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<document id="A5DD1FB58182701ABEFABA1B2F55BC9F" ID-CLB-Dataset="27706" ID-DOI="10.11646/zootaxa.4543.2.2" ID-GBIF-Dataset="7438332f-a925-43aa-b17f-059781bdfc12" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617757" ID-ZooBank="11F546CD-109F-4CBC-B87D-EA8D1AD0B96F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928261775" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos &amp; Cerdeño, Esperanza" docDate="2019" docId="B0798797FFA8287A72CBCD89FE45F91B" docLanguage="en" docName="zootaxa.4543.2.2.pdf" docOrigin="Zootaxa 4543 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Icochilus crassiramis Ameghino 1894" docType="treatment" docVersion="8" lastPageNumber="213" masterDocId="4C40FFEFFFBA2868725CCE7FFF8EFFA2" masterDocTitle="On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina" masterLastPageNumber="220" masterPageNumber="195" pageNumber="213" updateTime="1701997211094" updateUser="plazi">
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<mods:title id="FBA640D4EA75C3A626221FDB3B64CAEC">On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina</mods:title>
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<mods:namePart id="944842B2BDA2A212504C615503E1AB19">Fernicola, Juan Carlos</mods:namePart>
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<treatment id="B0798797FFA8287A72CBCD89FE45F91B" ID-DOI="http://doi.org/10.5281/zenodo.5931305" ID-GBIF-Taxon="154529773" ID-Zenodo-Dep="5931305" LSID="urn:lsid:plazi:treatment:B0798797FFA8287A72CBCD89FE45F91B" httpUri="http://treatment.plazi.org/id/B0798797FFA8287A72CBCD89FE45F91B" lastPageNumber="213" pageId="18" pageNumber="213">
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<paragraph id="386F3681FFA8287A72CBCD89FDF6FBB2" blockId="18.[151,632,1014,1040]" box="[151,632,1014,1040]" pageId="18" pageNumber="213">
<heading id="632781EDFFA8287A72CBCD89FDF6FBB2" bold="true" box="[151,632,1014,1040]" fontSize="11" level="1" pageId="18" pageNumber="213" reason="1">
<taxonomicName id="FFD04D02FFA8287A72CBCD89FDF6FBB2" authority="Ameghino, 1894 a" authorityName="Ameghino" authorityYear="1894" box="[151,632,1014,1040]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="18" pageNumber="213" phylum="Chordata" rank="species" species="crassiramis">
<emphasis id="0AA4EA93FFA8287A72CBCD89FDF6FBB2" bold="true" box="[151,632,1014,1040]" pageId="18" pageNumber="213">
<emphasis id="0AA4EA93FFA8287A72CBCD89FE17FBB2" bold="true" box="[151,409,1014,1040]" italics="true" pageId="18" pageNumber="213">Icochilus crassiramis</emphasis>
<bibRefCitation id="5C414B70FFA8287A73FCCD89FDF6FBB2" author="Ameghino, F." box="[416,632,1014,1040]" pageId="18" pageNumber="213" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino, 1894a</bibRefCitation>
</emphasis>
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<subSubSection id="70CA650AFFA8287A72CBCA42FB6AFA40" pageId="18" pageNumber="213" type="materials_examined">
<paragraph id="386F3681FFA8287A72CBCA42FBA5FBF4" blockId="18.[151,1437,1085,1722]" box="[151,1067,1085,1110]" pageId="18" pageNumber="213">
<materialsCitation id="88B83CDCFFA8287A72CBCA42FBA5FBF4" ID-GBIF-Occurrence="2147525032" box="[151,1067,1085,1110]" collectingDate="i 2" collectionCode="MACN-A" pageId="18" pageNumber="213" specimenCode="MACN-A 3636" specimenCount="1" typeStatus="holotype">
<emphasis id="0AA4EA93FFA8287A72CBCA42FE85FBF4" bold="true" box="[151,267,1085,1110]" pageId="18" pageNumber="213">
<typeStatus id="E76B8823FFA8287A72CBCA42FE88FBF4" box="[151,262,1085,1110]" pageId="18" pageNumber="213" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="68769EFAFFA8287A734ECA41FE4CFBF4" box="[274,450,1085,1110]" collectionCode="MACN-A" pageId="18" pageNumber="213">MACN-A 3636</specimenCode>
