treatments-xml/data/A1/4C/D5/A14CD52B1746FFB1FF56FB4FFE3C54E3.xml
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<document id="5E5C1FC0070AD093803388599C97A5FD" ID-DOI="10.5281/zenodo.210632" ID-GBIF-Dataset="d241173d-b901-4ba7-878d-04546e8d1200" ID-ISSN="1175-5326" ID-Zenodo-Dep="210632" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460503317629" checkinUser="plazi" docAuthor="Ocampo, Federico C. &amp; Mondaca, José" docDate="2012" docId="A14CD52B1746FFB1FF56FB4FFE3C54E3" docLanguage="en" docName="zt03409p029.pdf" docOrigin="Zootaxa 3409" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Aclopinae" docType="treatment" docVersion="9" lastPageNumber="3" masterDocId="5D75AD531744FFB3FFC1FF82FF94523F" masterDocTitle="Revision of the scarab subfamily Aclopinae Blanchard (Coleoptera: Scarabaeidae) in Argentina and Chile" masterLastPageNumber="29" masterPageNumber="1" pageNumber="3" updateTime="1698312366013" updateUser="plazi">
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<mods:title id="C5630ED3C7CD816C4905E5E5980FE48F">Revision of the scarab subfamily Aclopinae Blanchard (Coleoptera: Scarabaeidae) in Argentina and Chile</mods:title>
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<mods:namePart id="8AFC58983C989901417F7B0E1A3A3178">Ocampo, Federico C.</mods:namePart>
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<mods:namePart id="A5C570707F0E00332244C51DBC51FAB6">Mondaca, José</mods:namePart>
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<mods:date id="DBE56F9BA5DA8FD419340894DCEF3C33">2012</mods:date>
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<treatment id="A14CD52B1746FFB1FF56FB4FFE3C54E3" ID-DOI="http://doi.org/10.5281/zenodo.6167162" ID-GBIF-Taxon="119522185" ID-Zenodo-Dep="6167162" LSID="urn:lsid:plazi:treatment:A14CD52B1746FFB1FF56FB4FFE3C54E3" httpUri="http://treatment.plazi.org/id/A14CD52B1746FFB1FF56FB4FFE3C54E3" lastPageNumber="3" pageId="2" pageNumber="3">
<subSubSection id="61FF37B61746FFB1FF56FB4FFE3C54E3" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="295A643D1746FFB1FF56FB4FFE0856D8" blockId="2.[151,412,1229,1255]" box="[151,412,1229,1255]" pageId="2" pageNumber="3">
<heading id="7212D3511746FFB1FF56FB4FFE0856D8" bold="true" box="[151,412,1229,1255]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="1B91B82F1746FFB1FF56FB4FFE0856D8" bold="true" box="[151,412,1229,1255]" pageId="2" pageNumber="3">
Subfamily
<taxonomicName id="EEE51FBE1746FFB1FEDEFB4FFE0856D8" ID-CoL="J3K" box="[287,412,1229,1255]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Aclopinae">Aclopinae</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="295A643D1746FFB1FF56FA97FE3C54E3" blockId="2.[151,1436,1300,1756]" pageId="2" pageNumber="3">
Recent studies indicate that the
<taxonomicName id="EEE51FBE1746FFB1FDD1FA96FD175713" box="[528,643,1300,1324]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Aclopinae">Aclopinae</taxonomicName>
as previously considered does not constitute a monophyletic group (Smith
<emphasis id="1B91B82F1746FFB1FF2AFABBFE8A576F" box="[235,286,1336,1360]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. 2006;
<bibRefCitation id="4D7419CC1746FFB1FEB4FAB5FDC85770" author="Ocampo" box="[373,604,1335,1360]" pageId="2" pageNumber="3" refString="Ocampo, F. C., Ruiz-Manzanos E., &amp; Marvaldi A. E. (2010) Systematic revision, cladistics, and biogeography of the genus Neogutierrezia Martinez 1953 (Coleoptera: Scarabaeidae), and its phylogenetic placement in Rutelinae based on structural alignment of 28 S rDNA sequences. Invertebrate Systematics, 24, 81 - 111." type="journal article" year="2010">
Ocampo
<emphasis id="1B91B82F1746FFB1FE1CFABBFD84576F" box="[477,528,1336,1360]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. 2010
</bibRefCitation>
). In a recent publication,
<bibRefCitation id="4D7419CC1746FFB1FCBFFAB5FA905770" author="Ocampo" box="[894,1284,1335,1359]" pageId="2" pageNumber="3" refString="Ocampo, F. C. &amp; Vaz-de-Mello, F. (2008) The genus Xenaclopus Arrow (Coleoptera: Scarabaeidae): redescription and removal from the Aclopinae, with systematic notes. Zootaxa, 1916, 57 - 62." type="journal article" year="2008">Ocampo and Vaz-de-Mello (2008)</bibRefCitation>
excluded the Bornean genus
<taxonomicName id="EEE51FBE1746FFB1FE87FADEFE5D574B" box="[326,457,1372,1396]" class="Insecta" family="Melolonthidae" genus="Xenaclopus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="1B91B82F1746FFB1FE87FADEFE5D574B" box="[326,457,1372,1396]" italics="true" pageId="2" pageNumber="3">Xenaclopus</emphasis>
</taxonomicName>
from the
<taxonomicName id="EEE51FBE1746FFB1FDF8FADEFD38574B" box="[569,684,1372,1396]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Aclopinae">Aclopinae</taxonomicName>
and placed it within
<taxonomicName id="EEE51FBE1746FFB1FC5AFADEFBAA574B" box="[923,1086,1372,1396]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Melolonthinae">Melolonthinae</taxonomicName>
<emphasis id="1B91B82F1746FFB1FB84FADFFB75574B" box="[1093,1249,1372,1396]" italics="true" pageId="2" pageNumber="3">incertae sedis</emphasis>
, because further studies are needed to reliably place this genus in the context of the melolonthine classification. Although the protruding labrum and mandibles are characters shared with the other genera currently placed in the subfamily
<taxonomicName id="EEE51FBE1746FFB1FA87FA26FF5557E0" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Aclopinae">Aclopinae</taxonomicName>
, there is no evidence that supports that these characters are synapomorphic and are likely due to convergence. As a result of this study, we have removed both
<taxonomicName id="EEE51FBE1746FFB1FD10FA6EFCE0543B" box="[721,884,1516,1540]" class="Insecta" family="Aclopidae" genus="Phaenognatha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="1B91B82F1746FFB1FD10FA6EFCE0543B" box="[721,884,1516,1540]" italics="true" pageId="2" pageNumber="3">Phaenognatha</emphasis>
</taxonomicName>
and
<taxonomicName id="EEE51FBE1746FFB1FC73FA6EFBEB543B" box="[946,1151,1516,1540]" class="Insecta" family="Aclopidae" genus="Neophaenognatha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="1B91B82F1746FFB1FC73FA6EFBEB543B" box="[946,1151,1516,1540]" italics="true" pageId="2" pageNumber="3">Neophaenognatha</emphasis>
</taxonomicName>
from the
<taxonomicName id="EEE51FBE1746FFB1FB37FA6EFAFD543B" box="[1270,1385,1516,1540]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Aclopinae">Aclopinae</taxonomicName>
and place them in the tribe Phaenognathini within the subfamily
<taxonomicName id="EEE51FBE1746FFB1FC97F98DFC695418" box="[854,1021,1551,1575]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Melolonthinae">Melolonthinae</taxonomicName>
. The relationships of this tribe with other melolonthine tribes are uncertain. The most obvious difference between these genera and aclopines is the presence of a well developed genital capsule (the genital capsule is absent in aclopines). These data, along with morphological evidence, supports the hypothesis of a relationship between
<taxonomicName id="EEE51FBE1746FFB1FC30F9FEFB0154AB" box="[1009,1173,1660,1684]" class="Insecta" family="Aclopidae" genus="Phaenognatha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="1B91B82F1746FFB1FC30F9FEFB0154AB" box="[1009,1173,1660,1684]" italics="true" pageId="2" pageNumber="3">Phaenognatha</emphasis>
</taxonomicName>
and
<taxonomicName id="EEE51FBE1746FFB1FB0FF9FEFA0F54AB" box="[1230,1435,1660,1684]" class="Insecta" family="Aclopidae" genus="Neophaenognatha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="1B91B82F1746FFB1FB0FF9FEFA0F54AB" box="[1230,1435,1660,1684]" italics="true" pageId="2" pageNumber="3">Neophaenognatha</emphasis>
</taxonomicName>
and the Chilean Lichniini (currently in
<taxonomicName id="EEE51FBE1746FFB1FD9BF91DFC965488" box="[602,770,1695,1719]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Melolonthinae">Melolonthinae</taxonomicName>
), and they together might constitute a distinct and ancient lineage of
<taxonomicName id="EEE51FBE1746FFB1FECCF946FE3154E3" box="[269,421,1732,1756]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
.
</paragraph>
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