455 lines
51 KiB
XML
455 lines
51 KiB
XML
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<mods:title id="B3747FE98233DC4F89D95C58F777517A">Clostophis Benson, 1860, is not a monotypic diplommatinid but a speciose hypselostomatid (Gastropoda: Eupulmonata), with descriptions of six new species</mods:title>
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<mods:namePart id="CD6CBFBC8D2B2CF7DC5B872CE0DDDD23">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="92ABCD5FB10E75E17504EBC81C59E2B2">Hunyadi, András</mods:namePart>
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<mods:namePart id="29A81E4C2906EFEE97DAC487755D625B">Grego, Jozef</mods:namePart>
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<mods:namePart id="8F212CFB4E776A48A912256F5794D2ED">Reischütz, Alexander</mods:namePart>
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<mods:namePart id="14251F2AAF170F6D41C08494EC95E601">Buczkó, Krisztina</mods:namePart>
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<mods:namePart id="A91DB51A380A1B48E1791B779EA64D2F">Vermeulen, Jaap J.</mods:namePart>
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<mods:title id="75D501D3F5FB023CBD05A16E806E003C">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="7807DCD0811A2D1C3939EC832116DD07">2020</mods:date>
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<taxonomicName id="DACC1ACAFFC5FFC5FCDB5D34FA4C9C59" ID-CoL="7QQQ3" authority="Pall-Gergely & A. Reischutz" authorityName="Pall-Gergely & A. Reischutz" authorityYear="2020" box="[833,1426,1820,1845]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="358" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
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<emphasis id="2FB8BD5BFFC5FFC5FCDB5D34FBE19C59" bold="true" box="[833,1087,1821,1845]" italics="true" pageId="8" pageNumber="358">Clostophis multiformis</emphasis>
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Páll-Gergely & A. Reischütz
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,
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<paragraph id="1D736149FFC5FFC5FBAA5D74FB769C19" blockId="8.[833,1431,1820,1909]" box="[1072,1192,1885,1909]" pageId="8" pageNumber="358">
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(
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<figureCitation id="85F77DCCFFC5FFC5FBA25D74FB7E9C19" box="[1080,1184,1885,1909]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="10.[113,148,2035,2057]" captionStartId-1="11.[136,171,2035,2057]" captionStartId-2="12.[113,148,2035,2057]" captionTargetBox-0="[148,1415,190,2020]" captionTargetBox-1="[172,1438,189,2020]" captionTargetBox-2="[149,1415,189,2020]" captionTargetId-0="figure-58@10.[148,1415,190,2020]" captionTargetId-1="figure-8@11.[172,1438,189,2020]" captionTargetId-2="figure-58@12.[149,1415,189,2020]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="Fig. 7. Clostophis multiformis Páll-Gergely & A. Reischütz, new species (NHMW 113273, holotype). A, dorsal view; B, ventral view; C, apertural view; D, lateral view; E, aperture dentition; F, protoconch sculpture; G, sculpture of the ventral side; H, sculpture of body whorl from standard apertural view." captionText-1="Fig. 8. Clostophis multiformis Páll-Gergely & A. Reischütz, new species (NHMW 113274, paratype). A, dorsal view; B, ventral view; C, apertural view; D, lateral view; E, aperture dentition; F, protoconch sculpture; G, sculpture of the ventral side; H, sculpture of body whorl from standard apertural view." captionText-2="Fig. 9. Clostophis multiformis Páll-Gergely & A. Reischütz, new species (HNHM 104434, JG4). A, lateral view; B, ventral view; C, apertural view; D, dorsal view; E, protoconch sculpture; F, aperture dentition; G, sculpture of the ventral side; H, sculpture of body whorl from standard apertural view." figureDoi-0="http://doi.org/10.5281/zenodo.4576876" figureDoi-1="http://doi.org/10.5281/zenodo.4576880" figureDoi-2="http://doi.org/10.5281/zenodo.4576882" httpUri-0="https://zenodo.org/record/4576876/files/figure.png" httpUri-1="https://zenodo.org/record/4576880/files/figure.png" httpUri-2="https://zenodo.org/record/4576882/files/figure.png" pageId="8" pageNumber="358">Figs. 7–9</figureCitation>
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)
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</paragraph>
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<emphasis id="2FB8BD5BFFC5FFC5FCB75DB4FC049CD9" bold="true" box="[813,986,1949,1973]" pageId="8" pageNumber="358">Type material.</emphasis>
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<typeStatus id="C277DFEBFFC5FFC5FC7E5DB4FB979CD9" box="[996,1097,1949,1973]" pageId="8" pageNumber="358" type="holotype">Holotype</typeStatus>
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(SH:
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, SW:
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<quantity id="DA34CCACFFC5FFC5FAA55DB4FA7A9CD9" box="[1343,1444,1949,1973]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="8" pageNumber="358" unit="mm" value="1.03">1.03 mm</quantity>
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) (
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<specimenCode id="4D6AC932FFC5FFC5FCAE5D94FC299CB9" box="[820,1015,1981,2005]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="358" type="Museum">NHMW 113273</specimenCode>
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,
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<figureCitation id="85F77DCCFFC5FFC5FB925D94FB8C9CB9" box="[1032,1106,1981,2005]" captionStart="Fig" captionStartId="10.[113,148,2035,2057]" captionTargetBox="[148,1415,190,2020]" captionTargetId="figure-58@10.[148,1415,190,2020]" captionTargetPageId="10" captionText="Fig. 7. Clostophis multiformis Páll-Gergely & A. Reischütz, new species (NHMW 113273, holotype). A, dorsal view; B, ventral view; C, apertural view; D, lateral view; E, aperture dentition; F, protoconch sculpture; G, sculpture of the ventral side; H, sculpture of body whorl from standard apertural view." figureDoi="http://doi.org/10.5281/zenodo.4576876" httpUri="https://zenodo.org/record/4576876/files/figure.png" pageId="8" pageNumber="358">Fig. 7</figureCitation>
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),
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<collectingCountry id="65DB21D9FFC5FFC5FBF45D97FB779CB9" box="[1134,1193,1982,2005]" name="Laos" pageId="8" pageNumber="358">Laos</collectingCountry>
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,
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<collectingRegion id="DF08AFABFFC5FFC5FB235D94FA789CB9" box="[1209,1446,1981,2005]" country="Laos" name="Khammouan" pageId="8" pageNumber="358">Khammouane Prov.</collectingRegion>
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, limestone hill in rice fields east of the road 1E,
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<quantity id="DA34CCACFFC5FFC5FAA75DF4FA5D9C99" box="[1341,1411,2013,2037]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="358" unit="m" value="300.0">
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</quantity>
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SE
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<collectingMunicipality id="FD17FB33FFC5FFC5FCB75DD4FC709379" box="[813,942,2045,2069]" pageId="8" pageNumber="358">Gnommalat</collectingMunicipality>
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,
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<collectorName id="B039049FFFC5FFC5FC2D5DD7FBE19379" box="[951,1087,2045,2069]" pageId="8" pageNumber="358">
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NE
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, at the base of rocks,
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,
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(locality code: La.4), coll.