(
<figureCitation id="A0EB2A04FFA8287A738ECA42FD91FBF4" box="[466,543,1085,1110]" captionStart="FIGURE 5" captionStartId="17.[151,249,1387,1409]" captionTargetBox="[159,1432,215,1349]" captionTargetId="figure@17.[151,1437,193,1366]" captionTargetPageId="17" captionText="FIGURE 5. Icochilus lamellosus, holotype MACN-A 3631, incomplete skull, in ventral view (A). Icochilus trilineatus, MACN-A 3633, left M2, in occlusal view (B); MACN-A 3651 to MACN-A 3657 in occlusal view (CH): left P4 or M1/2 (C), left P4 or M1/2 (D), left M3 (E), right P4 or M1/2 (F), right M3 (G), right m3 (H). Icochilus anomalus, detail of photograph of the holotype MACN-A 3634 from Scotts catalogue (I). Icochilus truncus, holotype MACN-A 3635, right fragmented mandible with alveoli of i1dp1 and complete p2m1, in occlusal (J) and lateral (K) views. Icochilus crassiramis, holotype MACN-A 3636, right mandibular fragment with i2m3, in occlusal (L) and lateral (M) views. Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617767/files/figure.png" pageId="18" pageNumber="213">Figs. 5</figureCitation>
LM), right mandibular fragment with
<collectingDate id="5C2AE9A9FFA8287A7180CA42FC7CFBF4" box="[988,1010,1085,1110]" pageId="18" pageNumber="213">
<date id="4C6E1041FFA8287A7180CA42FC7CFBF4" box="[988,1010,1085,1110]" pageId="18" pageNumber="213">i2</date>
</collectingDate>
m3.
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</paragraph>
<paragraph id="386F3681FFA8287A729BCA1EFEAFFB3C" blockId="18.[151,1437,1085,1722]" pageId="18" pageNumber="213">
<emphasis id="0AA4EA93FFA8287A729BCA1EFD5DFBD8" bold="true" box="[199,723,1121,1146]" pageId="18" pageNumber="213">
Age and provenance of the
<materialsCitation id="88B83CDCFFA8287A707BCA1EFD5DFBD8" ID-GBIF-Occurrence="2147524932" box="[551,723,1121,1146]" collectionCode="SALMA" country="Argentina" location="Miocene" pageId="18" pageNumber="213" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<typeStatus id="E76B8823FFA8287A707BCA1EFDD4FBD8" box="[551,602,1121,1146]" pageId="18" pageNumber="213">type</typeStatus>
material.
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</emphasis>
Santacrucian SALMA, early
<location id="3D0F605AFFA8287A766ACA1EFB15FBD8" LSID="urn:lsid:plazi:treatment:B0798797FFA8287A72CBCD89FE45F91B:3D0F605AFFA8287A766ACA1EFB15FBD8" box="[1078,1179,1121,1146]" country="Argentina" name="Miocene" pageId="18" pageNumber="213" stateProvince="Santa Cruz">Miocene</location>
,
<collectingRegion id="FA14F863FFA8287A76F7CA1DFA12FBD8" box="[1195,1436,1121,1146]" country="Argentina" name="Santa Cruz" pageId="18" pageNumber="213">Santa Cruz Province</collectingRegion>
(
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).
</paragraph>
<paragraph id="386F3681FFA8287A729BCAD6FCD4FAF0" blockId="18.[151,1437,1085,1722]" pageId="18" pageNumber="213">
<emphasis id="0AA4EA93FFA8287A729BCAD6FE30FB60" bold="true" box="[199,446,1193,1218]" pageId="18" pageNumber="213">Original description</emphasis>
(
<bibRefCitation id="5C414B70FFA8287A738FCAD6FD75FB60" author="Ameghino, F." box="[467,763,1193,1218]" pageId="18" pageNumber="213" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino 1894a: 1617</bibRefCitation>
). “
<emphasis id="0AA4EA93FFA8287A717DCAD5FBDCFB60" box="[801,1106,1193,1218]" italics="true" pageId="18" pageNumber="213">De la même taille que 1</emphasis>
<taxonomicName id="FFD04D02FFA8287A760FCAD5FB5FFB63" authorityName="Ameghino" authorityYear="1889" box="[1107,1233,1194,1217]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="18" pageNumber="213" phylum="Chordata" rank="species" species="extensus">I. extensus</taxonomicName>
.
