNumber="213" phylum="Chordata" rank="species" species="extensus">
<emphasis id="0AA4EA93FFA8287A70E8CC13FCB8FD21" box="[692,822,620,643]" italics="true" pageId="18" pageNumber="213">Ic. extensus</emphasis>
</taxonomicName>
. It is distinguished by the presence of a very small first lower premolar, and by the absence of the lower canine. In the place of the lower canine there is a rather extensive diastema that separates the external incisor from the first premolar.
</paragraph>
</subSubSection>
<subSubSection id="70CA650AFFA8287A729BCCA9FB04FC01" pageId="18" pageNumber="213" type="discussion">
<paragraph id="386F3681FFA8287A729BCCA9FB04FC01" blockId="18.[151,1437,402,931]" pageId="18" pageNumber="213">
<emphasis id="0AA4EA93FFA8287A729BCCA9FEC1FD4D" bold="true" box="[199,335,726,751]" pageId="18" pageNumber="213">Comments.</emphasis>
<bibRefCitation id="5C414B70FFA8287A730ACCA9FDA5FD4D" author="Ameghino, F." box="[342,555,726,751]" pageId="18" pageNumber="213" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
did not add any illustration to the brief description of
<taxonomicName id="FFD04D02FFA8287A76D2CCA8FB74FD4C" authorityName="Ameghino" authorityYear="1894" box="[1166,1274,727,750]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="18" pageNumber="213" phylum="Chordata" rank="species" species="truncus">
<emphasis id="0AA4EA93FFA8287A76D2CCA8FB74FD4C" box="[1166,1274,727,750]" italics="true" pageId="18" pageNumber="213">I. truncus</emphasis>
</taxonomicName>
. According to Ameghinos catalogue and
<bibRefCitation id="5C414B70FFA8287A73B9CC84FD01FCB1" author="Mones A." box="[485,655,762,787]" pageId="18" pageNumber="213" pagination="1 - 625" refId="ref17293" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article" year="1986">Mones (1986)</bibRefCitation>
, MACN-A 3635, a broken symphysis with a fragmented right mandibular ramus, is the type material of the species. This specimen has a small alveolus of dp1 and a diastema between
<date id="4C6E1041FFA8287A72A7CD3DFE9EFCF9" box="[251,272,834,859]" pageId="18" pageNumber="213">i3</date>
and dp1, whose length (
<quantity id="FF289B64FFA8287A707DCD3CFDF4FCF9" box="[545,634,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="213" unit="mm" value="2.2">2.2 mm</quantity>
) falls within the range (
<quantity id="FF289B64FFA8287A71DACD3CFC79FCF9" box="[902,1015,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="18" pageNumber="213" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1 to 3 mm</quantity>
) that Ameghino provided for any gap between adjacent teeth (
<bibRefCitation id="5C414B70FFA8287A73F6CD19FD44FCDD" author="Ameghino, F." box="[426,714,870,895]" pageId="18" pageNumber="213" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino 1894a: 1518</bibRefCitation>
). These characteristics are consistent with
<bibRefCitation id="5C414B70FFA8287A76EFCD19FA12FCDD" author="Ameghino, F." box="[1203,1436,870,895]" pageId="18" pageNumber="213" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghinos (1894a)</bibRefCitation>
description and, consequently, MACN-A 3635 is confirmed as the
<typeStatus id="E76B8823FFA8287A71DFCDF5FC6BFC01" box="[899,997,906,931]" pageId="18" pageNumber="213" type="holotype">holotype</typeStatus>
of
<emphasis id="0AA4EA93FFA8287A7657CDF3FB04FC01" box="[1035,1162,908,931]" italics="true" pageId="18" pageNumber="213">
<taxonomicName id="FFD04D02FFA8287A7657CDF3FB0BFC01" authorityName="Ameghino" authorityYear="1894" box="[1035,1157,908,931]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="18" pageNumber="213" phylum="Chordata" rank="species" species="truncus">Ic. truncus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
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