145 lines
21 KiB
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145 lines
21 KiB
XML
<document id="EBA066906D6FEB13F6914F341D1C6520" ID-CLB-Dataset="300857" ID-DOI="10.11646/zootaxa.5482.1.1" ID-GBIF-Dataset="ac67f31a-36b5-4570-8356-c43cd0614252" ID-ISSN="1175-5334" ID-Zenodo-Dep="13209117" ID-ZooBank="AE4D9DE4-95E1-49CC-8104-1E7242523986" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1722785713910" checkinUser="plazi" docAuthor="Bohn, Horst" docDate="2024" docId="9538E406FFBEFF822AC3FF3AFC56CB58" docLanguage="en" docName="zootaxa.5482.1.1.pdf" docOrigin="Zootaxa 5482 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5482.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Heterogamodes (Heterogamisca) arenarum Bohn 2024, comb. nov." docType="treatment" docVersion="4" lastPageNumber="25" masterDocId="69019C7EFFA6FF9A2A54FFE5FFF5C909" masterDocTitle="The spine armament of the legs as an important means for the characterisation of the genera of Corydiinae and their relationships (Blattodea, Corydiidae)" masterLastPageNumber="79" masterPageNumber="1" pageNumber="25" updateTime="1722945181962" updateUser="ExternalLinkService">
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<mods:title id="5FF7E71631FB8F2D9837C6693A696EB5">The spine armament of the legs as an important means for the characterisation of the genera of Corydiinae and their relationships (Blattodea, Corydiidae)</mods:title>
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<mods:title id="59C15DA9ABE47CB18AEFF085A23B0764">Zootaxa</mods:title>
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<mods:part id="6EA251AB672BA0CB17F5FBB7B7FFCBAD">
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<mods:date id="C3DE3A604D71FB38B7FF43A6F637DA95">2024</mods:date>
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<mods:number id="35C4FA285D1363EBC327F0D24FB93AC4">2024-07-22</mods:number>
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<mods:number id="38ADD74EA4653CC34965096D65D2AFA5">5482</mods:number>
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<treatment id="9538E406FFBEFF822AC3FF3AFC56CB58" ID-DOI="http://doi.org/10.5281/zenodo.13209101" ID-Zenodo-Dep="13209101" LSID="urn:lsid:plazi:treatment:9538E406FFBEFF822AC3FF3AFC56CB58" httpUri="http://treatment.plazi.org/id/9538E406FFBEFF822AC3FF3AFC56CB58" lastPageNumber="25" pageId="24" pageNumber="25">
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<subSubSection id="558B069BFFBEFF822AC3FF3AFBAAC9F0" box="[151,1119,223,250]" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="1D2E5510FFBEFF822AC3FF3AFBAAC9F0" blockId="24.[151,1119,223,286]" box="[151,1119,223,250]" pageId="24" pageNumber="25">
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<emphasis id="2FE58902FFBEFF822AC3FF3AFBAAC9F0" bold="true" box="[151,1119,223,250]" pageId="24" pageNumber="25">
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No. 8.
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<taxonomicName id="DA912E93FFBEFF822AB3FF3AFC3EC9F0" ID-CoL="6M52V" authority="(Chopard, 1929)" authorityName="Bohn" authorityYear="2024" baseAuthorityName="Chopard" baseAuthorityYear="1929" box="[231,971,223,250]" class="Insecta" family="Corydiidae" genus="Heterogamodes" kingdom="Animalia" order="Blattodea" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="arenarum" status="comb. nov." subGenus="Heterogamisca">
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<emphasis id="2FE58902FFBEFF822AB3FF3AFD03C9F0" bold="true" box="[231,758,223,250]" italics="true" pageId="24" pageNumber="25">Heterogamodes (Heterogamisca) arenarum</emphasis>
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(
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<bibRefCitation id="790028E1FFBEFF822951FF3AFC36C9F0" author="Chopard, L." box="[773,963,223,249]" pageId="24" pageNumber="25" pagination="215 - 358" refId="ref52228" refString="Chopard, L. (1929) Orthoptera palaearctica critica. VII. Les Polyphagiens de la faune palearctique (Orth., Blatt.). Eos, 5, 215 - 358." type="journal article" year="1929">Chopard, 1929</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel id="34D63479FFBEFF82298DFF3AFBAAC9F0" box="[985,1119,223,249]" pageId="24" pageNumber="25" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="558B069BFFBEFF822AC3FEE6FEB6C817" box="[151,323,259,286]" pageId="24" pageNumber="25" type="description">
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<paragraph id="1D2E5510FFBEFF822AC3FEE6FEB6C817" blockId="24.[151,1119,223,286]" box="[151,323,259,286]" pageId="24" pageNumber="25">