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<collectorName id="B039049FFFC5FFC5FA955237FA789359" box="[1295,1446,2077,2102]" pageId="8" pageNumber="358">A. Reischütz</collectorName>
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,
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<date id="69724789FFC0FFC0FF125A94FECF9BB9" box="[136,273,189,213]" pageId="13" pageNumber="363" value="2010-03">
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<collectingDate id="7936BE61FFC0FFC0FF125A94FECF9BB9" box="[136,273,189,213]" pageId="13" pageNumber="363" value="2010-03">March 2010</collectingDate>
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</materialsCitation>
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.
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<materialsCitation id="ADA46B14FFC0FFC0FE845A97FECC9A39" collectingDate="2010-03" collectionCode="NHMW" collectorName="La. & A. Reischutz" country="Laos" county="Xe Bangfai river" latitude="17.410133" location="Thakhek" longLatPrecision="1" longitude="105.20198" municipality="Mahaxay" pageId="13" pageNumber="363" specimenCode="NHMW 113274" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
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<typeStatus id="C277DFEBFFC0FFC0FE845A97FE549BBA" box="[286,394,190,214]" pageId="13" pageNumber="363" type="paratype">Paratypes</typeStatus>
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: 1 shell (
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<specimenCode id="4D6AC932FFC0FFC0FE685A94FD769BB9" box="[498,680,189,213]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="363" type="Museum">NHMW 113274</specimenCode>
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,
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<figureCitation id="85F77DCCFFC0FFC0FD2F5A94FD269BB9" box="[693,760,189,213]" captionStart="Fig" captionStartId="11.[136,171,2035,2057]" captionTargetBox="[172,1438,189,2020]" captionTargetId="figure-8@11.[172,1438,189,2020]" captionTargetPageId="11" captionText="Fig. 8. Clostophis multiformis Páll-Gergely & A. Reischütz, new species (NHMW 113274, paratype). A, dorsal view; B, ventral view; C, apertural view; D, lateral view; E, aperture dentition; F, protoconch sculpture; G, sculpture of the ventral side; H, sculpture of body whorl from standard apertural view." figureDoi="http://doi.org/10.5281/zenodo.4576880" httpUri="https://zenodo.org/record/4576880/files/figure.png" pageId="13" pageNumber="363">Fig. 8</figureCitation>
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),
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<collectingCountry id="65DB21D9FFC0FFC0FF125AF7FF609B99" box="[136,190,222,245]" name="Laos" pageId="13" pageNumber="363">Laos</collectingCountry>
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,
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<collectingRegion id="DF08AFABFFC0FFC0FF505AF4FE7D9B99" box="[202,419,221,245]" country="Laos" name="Khammouan" pageId="13" pageNumber="363">Khammouane Prov.</collectingRegion>
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, limestone hill at the
|
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<collectingCounty id="F41219C5FFC0FFC0FD165AF7FF659A79" pageId="13" pageNumber="363">Xe Bangfai river</collectingCounty>
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city bridge from
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<collectingMunicipality id="FD17FB33FFC0FFC0FE1A5AD4FE389A79" box="[384,486,253,277]" pageId="13" pageNumber="363">Mahaxay</collectingMunicipality>
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, E
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<location id="18133792FFC0FFC0FD965AD4FDB29A79" LSID="urn:lsid:plazi:treatment:9565D05FFFC5FFC0FCDB5D34FBBB9FF9:18133792FFC0FFC0FD965AD4FDB29A79" box="[524,620,253,277]" country="Laos" county="Xe Bangfai river" latitude="17.410133" longLatPrecision="1" longitude="105.20198" municipality="Mahaxay" name="Thakhek" pageId="13" pageNumber="363" stateProvince="Khammouan">Thakhek</location>
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,
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,
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(locality code:
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<collectorName id="B039049FFFC0FFC0FE455B34FDD99A59" box="[479,519,285,309]" pageId="13" pageNumber="363">La.</collectorName>
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8), coll.
|
||
<collectorName id="B039049FFFC0FFC0FDF05B37FCDF9A59" box="[618,769,285,310]" pageId="13" pageNumber="363">A. Reischütz</collectorName>
|
||
,
|
||
<date id="69724789FFC0FFC0FF135B14FECC9A39" box="[137,274,317,341]" pageId="13" pageNumber="363" value="2010-03">
|
||
<collectingDate id="7936BE61FFC0FFC0FF135B14FECC9A39" box="[137,274,317,341]" pageId="13" pageNumber="363" value="2010-03">March 2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ADA46B14FFC0FFC0FEBA5B14FF1A9A99" collectingDate="2019-07-19" collectionCode="SA" collectorName="S. Aiken" country="Laos" county="Xe Bangfai river" latitude="17.580984" location="2 km N of Gnommalath" longLatPrecision="1" longitude="105.16818" municipality="Mahaxay" pageId="13" pageNumber="363" specimenCode="SA1" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
|
||
2 shells (coll. RE), same data as preceding; 1 shell (coll.
|
||
<collectionCode id="7BDDF98CFFC0FFC0FE845B74FE9A9A19" box="[286,324,349,373]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="13" pageNumber="363">SA</collectionCode>
|
||
),
|
||
<collectingCountry id="65DB21D9FFC0FFC0FEC05B77FE4F9A19" box="[346,401,350,373]" name="Laos" pageId="13" pageNumber="363">Laos</collectingCountry>
|
||
,
|
||
<collectingRegion id="DF08AFABFFC0FFC0FE055B74FD7C9A19" box="[415,674,349,373]" country="Laos" name="Khammouan" pageId="13" pageNumber="363">Khammouane Province</collectingRegion>
|
||
,
|
||
<location id="18133792FFC0FFC0FD2A5B74FEE49AF9" LSID="urn:lsid:plazi:treatment:9565D05FFFC5FFC0FCDB5D34FBBB9FF9:18133792FFC0FFC0FD2A5B74FEE49AF9" country="Laos" latitude="17.580984" longLatPrecision="1" longitude="105.16818" name="2 km N of Gnommalath" pageId="13" pageNumber="363" stateProvince="Khammouan">
|
||
<quantity id="DA34CCACFFC0FFC0FD2A5B74FD349A19" box="[688,746,349,373]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="363" unit="km" value="2.0">2 km</quantity>
|
||
N of Gnommalath
|
||
</location>
|
||
, in dry soil on rock at foot of limestone hill, near the weir of powerplant channel,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FDE95BB5FCDC9AD9" box="[627,770,412,437]" degrees="17" direction="north" minutes="34.859" orientation="latitude" pageId="13" pageNumber="363" precision="1" value="17.580984">17°34.859′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FF135B95FEC49AB9" box="[137,282,444,469]" degrees="105" direction="east" minutes="10.091" orientation="longitude" pageId="13" pageNumber="363" precision="1" value="105.16818">105°10.091′E</geoCoordinate>
|
||
(locality code:
|
||
<specimenCode id="4D6AC932FFC0FFC0FE525B94FE249AB9" box="[456,506,445,469]" collectionCode="SA" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="13" pageNumber="363">SA1</specimenCode>
|
||
), coll.
|
||
<collectorName id="B039049FFFC0FFC0FDD25B94FD749AB9" box="[584,682,445,469]" pageId="13" pageNumber="363">S. Aiken</collectorName>
|
||
,
|
||
<date id="69724789FFC0FFC0FD2D5B94FF1A9A99" pageId="13" pageNumber="363" value="2019-07-19">
|
||
<collectingDate id="7936BE61FFC0FFC0FD2D5B94FF1A9A99" pageId="13" pageNumber="363" value="2019-07-19">19 July 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ADA46B14FFC0FFC0FF4E5BF4FF1C99D9" collectingDate="2007-11-25" collectionCode="V" collectorName="A. Abdou & Muratov" country="Laos" county="South-Central" elevation="150" latitude="17.4326" location="Muang" longLatPrecision="1" longitude="105.16115" municipality="Thakhek" pageId="13" pageNumber="363" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
|
||
3 shells (MNHN-IM-2014-6404),
|
||
<collectingCounty id="F41219C5FFC0FFC0FDFC5BF4FCDB9A99" box="[614,773,477,501]" pageId="13" pageNumber="363">South-Central</collectingCounty>
|
||
<collectingCountry id="65DB21D9FFC0FFC0FF125BD7FF609979" box="[136,190,510,533]" name="Laos" pageId="13" pageNumber="363">Laos</collectingCountry>
|
||
,
|
||
<collectingRegion id="DF08AFABFFC0FFC0FF565BD4FE1C9979" box="[204,450,509,533]" country="Laos" name="Khammouan" pageId="13" pageNumber="363">Khammouan province</collectingRegion>
|
||
, ca.
|
||
<quantity id="DA34CCACFFC0FFC0FE635BD4FD9E9979" box="[505,576,509,533]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="13" pageNumber="363" unit="km" value="37.0">37 km</quantity>
|
||
ENE of
|
||
<collectingMunicipality id="FD17FB33FFC0FFC0FD335BD4FCD99979" box="[681,775,509,533]" pageId="13" pageNumber="363">Thakhek</collectingMunicipality>
|
||
(
|
||
<location id="18133792FFC0FFC0FF155834FF029959" LSID="urn:lsid:plazi:treatment:9565D05FFFC5FFC0FCDB5D34FBBB9FF9:18133792FFC0FFC0FF155834FF029959" box="[143,220,541,565]" country="Laos" county="South-Central" latitude="17.4326" longLatPrecision="1" longitude="105.16115" municipality="Thakhek" name="Muang" pageId="13" pageNumber="363" stateProvince="Khammouan">Muang</location>
|
||
<collectingRegion id="DF08AFABFFC0FFC0FF7D5834FEA89959" box="[231,374,541,565]" country="Laos" name="Khammouan" pageId="13" pageNumber="363">Khammouan</collectingRegion>
|
||
), ca.
|
||
<locationDeviation id="0CC1786BFFC0FFC0FE2E5834FD009959" box="[436,734,541,565]" location="Mahaxai" pageId="13" pageNumber="363">
|
||
<quantity id="DA34CCACFFC0FFC0FE2E5834FDDC9959" box="[436,514,541,565]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="363" unit="km" value="4.5">4.5 km</quantity>
|
||
WNW of Mahaxai
|
||
</locationDeviation>
|
||
, on and under rocks in dry secondary forest under E exposed cliff,
|
||
<elevation id="96E1867AFFC0FFC0FF535874FE8A9919" box="[201,340,605,629]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="363" unit="m" value="150.0">
|
||
<quantity id="DA34CCACFFC0FFC0FF535874FECD9919" box="[201,275,605,629]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="363" unit="m" value="150.0">150 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FEFF5875FE289919" box="[357,502,604,629]" degrees="17" direction="north" minutes="25.956" orientation="latitude" pageId="13" pageNumber="363" precision="1" value="17.4326">17°25.956′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FD9C5875FD419919" box="[518,671,604,629]" degrees="105" direction="east" minutes="09.669" orientation="longitude" pageId="13" pageNumber="363" precision="1" value="105.16115">105°09.669′E</geoCoordinate>
|
||
(locality code: 3L07), coll.
|
||
<collectorName id="B039049FFFC0FFC0FED45857FE6699F9" box="[334,440,637,662]" pageId="13" pageNumber="363">A. Abdou</collectorName>
|
||
& I.
|
||
<collectionCode id="7BDDF98CFFC0FFC0FE735857FDDF99F9" box="[489,513,638,661]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="363" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="B039049FFFC0FFC0FD915857FDB499F9" box="[523,618,638,661]" pageId="13" pageNumber="363">Muratov</collectorName>
|
||
,
|
||
<date id="69724789FFC0FFC0FDEF5854FF1C99D9" pageId="13" pageNumber="363" value="2007-11-25">
|
||
<collectingDate id="7936BE61FFC0FFC0FDEF5854FF1C99D9" pageId="13" pageNumber="363" value="2007-11-25">25 November 2007</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ADA46B14FFC0FFC0FF5658B4FD529839" collectingDate="2007-11-25" collectionCode="V" collectorName="A. Abdou & Muratov" country="Laos" county="South-Central" elevation="169" latitude="17.4456" location="Muang" longLatPrecision="1" longitude="105.139336" municipality="Thakhek" pageId="13" pageNumber="363" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
|
||
1 shell (MNHN-IM-2014-6403),
|
||
<collectingCounty id="F41219C5FFC0FFC0FDAB58B4FD1899D9" box="[561,710,669,693]" pageId="13" pageNumber="363">South-Central</collectingCounty>
|
||
<collectingCountry id="65DB21D9FFC0FFC0FD5758B7FCDD99D9" box="[717,771,670,693]" name="Laos" pageId="13" pageNumber="363">Laos</collectingCountry>
|
||
,
|
||
<collectingRegion id="DF08AFABFFC0FFC0FF125894FEA499B9" box="[136,378,701,725]" country="Laos" name="Khammouan" pageId="13" pageNumber="363">Khammouan province</collectingRegion>
|
||
, ca.
|
||
<quantity id="DA34CCACFFC0FFC0FE305894FE3099B9" box="[426,494,701,725]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="363" unit="km" value="35.0">35 km</quantity>
|
||
ENE of
|
||
<collectingMunicipality id="FD17FB33FFC0FFC0FDD45894FD7599B9" box="[590,683,701,725]" pageId="13" pageNumber="363">Thakhek</collectingMunicipality>
|
||
(
|
||
<location id="18133792FFC0FFC0FD205894FCD999B9" LSID="urn:lsid:plazi:treatment:9565D05FFFC5FFC0FCDB5D34FBBB9FF9:18133792FFC0FFC0FD205894FCD999B9" box="[698,775,701,725]" country="Laos" county="South-Central" latitude="17.4456" longLatPrecision="1" longitude="105.139336" municipality="Thakhek" name="Muang" pageId="13" pageNumber="363" stateProvince="Khammouan">Muang</location>
|
||
<collectingRegion id="DF08AFABFFC0FFC0FF1358F4FEC99999" box="[137,279,733,757]" country="Laos" name="Khammouan" pageId="13" pageNumber="363">Khammouan</collectingRegion>
|
||
), ca.
|
||
<locationDeviation id="0CC1786BFFC0FFC0FECE58F7FDB99999" box="[340,615,733,757]" location="Mahaxai" pageId="13" pageNumber="363">
|
||
<quantity id="DA34CCACFFC0FFC0FECE58F7FE539999" box="[340,397,733,757]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="363" unit="km" value="7.0">7 km</quantity>
|
||
WNW of Mahaxai
|
||
</locationDeviation>
|
||
, on and under rocks in dry secondary forest under S exposed cliff,
|
||
<elevation id="96E1867AFFC0FFC0FD5858D4FF629859" metricMagnitude="2" metricUnit="m" metricValue="1.69" pageId="13" pageNumber="363" unit="m" value="169.0">
|
||
<quantity id="DA34CCACFFC0FFC0FD5858D4FCD99879" box="[706,775,765,789]" metricMagnitude="2" metricUnit="m" metricValue="1.69" pageId="13" pageNumber="363" unit="m" value="169.0">169 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FF5D5935FE8E9859" box="[199,336,796,821]" degrees="17" direction="north" minutes="26.736" orientation="latitude" pageId="13" pageNumber="363" precision="1" value="17.4456">17°26.736′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FEC05935FE329859" box="[346,492,796,821]" degrees="105" direction="east" minutes="08.360" orientation="longitude" pageId="13" pageNumber="363" precision="1" value="105.139336">105°08.360′E</geoCoordinate>
|
||
(locality code: 4L07), coll.
|
||
<collectorName id="B039049FFFC0FFC0FF125917FF2D9839" box="[136,243,829,854]" pageId="13" pageNumber="363">A. Abdou</collectorName>
|
||
& I.
|
||
<collectionCode id="7BDDF98CFFC0FFC0FEBD5917FEE19839" box="[295,319,830,853]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="363" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="B039049FFFC0FFC0FED15917FE749839" box="[331,426,830,853]" pageId="13" pageNumber="363">Muratov</collectorName>
|
||
,
|
||
<date id="69724789FFC0FFC0FE2C5914FD529839" box="[438,652,829,853]" pageId="13" pageNumber="363" value="2007-11-25">
|
||
<collectingDate id="7936BE61FFC0FFC0FE2C5914FD529839" box="[438,652,829,853]" pageId="13" pageNumber="363" value="2007-11-25">25 November 2007</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="49B331C1FFC4FFC4FF125DDAFE97932E" ID-DOI="http://doi.org/10.5281/zenodo.4576874" ID-Zenodo-Dep="4576874" httpUri="https://zenodo.org/record/4576874/files/figure.png" pageId="9" pageNumber="359" startId="9.[136,171,2035,2057]" targetBox="[172,1438,190,2020]" targetPageId="9">
|
||
<paragraph id="1D736149FFC4FFC4FF125DDAFE97932E" blockId="9.[136,1476,2035,2114]" pageId="9" pageNumber="359">
|
||
Fig. 6.
|
||
<taxonomicName id="DACC1ACAFFC4FFC4FF4A5DDAFD709365" authority="Pall-Gergely & Vermeulen" authorityName="Pall-Gergely & Vermeulen" authorityYear="2020" box="[208,686,2035,2057]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="359" phylum="Mollusca" rank="species" species="koilobasis" status="sp. nov.">
|
||
<emphasis id="2FB8BD5BFFC4FFC4FF4A5DDAFE419364" box="[208,415,2035,2056]" italics="true" pageId="9" pageNumber="359">Clostophis koilobasis</emphasis>
|
||
Páll-Gergely & Vermeulen
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="348B0020FFC4FFC4FD235DDAFCF39365" box="[697,813,2035,2057]" pageId="9" pageNumber="359" rank="species">new species</taxonomicNameLabel>
|
||
(NHMUK 20191126, holotype). A, apertural view; B, dorsal view; C, ventral view; D, lateral view; E–F, apertural dentition; G, protoconch sculpture; H, sculpture of the ventral side; I, sculpture of body whorl (lateral side).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="49B331C1FFC7FFC7FFEB5DDAFE0D932E" ID-DOI="http://doi.org/10.5281/zenodo.4576876" ID-Zenodo-Dep="4576876" httpUri="https://zenodo.org/record/4576876/files/figure.png" pageId="10" pageNumber="360" startId="10.[113,148,2035,2057]" targetBox="[148,1415,190,2020]" targetPageId="10">
|
||
<paragraph id="1D736149FFC7FFC7FFEB5DDAFE0D932E" blockId="10.[113,1451,2035,2114]" pageId="10" pageNumber="360">
|
||
Fig. 7.
|
||
<taxonomicName id="DACC1ACAFFC7FFC7FF205DDAFD1E9365" authority="Pall-Gergely & A. Reischutz" authorityName="Pall-Gergely & A. Reischutz" authorityYear="2020" box="[186,704,2035,2057]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="360" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
|
||
<emphasis id="2FB8BD5BFFC7FFC7FF205DDAFE469364" box="[186,408,2035,2056]" italics="true" pageId="10" pageNumber="360">Clostophis multiformis</emphasis>
|
||
Páll-Gergely & A. Reischütz
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="348B0020FFC7FFC7FD565DDAFC9F9365" box="[716,833,2035,2057]" pageId="10" pageNumber="360" rank="species">new species</taxonomicNameLabel>
|
||
(NHMW 113273, holotype). A, dorsal view; B, ventral view; C, apertural view; D, lateral view; E, aperture dentition; F, protoconch sculpture; G, sculpture of the ventral side; H, sculpture of body whorl from standard apertural view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="49B331C1FFC6FFC6FF125DDAFE34932F" ID-DOI="http://doi.org/10.5281/zenodo.4576880" ID-Zenodo-Dep="4576880" httpUri="https://zenodo.org/record/4576880/files/figure.png" pageId="11" pageNumber="361" startId="11.[136,171,2035,2057]" targetBox="[172,1438,189,2020]" targetPageId="11">
|
||
<paragraph id="1D736149FFC6FFC6FF125DDAFE34932F" blockId="11.[136,1474,2035,2115]" pageId="11" pageNumber="361">
|
||
Fig. 8.
|
||
<taxonomicName id="DACC1ACAFFC6FFC6FF4B5DDAFD099365" authority="Pall-Gergely & A. Reischutz" authorityName="Pall-Gergely & A. Reischutz" authorityYear="2020" box="[209,727,2035,2057]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="361" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
|
||
<emphasis id="2FB8BD5BFFC6FFC6FF4B5DDAFE719364" box="[209,431,2035,2056]" italics="true" pageId="11" pageNumber="361">Clostophis multiformis</emphasis>
|
||
Páll-Gergely & A. Reischütz
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="348B0020FFC6FFC6FD7E5DDDFC879365" box="[740,857,2035,2057]" pageId="11" pageNumber="361" rank="species">new species</taxonomicNameLabel>
|
||
(NHMW 113274, paratype). A, dorsal view; B, ventral view; C, apertural view; D, lateral view; E, aperture dentition; F, protoconch sculpture; G, sculpture of the ventral side; H, sculpture of body whorl from standard apertural view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="49B331C1FFC1FFC1FFEB5DDAFE4A932F" ID-DOI="http://doi.org/10.5281/zenodo.4576882" ID-Zenodo-Dep="4576882" httpUri="https://zenodo.org/record/4576882/files/figure.png" pageId="12" pageNumber="362" startId="12.[113,148,2035,2057]" targetBox="[149,1415,189,2020]" targetPageId="12">
|
||
<paragraph id="1D736149FFC1FFC1FFEB5DDAFE4A932F" blockId="12.[113,1451,2035,2115]" pageId="12" pageNumber="362">
|
||
Fig. 9.
|
||
<taxonomicName id="DACC1ACAFFC1FFC1FF215DDAFD169365" authority="Pall-Gergely & A. Reischutz" authorityName="Pall-Gergely & A. Reischutz" authorityYear="2020" box="[187,712,2035,2057]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="362" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
|
||
<emphasis id="2FB8BD5BFFC1FFC1FF215DDAFE429364" box="[187,412,2035,2056]" italics="true" pageId="12" pageNumber="362">Clostophis multiformis</emphasis>
|
||
Páll-Gergely & A. Reischütz
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="348B0020FFC1FFC1FD4E5DDDFC929365" box="[724,844,2035,2057]" pageId="12" pageNumber="362" rank="species">new species</taxonomicNameLabel>
|
||
(HNHM 104434, JG4). A, lateral view; B, ventral view; C, apertural view; D, dorsal view; E, protoconch sculpture; F, aperture dentition; G, sculpture of the ventral side; H, sculpture of body whorl from standard apertural view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1D736149FFC0FFC0FF125954FEF09E79" blockId="13.[134,776,893,1301]" pageId="13" pageNumber="363">
|
||
<emphasis id="2FB8BD5BFFC0FFC0FF125954FEB298F9" bold="true" box="[136,364,893,917]" pageId="13" pageNumber="363">Additional material.</emphasis>
|
||
<materialsCitation id="ADA46B14FFC0FFC0FEEA5954FEEA9F79" collectingDate="2017-02-11" collectorName="J. Grego" country="Laos" elevation="160" latitude="17.563583" location="Earthquake Dome" longLatPrecision="1" longitude="104.871635" municipality="Tham Nam Don Cave" pageId="13" pageNumber="363" specimenCount="1" stateProvince="Khammouan">
|
||
2 shells (coll. JG),
|
||
<collectingCountry id="65DB21D9FFC0FFC0FDAB5957FDB998F9" box="[561,615,894,917]" name="Laos" pageId="13" pageNumber="363">Laos</collectingCountry>
|
||
,
|
||
<collectingRegion id="DF08AFABFFC0FFC0FDEB5954FF3798D9" country="Laos" name="Khammouan" pageId="13" pageNumber="363">Khammouane Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="FD17FB33FFC0FFC0FF6D59B4FE2E98D9" box="[247,496,925,950]" pageId="13" pageNumber="363">Tham Nam Dôn Cave</collectingMunicipality>
|
||
,
|
||
<location id="18133792FFC0FFC0FE6459B4FD1898D9" LSID="urn:lsid:plazi:treatment:9565D05FFFC5FFC0FCDB5D34FBBB9FF9:18133792FFC0FFC0FE6459B4FD1898D9" box="[510,710,925,949]" country="Laos" latitude="17.563583" longLatPrecision="1" longitude="104.871635" municipality="Tham Nam Don Cave" name="Earthquake Dome" pageId="13" pageNumber="363" stateProvince="Khammouan">Earthquake Dome</location>
|
||
, sand sediments at bank of cave river,
|
||
<elevation id="96E1867AFFC0FFC0FE715994FDB498B9" box="[491,618,957,981]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="363" unit="m" value="160.0">
|
||
<quantity id="DA34CCACFFC0FFC0FE715994FDEE98B9" box="[491,560,957,981]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="363" unit="m" value="160.0">160 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FDE25995FCDD98B9" box="[632,771,956,981]" degrees="17" direction="north" minutes="33.815" orientation="latitude" pageId="13" pageNumber="363" precision="1" value="17.563583">17°33.815′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FF1259F5FEFC9899" box="[136,290,988,1013]" degrees="104" direction="east" minutes="52.298" orientation="longitude" pageId="13" pageNumber="363" precision="1" value="104.871635">104°52.298′E</geoCoordinate>
|
||
(locality code: JG2B), coll.
|
||
<collectorName id="B039049FFFC0FFC0FDEC59F7FD029899" box="[630,732,989,1013]" pageId="13" pageNumber="363">J. Grego</collectorName>
|
||
,
|
||
<date id="69724789FFC0FFC0FD7159F4FEEA9F79" pageId="13" pageNumber="363" value="2017-02-11">
|
||
<collectingDate id="7936BE61FFC0FFC0FD7159F4FEEA9F79" pageId="13" pageNumber="363" value="2017-02-11">11 February 2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ADA46B14FFC0FFC0FED959D4FF1F9FF9" collectingDate="2017-02-08" collectionCode="HNHM, JG" collectorName="J. Grego" country="Laos" elevation="153" latitude="17.5518" location="Mt. Pha Soung" longLatPrecision="1" longitude="104.87168" pageId="13" pageNumber="363" specimenCode="HNHM 104434, JG4" specimenCount="1" stateProvince="Khammouan">
|
||
1 shell (
|
||
<specimenCode id="4D6AC932FFC0FFC0FE3959D4FD849F79" box="[419,602,1021,1045]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="13" pageNumber="363">HNHM 104434</specimenCode>
|
||
,
|
||
<figureCitation id="85F77DCCFFC0FFC0FDF059D4FD6F9F79" box="[618,689,1021,1045]" captionStart="Fig" captionStartId="12.[113,148,2035,2057]" captionTargetBox="[149,1415,189,2020]" captionTargetId="figure-58@12.[149,1415,189,2020]" captionTargetPageId="12" captionText="Fig. 9. Clostophis multiformis Páll-Gergely & A. Reischütz, new species (HNHM 104434, JG4). A, lateral view; B, ventral view; C, apertural view; D, dorsal view; E, protoconch sculpture; F, aperture dentition; G, sculpture of the ventral side; H, sculpture of body whorl from standard apertural view." figureDoi="http://doi.org/10.5281/zenodo.4576882" httpUri="https://zenodo.org/record/4576882/files/figure.png" pageId="13" pageNumber="363">Fig. 9</figureCitation>
|
||
),
|
||
<collectingCountry id="65DB21D9FFC0FFC0FD5159D7FCDD9F79" box="[715,771,1022,1045]" name="Laos" pageId="13" pageNumber="363">Laos</collectingCountry>
|
||
,
|
||
<collectingRegion id="DF08AFABFFC0FFC0FF125E34FEA09F59" box="[136,382,1053,1077]" country="Laos" name="Khammouan" pageId="13" pageNumber="363">Khammouane Province</collectingRegion>
|
||
, cave at the NE foot of
|
||
<location id="18133792FFC0FFC0FDF25E37FCDC9F59" LSID="urn:lsid:plazi:treatment:9565D05FFFC5FFC0FCDB5D34FBBB9FF9:18133792FFC0FFC0FDF25E37FCDC9F59" box="[616,770,1053,1077]" country="Laos" latitude="17.5518" longLatPrecision="1" longitude="104.87168" name="Mt. Pha Soung" pageId="13" pageNumber="363" stateProvince="Khammouan">Mt. Pha Soung</location>
|
||
, caverns among slope boulders,
|
||
<quantity id="DA34CCACFFC0FFC0FE7D5E14FDF19F39" box="[487,559,1085,1109]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="13" pageNumber="363" unit="m" value="153.0">
|
||
<elevation id="96E1867AFFC0FFC0FE7D5E14FDF19F39" box="[487,559,1085,1109]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="13" pageNumber="363" unit="m" value="153.0">153 m</elevation>
|
||
</quantity>
|
||
a.sl.,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FDEF5E15FCDF9F39" box="[629,769,1084,1109]" degrees="17" direction="north" minutes="33.108" orientation="latitude" pageId="13" pageNumber="363" precision="1" value="17.5518">17°33.108′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FF125E75FEC69F19" box="[136,280,1116,1141]" degrees="104" direction="east" minutes="52.301" orientation="longitude" pageId="13" pageNumber="363" precision="1" value="104.87168">104°52.301′E</geoCoordinate>
|
||
(locality code:
|
||
<specimenCode id="4D6AC932FFC0FFC0FE2C5E74FE3A9F19" box="[438,484,1117,1141]" collectionCode="JG" pageId="13" pageNumber="363">JG4</specimenCode>
|
||
), coll.
|
||
<collectorName id="B039049FFFC0FFC0FDBD5E77FD5E9F19" box="[551,640,1117,1141]" pageId="13" pageNumber="363">J. Grego</collectorName>
|
||
,
|
||
<date id="69724789FFC0FFC0FD135E74FF1F9FF9" pageId="13" pageNumber="363" value="2017-02-08">
|
||
<collectingDate id="7936BE61FFC0FFC0FD135E74FF1F9FF9" pageId="13" pageNumber="363" value="2017-02-08">08 February 2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ADA46B14FFC0FFC0FF545E54FEF59E79" collectingDate="2017-02-09" collectionCode="HNHM, JG" collectorName="Grego, J." country="Laos" elevation="153" latitude="17.5509" location="Pha Soung Cave" longLatPrecision="1" longitude="104.873566" pageId="13" pageNumber="363" specimenCount="1" stateProvince="Khammouan">
|
||
2 shells (coll. JG), same data as preceding; 3 shells (coll. JG),
|
||
<collectingCountry id="65DB21D9FFC0FFC0FF655EB7FEEB9FD9" box="[255,309,1182,1205]" name="Laos" pageId="13" pageNumber="363">Laos</collectingCountry>
|
||
,
|
||
<collectingRegion id="DF08AFABFFC0FFC0FED95EB4FDC09FD9" box="[323,542,1181,1205]" country="Laos" name="Khammouan" pageId="13" pageNumber="363">Khammouane Prov.</collectingRegion>
|
||
,
|
||
<locationDeviation id="0CC1786BFFC0FFC0FDB15EB4FEDC9FB9" location="Ban Na village" pageId="13" pageNumber="363">
|
||
<quantity id="DA34CCACFFC0FFC0FDB15EB4FDBA9FD9" box="[555,612,1181,1205]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="363" unit="km" value="2.0">2 km</quantity>
|
||
WNW of Ban Na village
|
||
</locationDeviation>
|
||
,
|
||
<location id="18133792FFC0FFC0FE8B5E94FE119FB9" LSID="urn:lsid:plazi:treatment:9565D05FFFC5FFC0FCDB5D34FBBB9FF9:18133792FFC0FFC0FE8B5E94FE119FB9" box="[273,463,1213,1237]" country="Laos" latitude="17.5509" longLatPrecision="1" longitude="104.873566" name="Pha Soung Cave" pageId="13" pageNumber="363" stateProvince="Khammouan">Pha Soung Cave</location>
|
||
– Frog lake,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FDE95E95FCDC9FB9" box="[627,770,1212,1237]" degrees="17" direction="north" minutes="33.054" orientation="latitude" pageId="13" pageNumber="363" precision="1" value="17.5509">17°33.054′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78F8078EFFC0FFC0FF135EF5FEFE9F99" box="[137,288,1244,1269]" degrees="104" direction="east" minutes="52.414" orientation="longitude" pageId="13" pageNumber="363" precision="1" value="104.873566">104°52.414′E</geoCoordinate>
|
||
(locality code: JG3A), coll.
|
||
<collectorName id="B039049FFFC0FFC0FDEB5EF4FD059F99" box="[625,731,1245,1269]" pageId="13" pageNumber="363">Grego, J.</collectorName>
|
||
,
|
||
<date id="69724789FFC0FFC0FD765EF4FEF59E79" pageId="13" pageNumber="363" value="2017-02-09">
|
||
<collectingDate id="7936BE61FFC0FFC0FD765EF4FEF59E79" pageId="13" pageNumber="363" value="2017-02-09">09 February 2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55D632C2FFC0FFC0FF125F14FE4F9ED9" pageId="13" pageNumber="363" type="diagnosis">
|
||
<paragraph id="1D736149FFC0FFC0FF125F14FE4F9ED9" blockId="13.[136,777,1341,1461]" pageId="13" pageNumber="363">
|
||
<emphasis id="2FB8BD5BFFC0FFC0FF125F14FF239E39" bold="true" box="[136,253,1341,1365]" pageId="13" pageNumber="363">Diagnosis.</emphasis>
|
||
This variable
|
||
<taxonomicName id="DACC1ACAFFC0FFC0FE0F5F14FDD99E39" authorityName="Benson" authorityYear="1860" box="[405,519,1341,1365]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="363" phylum="Mollusca" rank="genus">
|
||
<emphasis id="2FB8BD5BFFC0FFC0FE0F5F14FDD99E39" box="[405,519,1341,1365]" italics="true" pageId="13" pageNumber="363">Clostophis</emphasis>
|
||
</taxonomicName>
|
||
species is characterised by a conical or ovoid-conical shell, a constricted, teardrop- or kidney-shaped aperture with a weak parietal tooth, and a rather wide umbilicus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55D632C2FFC0FFC0FF125FF4FD719CB9" pageId="13" pageNumber="363" type="description">
|
||
<paragraph id="1D736149FFC0FFC0FF125FF4FD329C19" blockId="13.[135,776,1501,1909]" pageId="13" pageNumber="363">
|
||
<emphasis id="2FB8BD5BFFC0FFC0FF125FF4FECB9E99" bold="true" box="[136,277,1501,1525]" pageId="13" pageNumber="363">Description.</emphasis>
|
||
Shell conical or ovoid, higher than wide to wider than high. Whorls rounded or very slightly shouldered. Tuba slightly (usually) or strongly (rarely) detached from penultimate whorl. Protoconch consisting of 1.25–1.5 whorls, pitted and spirally striated. Entire shell with 4.25–4.5 whorls. Umbilicus wide, occupies ca. one third of the shell width (see also Remarks). Entire teleoconch with strong, elevated, rather equidistant spiral striae (see also Remarks), and some weak, irregular radial growth lines. Aperture teardrop- or kidney-shaped. Sinulus wide. Parietal tooth weak and relatively short, in some shells reaches, in others does not reach peristome. In some shells there is a slight indication of a palatal tooth. Peristome expanded but not reflected.
|
||
</paragraph>
|
||
<paragraph id="1D736149FFC0FFC0FF125DB4FD719CB9" blockId="13.[136,774,1949,2005]" pageId="13" pageNumber="363">
|
||
<emphasis id="2FB8BD5BFFC0FFC0FF125DB4FE429CD9" bold="true" box="[136,412,1949,1973]" pageId="13" pageNumber="363">Measurements (in mm).</emphasis>
|
||
SH = 1.02–1.4, SW = 1.03–1.39 (n = 11, shells belonging to multiple populations).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55D632C2FFC0FFC0FCD95A94FB4A9A59" pageId="13" pageNumber="363" type="diagnosis">
|
||
<paragraph id="1D736149FFC0FFC0FCD95A94FB4A9A59" blockId="13.[835,1474,189,309]" pageId="13" pageNumber="363">
|
||
<emphasis id="2FB8BD5BFFC0FFC0FCD95A94FB9C9BB9" bold="true" box="[835,1090,189,213]" pageId="13" pageNumber="363">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="DACC1ACAFFC0FFC0FBD65A94FAF99BB9" authority="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen, 2020" authorityName="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen" authorityYear="2020" box="[1100,1319,189,213]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="363" phylum="Mollusca" rank="species" species="infantilis" status="sp. nov.">
|
||
<emphasis id="2FB8BD5BFFC0FFC0FBD65A94FAF99BB9" box="[1100,1319,189,213]" italics="true" pageId="13" pageNumber="363">Clostophis infantilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="348B0020FFC0FFC0FAA25A94FA609BB9" box="[1336,1470,189,213]" pageId="13" pageNumber="363" rank="species">new species</taxonomicNameLabel>
|
||
, has a smaller shell, a more pointed apex, stronger radial sculptural elements, and a narrower umbilicus. See also under
|
||
<taxonomicName id="DACC1ACAFFC0FFC0FC135B34FC259A59" authority="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen, 2020" authorityName="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen" authorityYear="2020" box="[905,1019,285,309]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="363" phylum="Mollusca" rank="species" species="obtusus" status="sp. nov.">
|
||
<emphasis id="2FB8BD5BFFC0FFC0FC135B34FC259A59" box="[905,1019,285,309]" italics="true" pageId="13" pageNumber="363">C. obtusus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="348B0020FFC0FFC0FB915B34FB4E9A59" box="[1035,1168,285,309]" pageId="13" pageNumber="363" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55D632C2FFC0FFC0FCD95B74FB0B9AF9" pageId="13" pageNumber="363" type="etymology">
|
||
<paragraph id="1D736149FFC0FFC0FCD95B74FB0B9AF9" blockId="13.[835,1474,349,405]" pageId="13" pageNumber="363">
|
||
<emphasis id="2FB8BD5BFFC0FFC0FCD95B74FC1B9A19" bold="true" box="[835,965,349,373]" pageId="13" pageNumber="363">Etymology.</emphasis>
|
||
This new species is named after the variability of its shell shape across populations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55D632C2FFC0FFC0FCD95B94FBBB9FF9" pageId="13" pageNumber="363" type="discussion">
|
||
<paragraph id="1D736149FFC0FFC0FCD95B94FBBB9FF9" blockId="13.[834,1476,445,1173]" pageId="13" pageNumber="363">
|
||
<emphasis id="2FB8BD5BFFC0FFC0FCD95B94FC6C9AB9" bold="true" box="[835,946,445,469]" pageId="13" pageNumber="363">Remarks.</emphasis>
|
||
This is a variable species in terms of shape and size, and the density of spiral striae. Future investigations including more detailed sampling and larger sample sizes might reveal that the populations listed here belong to multiple species. The populations east of Thakhek (localities: 3L07, 4L07, La.4, La.8, SA1) are slightly smaller (SH:
|
||
<quantity id="DA34CCACFFC0FFC0FAC35874FCAE99F8" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.21" metricValueMin="1.09" pageId="13" pageNumber="363" unit="mm" value="1.15" valueMax="1.21" valueMin="1.09">1.09–1.21 mm</quantity>
|
||
, SW:
|
||
<quantity id="DA34CCACFFC0FFC0FC2E5854FB8E99F8" box="[948,1104,637,661]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.12" metricValueMin="1.03" pageId="13" pageNumber="363" unit="mm" value="1.075" valueMax="1.12" valueMin="1.03">1.03–1.12 mm</quantity>
|
||
vs. SH:
|
||
<quantity id="DA34CCACFFC0FFC0FB2B5854FA9D99F8" box="[1201,1347,637,661]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" metricValueMax="1.4" metricValueMin="1.14" pageId="13" pageNumber="363" unit="mm" value="1.27" valueMax="1.4" valueMin="1.14">1.14–1.4 mm</quantity>
|
||
, SW:
|
||
<quantity id="DA34CCACFFC0FFC0FA1D5854FC7A99D9" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.39" metricValueMin="1.21" pageId="13" pageNumber="363" unit="mm" value="1.2999999999999998" valueMax="1.39" valueMin="1.21">1.21– 1.39 mm</quantity>
|
||
) and have a higher spire and a narrower umbilicus than the populations northeast of Thakhek (localities: JG2B, JG3A, JG4). The sculpture and the aperture dentition of the two groups are similar. We have no samples from the area between the two known areas, therefore, at the moment, we see no necessity of splitting this species into subspecies. The tube (terminal part of the body whorl) is slightly detached from the penultimate whorl, but in one population (locality: La.8) it was strongly detached. Nearly all examined shells have dense spiral striation (26–30 striae counted in the middle line of the body whorl from standard apertural view), whereas the single shell from population JG4 has only 20 striae. The shell and aperture shape are similar to the neighbouring population (locality: JG3A) and therefore, we consider it conspecific with the neighbouring population (locality: JG4). See
|
||
<figureCitation id="85F77DCCFFC0FFC0FB855E54FBBF9FF9" box="[1055,1121,1149,1173]" captionStart="Fig" captionStartId="4.[113,149,1778,1800]" captionTargetBox="[113,1451,387,1763]" captionTargetId="figure-11@4.[113,1451,387,1763]" captionTargetPageId="4" captionText="Fig. 2. Known distribution of Clostophis species. Empty triangle: Clostophis stochi; inverted empty triangle: C. proboscideus; filled triangle: Clostophis sankeyi; inverted filled triangle: Clostophis koilobasis, new species, and Clostophis incurvus, new species; filled square: Clostophis neglectus; filled diamond: Clostophis platytrochus, new species. Inset map shows the distribution of Clostophis multiformis, new species (circles with labels, see explanation in the text), Clostophis obtusus, new species (empty diamond), and Clostophis infantilis, new species (empty square), east of Thakhek." figureDoi="http://doi.org/10.5281/zenodo.4576865" httpUri="https://zenodo.org/record/4576865/files/figure.png" pageId="13" pageNumber="363">Fig. 2</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |