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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="226E6D43C4A525B4EB9F1119B449C9CF" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus kerga Marseul 1870" docType="treatment" docVersion="3" lastPageNumber="51" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="48" updateTime="1668155382655" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>271</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>401</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152040995" LSID="urn:lsid:plazi:treatment:226E6D43C4A525B4EB9F1119B449C9CF" httpUri="http://treatment.plazi.org/id/226E6D43C4A525B4EB9F1119B449C9CF" lastPageId="50" lastPageNumber="51" pageId="47" pageNumber="48">
<subSubSection pageId="47" pageNumber="48" type="nomenclature">
<paragraph pageId="47" pageNumber="48">
<taxonomicName LSID="http://species-id.net/wiki/Operclipygus_kerga" authority="Marseul, 1870" authorityName="Marseul" authorityYear="1870" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga (Marseul, 1870)</taxonomicName>
Figs 11
<normalizedToken originalValue="A–B12A–C">A-B12A-C</normalizedToken>
, FMap 3
</paragraph>
</subSubSection>
<subSubSection pageId="47" pageNumber="48" type="reference_group">
<paragraph pageId="47" pageNumber="48">
<taxonomicName class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister kerga" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kerga">Phelister kerga</taxonomicName>
Marseul, 1870: 77;
<taxonomicName class="Insecta" family="Histeridae" genus="Pseudister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudister kerga" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kerga">Pseudister kerga</taxonomicName>
:
<bibRefCitation pageId="47" pageNumber="48">Bickhardt 1917</bibRefCitation>
: 165;
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
:
<bibRefCitation author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel 1976</bibRefCitation>
: 259.
</paragraph>
</subSubSection>
<subSubSection pageId="47" pageNumber="48" type="type locality">
<paragraph pageId="47" pageNumber="48">Type locality.</paragraph>
<paragraph pageId="47" pageNumber="48">Unspecified, somewhere along the Amazon in Brazil.</paragraph>
</subSubSection>
<subSubSection lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="type material">
<paragraph pageId="47" pageNumber="48">Type material.</paragraph>
<paragraph lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
Lectotype, here designated, of undetermined sex: &quot;
<taxonomicName class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister kerga" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kerga">Phelister kerga</taxonomicName>
, Amazon&quot; / &quot;LECTOTYPE
<taxonomicName class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister kerga" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kerga">Phelister kerga</taxonomicName>
<bibRefCitation author="Marseul, SA" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="377" pageNumber="378" pagination="55 - 138" title="Description d'especes nouvelles d'Histerides." volume="13" year="1870">Marseul 1870</bibRefCitation>
, M.S.Caterino &amp; A.K.Tishechkin des. 2010&quot; (MNHN). This species was described from an unspecified
<pageBreakToken pageId="48" pageNumber="49" start="start">number</pageBreakToken>
of specimens, and the lectotype designation fixes primary type status on the only known original specimen.
</paragraph>
</subSubSection>
<subSubSection pageId="48" pageNumber="49" type="other material">
<paragraph pageId="48" pageNumber="49">Other material.</paragraph>
<paragraph pageId="48" pageNumber="49">
BOLIVIA: Santa Cruz: 6: 4-5km SSE Buena Vista, Hotel Flora y Fauna,
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-17.498333">17°29.9'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-63.651665">63°39.1'W</geoCoordinate>
, 15-24.xii.2003, FIT, S. &amp; J. Peck (AKTC); 2: [country record only] (ZMHB). BRAZIL:
<normalizedToken originalValue="Pará">Para</normalizedToken>
: 1: Monte Alegre,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.15">3°9'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-52.05">52°3'W</geoCoordinate>
, 17.vi.1992, 3.vii.1992, FIT (CHND); 1: 10-27.vi.1992, FIT (CHND). 5:
<normalizedToken originalValue="Tucuruí">Tucurui</normalizedToken>
,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.75">3°45'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-49.666668">49°40'W</geoCoordinate>
, 10-29.vii.1985, FIT (CHND); 1: 19.vi.1986, 7.vii.1986, FIT (CHND). FRENCH GUIANA: 1:
<normalizedToken originalValue="Belvèdére">Belvedere</normalizedToken>
de
<normalizedToken originalValue="Saül">Sauel</normalizedToken>
, point de vue,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="3.0227778">3°1'22&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-53.209446">53°12'34&quot;W</geoCoordinate>
, 30.xi.2010, FIT, SEAG (CHND). PERU: Cusco: 1: Cock of the Rock Lodge, NE Paucartambo,
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-13.055">13°3.3'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-71.545">71°32.7'W</geoCoordinate>
, 1120m, 4.xi.2007, 9.xi.2007, FIT, D. Brzoska (SEMC). 1: Manu rd. km 165, Consuelo, 10.x.1982, beating dead branches, L. Watrous &amp; G. Mazurek (FMNH); 1: 12.x.1982, rotten palm, L. Watrous &amp; G. Mazurek (FMNH).
<normalizedToken originalValue="Junín">Junin</normalizedToken>
: 4: Pampa Hermosa Lodge, 22km N San Ramon,
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-10.988334">10°59.3'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-75.425">75°25.5'W</geoCoordinate>
, 1220m, 24-27.xi.2007, FIT, D. Brzoska (SEMC). 1: 11km NE Puerto Ocopa, Los Olivos,
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-11.116667">11°7.00'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-74.25867">74°15.52'W</geoCoordinate>
, 1200m, 26-28.iii.2009, Window trap, A.V. Petrov (AKTC); 1: 24-26.iii.2009, Window trap, A.V. Petrov (MUSM). 1: 7.5km NE Puerto Ocopa, Cananeden,
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-11.081667">11°4.9'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-74.26833">74°16.1'W</geoCoordinate>
, 1180m, 11.i.2007, A.V. Petrov (AKTC); 1: 15km NW Satipo, Rio Venado,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-11.19515">11°11.709'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-74.76865">74°46.119'W</geoCoordinate>
, 1150m, 17-24.v.2012, A.V. Petrov, DNA Extract MSC-2322 (AKTC). Loreto: 2: km 63, rd. Iquitos - Nauta, Rio Itaya,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-4.2534165">4°15.205'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-73.43457">73°26.074'W</geoCoordinate>
, 140m, 10-14.ii.2010, A.V. Petrov (AKTC). 1: 9-13.i.2011, A.V. Petrov (MUSM). Madre de Dios: 1: Los Amigos Field Sta., Luisa Bamboo forest,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-12.5422">12.5422°S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-74.117">74.1170°W</geoCoordinate>
, 274m, 2-11.i.2007, pitfall, bamboo forest, J. Jacobs (CASC). 1: Manu National Park, Zona res., Rio Manu, Cocha Juarez, trail nr. Manu Loge, 18-24.ix.1991, FIT, A. Hartman (FMNH). 16: Manu National Park, Pantiacolla Lodge, Alto Madre de Dios River,
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-12.655">12°39.3'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-71.23167">71°13.9'W</geoCoordinate>
, 420m, 14-19.xi.2007, FIT, D. Brzoska (SEMC). 1: Manu National Park, Pantiacolla Lodge, 5.5km NW El Mirador Trail, Alto Madre de Dios River,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.652778">12°39'10&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-71.257774">71°15'28&quot;W</geoCoordinate>
, 500m, 23-26.x.2000, FIT, R. Brooks (SEMC). 6: Manu National Park, Cocha Cashu Bio. Sta.,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-11.895833">11°53'45&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-71.40667">71°24'24&quot;W</geoCoordinate>
, 350m, 17-19.x.2000, FIT, R. Brooks (SEMC). 2: Manu National Park, Cocha Salvador,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.003333">12°0'12&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-71.52667">71°31'36&quot;W</geoCoordinate>
, 310m, 20-21.x.2000, FIT, R. Brooks (SEMC). 2: Tambopata, Reserva Cuzco Amazonico, 15km NE Puerto Maldonado, Quebrada Mariposa,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
, 200m, 13.vi.1989, FIT, R. A. Leschen (SEMC). VENEZUELA: Monagas: 11: Caripe Cueva
<normalizedToken originalValue="Guácharo">Guacharo</normalizedToken>
, 700m, 20-30.vii.1987, malaise trap, forest over coffee, S. &amp; J. Peck (FMNH).
</paragraph>
</subSubSection>
<subSubSection lastPageId="50" lastPageNumber="51" pageId="48" pageNumber="49" type="description">
<paragraph pageId="48" pageNumber="49">Diagnostic description.</paragraph>
<paragraph lastPageId="50" lastPageNumber="51" pageId="48" pageNumber="49">
Length: 2.40-2.87 mm, width: 2.18-2.46 mm; body rounded at sides, moderately convex dorsally; frons slightly convex; frontal stria interrupted on each side; supraorbital stria complete across vertex, barely separated from portion of frontal stria in front of eyes; pronotum with lateral submarginal stria complete, free anteriorly; anterior submarginal pronotal stria present; median pronotal gland openings slightly behind anterior margin, near apices of recurved submarginal stria; pronotal disk with few or no coarse lateral punctures; antescutellar fovea present, small, ovoid; elytra with outer subhumeral, inner subhumeral, and striae 1-3 complete, striae 4 and 5 usually subequal, present in apical third to one-half, sutural stria present in apical two-thirds; prosternal keel striae convergent and connected anteriorly, keel project
<pageBreakToken pageId="49" pageNumber="50" start="start">ing</pageBreakToken>
posteriorly; marginal mesoventral stria complete; mesometaventral stria outwardly arcuate, nearly in contact with marginal mesoventral stria at middle; central portion of metaventral disk impunctate, small fragment of secondary lateral metaventral stria gen
<pageBreakToken pageId="50" pageNumber="51" start="start">erally</pageBreakToken>
present; propygidium uniformly covered with round punctures separated by one-fourth their diameters; pygidium with dense, fine ground punctation, with small but coarser punctures sparsely intermingled, apical marginal stria complete, finely crenulate. Male genitalia (Figs 12
<normalizedToken originalValue="A–C">A-C</normalizedToken>
, F): accessory sclerites present; T8 short, basal emargination broad, apical emargination narrow, basal membrane attachment line separated from basal emargination by one-third its depth, ventrolateral apodemes not meeting at midline, most strongly developed basally, narrowed toward apex; S8 with narrow apical guides gradually widened to apex, halves approximate in basal half, weakly diverging to apex; T9 with apices bluntly rounded, ventrolateral tooth well developed, just beyond midpoint; T10 halves separate; S9 narrowest near apex, expanded toward base, parallel sided in basal fourth, base truncate to weakly emarginate, lacking apical emargination, apical flanges separate; tegmen with sides evenly rounded to base and apex, medioventral process fine, narrowly rounded, weakly projecting beneath just basad midpoint; basal piece just less than one-third tegmen length; median lobe about one-half tegmen length.
</paragraph>
</subSubSection>
<subSubSection pageId="50" pageNumber="51" type="remarks">
<paragraph pageId="50" pageNumber="51">Remarks.</paragraph>
<paragraph pageId="50" pageNumber="51">
Of two specimens in ZMHB, one bears a label 'Phelister
<normalizedToken originalValue="bimarginatus’">bimarginatus'</normalizedToken>
, apparently in Bickhardt's handwriting. However, this is a nomen nudum. Presumably, Bickhardt intended to describe this species before deciding it was
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
.
</paragraph>
<paragraph pageId="50" pageNumber="51">The type locality is merely the Amazon region. Beyond that it is widespread in much of the Amazonian basin, except where it is replaced by its close relatives, described below.</paragraph>
<caption pageId="50" pageNumber="51">
<paragraph pageId="50" pageNumber="51">
Figure 11.
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
group. A Dorsal habitus
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
(lectotype) B Frons
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
C Pygidia of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus punctistrius" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="punctistrius">Operclipygus punctistrius</taxonomicName>
D Pygidia of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus planifrons" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="planifrons">Operclipygus planifrons</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="50" pageNumber="51">
<paragraph pageId="50" pageNumber="51">
Figure 12. Male genitalia of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
group. A T8 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
B S8 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
C S9 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
D S9 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus punctistrius" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="punctistrius">Operclipygus punctistrius</taxonomicName>
E S9 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus planifrons" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="planifrons">Operclipygus planifrons</taxonomicName>
F Aedeagus, dorsal and lateral views, of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
G Aedeagus, dorsal and lateral views, of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus punctistrius" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="punctistrius">Operclipygus punctistrius</taxonomicName>
H Aedeagus, lateral view, of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus planifrons" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="planifrons">Operclipygus planifrons</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="50" pageNumber="51">
<paragraph pageId="50" pageNumber="51">
Map 3. Records of the
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
group.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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