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<figureCitation id="85AA4995FFBEFF822AC3FEE6FF0FC814" box="[151,250,259,285]" captionStart="FIGURE 3" captionStartId="53.[151,250,186,210]" captionTargetId="figure-33@52.[220,1367,181,1919]" captionTargetPageId="52" captionText="FIGURE 3. The right phallomere and its parts. A: Complete right phallomere of Type 1 in standard view, seen from the left (inner face) towards the right (outer face); the position of the other two axes of the body are indicated by the labels ant(erior)— post(erior) and dors(al)—ventr(al). The three sclerites of the phallomere are R1M, R2, and R3. The largest dent of R2 is called main dent (MD), in this case stepped in two etages, upper etage (ue). R1M with a tubercular ridge (tr), the two arrows point to an edge where R1M and R2 articulate. B: Most posterior part of Type 6 phallomere in ventral view, showing the frontside of R1M and the backside of the main dent, with its vertical arm (va) in focus, the horizontal arm (ha) out of focus. C,D: Isolated R1M sclerite, with the sclerotised part (sc) at the left, in D (Type 1) with a tubercular ridge (tr), the membraneous part at the right, in C (Type 2) with a gutterlike deepening (g), arrows indicate where R1M articulates with R2. E–G. Phallomeres in ventral view showing the backside of the main dent, R1M with a ridge (r). E: Phallomere Type 1/2, r of R1M as in Type 2 without tubercular elevations, ue in outline rounded as in Type 1, but with a small lobelike projection (l) recalling the pointed tip of Type 2. F,G: Type 3 phallomeres, main dent with a deep incision (arrow).—Species: A: Heterogamisca arenarum, Type; B: Psammoblatta minuta, Type; C: Mollidentoblatta persica, Holotype; D: Heterogamisca longipilosa, Holotype; E: Atuberculoblatta intermedia, B 1416; F: Fissidentoblatta afghana, Paratype; G: F. polyphagella, Holotype.—Enlargements: Scale in B 0.1 mm, in all others 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13235904" httpUri="https://zenodo.org/record/13235904/files/figure.png" pageId="24" pageNumber="25">Figs. 3A</figureCitation>
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,
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<figureCitation id="85AA4995FFBEFF822B52FEE6FEB6C817" box="[262,323,259,286]" captionStart="FIGURE 5" captionStartId="55.[151,250,1873,1897]" captionTargetBox="[220,1368,181,1849]" captionTargetId="figure-16@55.[220,1368,181,1849]" captionTargetPageId="55" captionText="FIGURE 5. Subgenital plate (at the left, ventral view) and posterior part of the right phallomere (at the right, standard view) of male.—Species: A,B: Heterogamisca longipilosa. (A) Holotype, Bo 1329; (B) Paratype, Bo 1358, dents of R2 sclerite isolated; C,D: H. jeffreyana, Holotype, Bo 1328; E,F: H. sulcata, Holotype, Bo 1417; G,H: Heterogamisca arenarum, Type, Bo 1430.— Phallomere type: Type 1 in all corresponding figures.—Explanations: R1M, R2, R3 sclerites of the right phallomere, MD main dent of R2, ue its upper etage; tr tubercular ridge.—Enlargements: Scale 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13209133" httpUri="https://zenodo.org/record/13209133/files/figure.png" pageId="24" pageNumber="25">5G,H</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="558B069BFFBEFF822AC3FEADFD50C894" pageId="24" pageNumber="25" type="reference_group">
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||
<paragraph id="1D2E5510FFBEFF822AC3FEADFCA3C856" blockId="24.[151,961,327,413]" box="[151,854,327,352]" pageId="24" pageNumber="25">
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<treatmentCitationGroup id="3D81723EFFBEFF822AC3FEADFCA3C856" box="[151,854,327,352]" pageId="24" pageNumber="25">
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<treatmentCitation id="9C307301FFBEFF822AC3FEADFD87C856" author="Chopard, L." box="[151,626,327,352]" page="335" pageId="24" pageNumber="25" year="1929">
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<taxonomicName id="DA912E93FFBEFF822AC3FEADFD87C856" ID-CoL="6M52V" authority="Chopard, 1929: 335" authorityName="Chopard" authorityPageNumber="335" authorityYear="1929" box="[151,626,327,352]" class="Insecta" family="Corydiidae" genus="Heterogamodes" kingdom="Animalia" order="Blattodea" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="arenarum">
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<emphasis id="2FE58902FFBEFF822AC3FEADFE55C856" box="[151,416,328,352]" italics="true" pageId="24" pageNumber="25">Heterogamodes arenarum</emphasis>
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<bibRefCitation id="790028E1FFBEFF822BF2FEADFD87C856" author="Chopard, L." box="[422,626,327,352]" pageId="24" pageNumber="25" pagination="215 - 358" refId="ref52228" refString="Chopard, L. (1929) Orthoptera palaearctica critica. VII. Les Polyphagiens de la faune palearctique (Orth., Blatt.). Eos, 5, 215 - 358." type="journal article" year="1929">Chopard, 1929: 335</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, figs. 81,89; 1943, 50.