<emphasis id="0AA4EA93FFA8287A76B9CAD5FCC8FAF3" italics="true" pageId="18" pageNumber="213">Cette espèce se distingue très bien par la première prémolaire inférieure qui a la forme d'une canine bien développée, étant isolée en avant et en arrière par des diastèmes assez longs. La canine inférieure a la forme d'une incisive, étant couchée en avant sur l'incisive externe. La deuxième prémolaire inférieure est elliptique, sans sillon perpendiculaire interne; de celui du côté externe on en voit à peine des traces</emphasis>
”.
</paragraph>
<paragraph id="386F3681FFA8287A729BCB22FB6AFA40" blockId="18.[151,1437,1085,1722]" pageId="18" pageNumber="213">
English translation. Same size as
<taxonomicName id="FFD04D02FFA8287A7007CB21FD6AFAD7" authorityName="Ameghino" authorityYear="1889" box="[603,740,1374,1397]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="18" pageNumber="213" phylum="Chordata" rank="species" species="extensus">
<emphasis id="0AA4EA93FFA8287A7007CB21FD6AFAD7" box="[603,740,1374,1397]" italics="true" pageId="18" pageNumber="213">Ic. extensus</emphasis>
</taxonomicName>
. This species is very well distinguished by its first lower premolar that has the morphology of a well-developed canine, being anteriorly and posteriorly isolated by rather long diastemata. The lower canine is incisor-shaped, anteriorly leaned on the external incisor. The second lower premolar is elliptical, without internal perpendicular groove; no trace of external groove is visible.
</paragraph>
</subSubSection>
<subSubSection id="70CA650AFFA8287A729BCB92FE45F91B" pageId="18" pageNumber="213" type="discussion">
<paragraph id="386F3681FFA8287A729BCB92FE45F91B" blockId="18.[151,1437,1085,1722]" pageId="18" pageNumber="213">
<emphasis id="0AA4EA93FFA8287A729BCB92FEC1F9A4" bold="true" box="[199,335,1517,1542]" pageId="18" pageNumber="213">Comments.</emphasis>
<bibRefCitation id="5C414B70FFA8287A7307CB92FDBAF9A4" author="Ameghino, F." box="[347,564,1517,1542]" pageId="18" pageNumber="213" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
did not illustrate any specimen of this species. According to Ameghinos catalogue and
<bibRefCitation id="5C414B70FFA8287A731FC86DFE69F988" author="Mones A." box="[323,487,1553,1578]" pageId="18" pageNumber="213" pagination="1 - 625" refId="ref17293" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article" year="1986">Mones (1986)</bibRefCitation>
, MACN-A 3636, a right mandibular ramus, is the type specimen. This material matches the original description (
<bibRefCitation id="5C414B70FFA8287A7057C84AFD41F9EC" author="Ameghino, F." box="[523,719,1589,1614]" pageId="18" pageNumber="213" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino 1894a</bibRefCitation>
) because it exhibits a canine-like dp1 separated by anterior (L =
<quantity id="FF289B64FFA8287A72CBC826FF7EF9D0" box="[151,240,1625,1650]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="213" unit="mm" value="1.5">1.5 mm</quantity>
) and posterior (L =
<quantity id="FF289B64FFA8287A738AC825FDA1F9D0" box="[470,559,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="213" unit="mm" value="1.2">1.2 mm</quantity>
) diastemata, and an elliptical p2 with very shallow ectoflexid and entoflexid. These characteristics match
<bibRefCitation id="5C414B70FFA8287A7384C802FD4EF934" author="Ameghino, F." box="[472,704,1661,1686]" pageId="18" pageNumber="213" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghinos (1894a)</bibRefCitation>
description and, as a result, MACN-A 3636 is confirmed as the
<typeStatus id="E76B8823FFA8287A72CBC8DEFF77F918" box="[151,249,1697,1722]" pageId="18" pageNumber="213" type="holotype">holotype</typeStatus>
of
<taxonomicName id="FFD04D02FFA8287A7342C8DDFE4BF91B" authorityName="Ameghino" authorityYear="1894" box="[286,453,1698,1721]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="18" pageNumber="213" phylum="Chordata" rank="species" species="crassiramis">
<emphasis id="0AA4EA93FFA8287A7342C8DDFE4BF91B" box="[286,453,1698,1721]" italics="true" pageId="18" pageNumber="213">Ic. crassiramis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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