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="1D2E5510FFBEFF822AC3FE82FC34C877" blockId="24.[151,961,327,413]" box="[151,961,358,382]" pageId="24" pageNumber="25">
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<treatmentCitationGroup id="3D81723EFFBEFF822AC3FE82FC34C877" box="[151,961,358,382]" pageId="24" pageNumber="25">
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||
<taxonomicName id="DA912E93FFBEFF822AC3FE82FDE6C877" ID-CoL="GFKM" baseAuthorityName="Chopard" baseAuthorityYear="1929" box="[151,531,358,382]" class="Insecta" family="Corydiidae" genus="Arenivaga" kingdom="Animalia" order="Blattodea" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="arenarum" subGenus="Psammoblatta">
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<emphasis id="2FE58902FFBEFF822AC3FE82FDE6C877" box="[151,531,358,382]" italics="true" pageId="24" pageNumber="25">Arenivaga (Psammoblatta) arenarum</emphasis>
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||
</taxonomicName>
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||
:
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||
<treatmentCitation id="9C307301FFBEFF822874FE83FCFFC877" author="Bey-Bienko, G. Y." box="[544,778,358,382]" page="307" pageId="24" pageNumber="25" year="1950">
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||
<bibRefCitation id="790028E1FFBEFF822874FE83FCFFC877" author="Bey-Bienko, G. Y." box="[544,778,358,382]" pageId="24" pageNumber="25" pagination="1 - 342" refId="ref51717" refString="Bey-Bienko, G. Y. (1950) Fauna of the U. S. S. R. Insects. Blattodea. Zoologicheskogo Instituta Akademija Nauk. S. S. S. R., Moskva, New Series, 40, 1 - 342. [in Russian]" type="journal article" year="1950">Bey-Bienko 1950: 307</bibRefCitation>
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="9C307301FFBEFF822940FE83FC48C877" author="Princis, K." box="[788,957,358,382]" page="64" pageId="24" pageNumber="25" year="1962">
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||
<bibRefCitation id="790028E1FFBEFF822940FE83FC48C877" author="Princis, K." box="[788,957,358,382]" pageId="24" pageNumber="25" pagination="1 - 74" refId="ref53728" refString="Princis, K. (1962) Blattariae: Subordo Polyphagoidea, Fam. Polyphagidae. In: Beier, M. (Ed.), Orthopterorum Catalogus. Pars 3. Junk, s'Gravenhage, pp. 1 - 74." type="book chapter" year="1962">Princis 1962: 64</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="1D2E5510FFBEFF822AC3FE60FD50C894" blockId="24.[151,961,327,413]" box="[151,677,389,413]" pageId="24" pageNumber="25">
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<treatmentCitationGroup id="3D81723EFFBEFF822AC3FE60FD50C894" box="[151,677,389,413]" pageId="24" pageNumber="25">
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<taxonomicName id="DA912E93FFBEFF822AC3FE60FE53C894" ID-CoL="6LMTV" baseAuthorityName="Chopard" baseAuthorityYear="1929" box="[151,422,389,413]" class="Insecta" family="Corydiidae" genus="Hemelytroblatta" kingdom="Animalia" order="Blattodea" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="arenarum">
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<emphasis id="2FE58902FFBEFF822AC3FE60FE53C894" box="[151,422,389,413]" italics="true" pageId="24" pageNumber="25">Hemelytroblatta arenarum</emphasis>
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||
</taxonomicName>
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||
:
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||
<treatmentCitation id="9C307301FFBEFF822BE7FE60FD54C894" author="Grandcolas, P." box="[435,673,389,413]" page="157" pageId="24" pageNumber="25" year="1994">
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<bibRefCitation id="790028E1FFBEFF822BE7FE60FD54C894" author="Grandcolas, P." box="[435,673,389,413]" pageId="24" pageNumber="25" pagination="145 - 158" refId="ref52889" refString="Grandcolas, P. (1994 a) Phylogenetic systematics of the subfamily Polyphaginae, with the assignment of Cryptocercus Scudder, 1862 to this taxon (Blattaria, Blaberoidea, Polyphagidae). Systematic Entomology, 19, 145 - 158. https: // doi. org / 10.1111 / j. 1365 - 3113.1994. tb 00584. x" type="journal article" year="1994">Grandcolas 1994a: 157</bibRefCitation>
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</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="558B069BFFBEFF822AC3FE2EFDF8CB24" pageId="24" pageNumber="25" type="description">
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<paragraph id="1D2E5510FFBEFF822AC3FE2EFC61CB03" blockId="24.[151,1436,459,594]" pageId="24" pageNumber="25">
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<emphasis id="2FE58902FFBEFF822AC3FE2EFE91C8EC" bold="true" box="[151,356,459,485]" pageId="24" pageNumber="25">Male characters.</emphasis>
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Tibia armament: [1.8.0][6.8.1][8.7.3–4], dorsal spines of hindtibia distributed in two groups (5+3).—Subgenital plate:
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<figureCitation id="85AA4995FFBEFF822BE9FE0AFDE7CB00" box="[445,530,495,522]" captionStart="FIGURE 5" captionStartId="55.[151,250,1873,1897]" captionTargetBox="[220,1368,181,1849]" captionTargetId="figure-16@55.[220,1368,181,1849]" captionTargetPageId="55" captionText="FIGURE 5. Subgenital plate (at the left, ventral view) and posterior part of the right phallomere (at the right, standard view) of male.—Species: A,B: Heterogamisca longipilosa. (A) Holotype, Bo 1329; (B) Paratype, Bo 1358, dents of R2 sclerite isolated; C,D: H. jeffreyana, Holotype, Bo 1328; E,F: H. sulcata, Holotype, Bo 1417; G,H: Heterogamisca arenarum, Type, Bo 1430.— Phallomere type: Type 1 in all corresponding figures.—Explanations: R1M, R2, R3 sclerites of the right phallomere, MD main dent of R2, ue its upper etage; tr tubercular ridge.—Enlargements: Scale 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13209133" httpUri="https://zenodo.org/record/13209133/files/figure.png" pageId="24" pageNumber="25">Fig. 5G</figureCitation>
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.—Right phallomere:
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<figureCitation id="85AA4995FFBEFF822953FE0AFCAACB00" box="[775,863,495,521]" captionStart="FIGURE 3" captionStartId="53.[151,250,186,210]" captionTargetId="figure-33@52.[220,1367,181,1919]" captionTargetPageId="52" captionText="FIGURE 3. The right phallomere and its parts. A: Complete right phallomere of Type 1 in standard view, seen from the left (inner face) towards the right (outer face); the position of the other two axes of the body are indicated by the labels ant(erior)— post(erior) and dors(al)—ventr(al). The three sclerites of the phallomere are R1M, R2, and R3. The largest dent of R2 is called main dent (MD), in this case stepped in two etages, upper etage (ue). R1M with a tubercular ridge (tr), the two arrows point to an edge where R1M and R2 articulate. B: Most posterior part of Type 6 phallomere in ventral view, showing the frontside of R1M and the backside of the main dent, with its vertical arm (va) in focus, the horizontal arm (ha) out of focus. C,D: Isolated R1M sclerite, with the sclerotised part (sc) at the left, in D (Type 1) with a tubercular ridge (tr), the membraneous part at the right, in C (Type 2) with a gutterlike deepening (g), arrows indicate where R1M articulates with R2. E–G. Phallomeres in ventral view showing the backside of the main dent, R1M with a ridge (r). E: Phallomere Type 1/2, r of R1M as in Type 2 without tubercular elevations, ue in outline rounded as in Type 1, but with a small lobelike projection (l) recalling the pointed tip of Type 2. F,G: Type 3 phallomeres, main dent with a deep incision (arrow).—Species: A: Heterogamisca arenarum, Type; B: Psammoblatta minuta, Type; C: Mollidentoblatta persica, Holotype; D: Heterogamisca longipilosa, Holotype; E: Atuberculoblatta intermedia, B 1416; F: Fissidentoblatta afghana, Paratype; G: F. polyphagella, Holotype.—Enlargements: Scale in B 0.1 mm, in all others 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13235904" httpUri="https://zenodo.org/record/13235904/files/figure.png" pageId="24" pageNumber="25">Fig. 3A</figureCitation>
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||
,
|
||
<figureCitation id="85AA4995FFBEFF82293FFE0AFC65CB03" box="[875,912,495,522]" captionStart="FIGURE 5" captionStartId="55.[151,250,1873,1897]" captionTargetBox="[220,1368,181,1849]" captionTargetId="figure-16@55.[220,1368,181,1849]" captionTargetPageId="55" captionText="FIGURE 5. Subgenital plate (at the left, ventral view) and posterior part of the right phallomere (at the right, standard view) of male.—Species: A,B: Heterogamisca longipilosa. (A) Holotype, Bo 1329; (B) Paratype, Bo 1358, dents of R2 sclerite isolated; C,D: H. jeffreyana, Holotype, Bo 1328; E,F: H. sulcata, Holotype, Bo 1417; G,H: Heterogamisca arenarum, Type, Bo 1430.— Phallomere type: Type 1 in all corresponding figures.—Explanations: R1M, R2, R3 sclerites of the right phallomere, MD main dent of R2, ue its upper etage; tr tubercular ridge.—Enlargements: Scale 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13209133" httpUri="https://zenodo.org/record/13209133/files/figure.png" pageId="24" pageNumber="25">5H</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1D2E5510FFBEFF822A93FDF6FDF8CB24" blockId="24.[151,1436,459,594]" box="[199,525,531,557]" pageId="24" pageNumber="25">
|
||
<emphasis id="2FE58902FFBEFF822A93FDF6FEDDCB24" bold="true" box="[199,296,531,557]" pageId="24" pageNumber="25">Female:</emphasis>
|
||
<bibRefCitation id="790028E1FFBEFF822B7BFDF6FE24CB24" author="Chopard, L." box="[303,465,531,557]" pageId="24" pageNumber="25" pagination="215 - 358" refId="ref52228" refString="Chopard, L. (1929) Orthoptera palaearctica critica. VII. Les Polyphagiens de la faune palearctique (Orth., Blatt.). Eos, 5, 215 - 358." type="journal article" year="1929">Chopard 1929</bibRefCitation>
|
||
, 337.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="558B069BFFBEFF822A93FDD2FC56CB58" box="[199,931,567,594]" pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph id="1D2E5510FFBEFF822A93FDD2FC56CB58" blockId="24.[151,1436,459,594]" box="[199,931,567,594]" pageId="24" pageNumber="25">
|
||
<emphasis id="2FE58902FFBEFF822A93FDD2FE60CB58" bold="true" box="[199,405,567,593]" pageId="24" pageNumber="25">Material studied.</emphasis>
|
||
<materialsCitation id="ADF95F4DFFBEFF822BCFFDDDFC6ACB58" box="[411,927,567,594]" collectorName="Bo & M. Paris" location="Biskra" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Biskra" typeStatus="holotype">
|
||
<typeStatus id="C22AEBB2FFBEFF822BCFFDDDFE20CB5B" box="[411,469,568,594]" pageId="24" pageNumber="25">Type</typeStatus>
|
||
,
|
||
<specimenCount id="0B979E99FFBEFF822BB5FDD2FDF0CB58" box="[481,517,567,593]" count="1" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
|
||
,
|
||
<collectingRegion id="DF559BF2FFBEFF822846FDD2FDAECB58" box="[530,603,567,593]" country="Algeria" name="Biskra" pageId="24" pageNumber="25">Biskra</collectingRegion>
|
||
(prep.:
|
||
<collectorName id="B06430C6FFBEFF8228E7FDDDFD21CB5B" box="[691,724,568,594]" pageId="24" pageNumber="25">Bo</collectorName>
|
||
1430). (
|
||
<collectorName id="B06430C6FFBEFF822967FDD2FC62CB58" box="[819,919,567,593]" pageId="24" pageNumber="25">M. Paris</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |