433 lines
46 KiB
XML
433 lines
46 KiB
XML
<document id="DD43CF017D85931E8242D7144604468B" ID-DOI="10.3897/dez.71.118923" ID-ZooBank="73CD46B3-AA8E-42CB-8640-3C04D799D1F6" ID-publisher-id="118923" URI-arpha="D3663AC6-0EB2-5536-963F-32F61189DD89" URI-zoobank="http://zoobank.org/73CD46B3-AA8E-42CB-8640-3C04D799D1F6" article-type="research-article" checkinTime="1722350525570" checkinUser="pensoft" docAuthor="Clavijo-Bustos, Julián, Castro-Vargas, María I. & Neita Moreno, Jhon César" docDate="2024" docId="FE891834304252368DBA79D5AA7866F3" docLanguage="en" docName="Deutsche Entomologische Zeitschrift 71(1): 209-217" docOrigin="Deutsche Entomologische Zeitschrift 71 (1)" docSource="https://dez.pensoft.net/article/118923/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Manodactylus paramicola Clavijo-Bustos, Castro-Vargas & Moreno, 2024, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="217" masterDocId="D3663AC60EB25536963F32F61189DD89" masterDocTitle="A second species of the genus Manodactylus Moser, 1919 (Coleoptera, Scarabaeidae, Melolonthinae, Macrodactylini) from the highlands of Colombia" masterLastPageNumber="217" masterPageNumber="209" pageNumber="209" updateTime="1722351143611" updateUser="ExternalLinkService">
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<mods:title id="8A808B900B33B4357741F96E54676AE5">A second species of the genus Manodactylus Moser, 1919 (Coleoptera, Scarabaeidae, Melolonthinae, Macrodactylini) from the highlands of Colombia</mods:title>
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<mods:name id="69D03AB2522D11BBBCA570BB9506B25B" type="personal">
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<mods:roleTerm id="FE6FF030D22AAD94ECF169056CA8AFAC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7AC1DA18EFC11F0C77C9C5459BE1BFB6">Clavijo-Bustos, Julián</mods:namePart>
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<mods:nameIdentifier id="A5481BBBC515ACCB616A8722B8B82F75" type="ORCID">0000-0002-7453-9792</mods:nameIdentifier>
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<mods:affiliation id="B24A87F06DED8570856206F278965D5E">Sección Entomología, Colecciones Biológicas, Centro Colecciones y Gestión de Especies, Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Villa de Leyva, Boyacá, Colombia</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="A961260293C6D590117AD94570DD8836">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="720AD09BAB2F81D2780ACE9B6D13D715">Castro-Vargas, María I.</mods:namePart>
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<mods:nameIdentifier id="3D7AED59FF6D4B95579C2A12BB63BB99" type="ORCID">0000-0003-0401-6301</mods:nameIdentifier>
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<mods:affiliation id="35AA28331983B58A4C0ECB4F47A18FF8">Maestría en Ciencias Agrarias – Entomología, Facultad de Ciencias Agrarias, Universidad Nacional de Colombia sede Bogotá, Bogotá D. C., Colombia</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="FC1439BB398B884C392A9E20B2F699F2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8D571CA1336A05DA2E66E5B2D3E1E0F9">Neita Moreno, Jhon César</mods:namePart>
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<mods:nameIdentifier id="5466025323AAA8F510292DE02C29D584" type="ORCID">0000-0003-2998-2063</mods:nameIdentifier>
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<mods:affiliation id="F332656ED14925AC465EFB32AD868BD6">Sección Entomología, Colecciones Biológicas, Centro Colecciones y Gestión de Especies, Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Villa de Leyva, Boyacá, Colombia</mods:affiliation>
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<mods:title id="C0642C57C5554AA6FAB5CA6239F1758F">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part id="36F08902173C2EEEB7CE8D6527BED5C3">
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<mods:date id="8CE69B28AF0F430E5DA2FBA0341CE741">2024</mods:date>
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<mods:number id="6A87A3E33A35AE57DA434CA64BC1498F">2024-07-30</mods:number>
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<mods:detail id="AAC878BE40CD869230C45C42105B399B" type="volume">
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<mods:number id="B309EE443018D6782B06BCADDF2CB1CB">71</mods:number>
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<mods:detail id="9546EEA4E5D1BFAADA25BF346E224A1E" type="issue">
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<mods:identifier id="2E9A91AEA5680216CC9ED60903ECA2E6" type="DOI">10.3897/dez.71.118923</mods:identifier>
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<mods:identifier id="43575945B2D564C2FB00368A40BFC0BA" type="ZooBank">73CD46B3-AA8E-42CB-8640-3C04D799D1F6</mods:identifier>
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<treatment id="FE891834304252368DBA79D5AA7866F3" ID-DOI="http://doi.org/10.5281/zenodo.13134905" ID-Zenodo-Dep="13134905" ID-arpha="FE891834-3042-5236-8DBA-79D5AA7866F3" ID-zoobank="https://zoobank.org/A4593819-8F5D-4DD9-A501-BE262E0C1F8A" LSID="urn:lsid:plazi:treatment:FE891834304252368DBA79D5AA7866F3" LSID-ZBK="urn:lsid:zoobank.org:act:A4593819-8F5D-4DD9-A501-BE262E0C1F8A" httpUri="http://treatment.plazi.org/id/FE891834304252368DBA79D5AA7866F3">
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<subSubSection id="CC4246243550A6D7B5D6C746554BD587" type="nomenclature">
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||
<paragraph id="45E4FADCE541682BB8A4FF33BB4F13E4">
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<taxonomicName id="715682ECB8758B5AA15CC6D75C3F8ADF" authorityName="Clavijo-Bustos & Castro-Vargas & Neita Moreno" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="paramicola" status="sp. nov.">
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<emphasis id="CA5263EA06F3A50BC14073843D7DAB47" italics="true">Manodactylus paramicola</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6E7BF16D084025970CDF008841E56389" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="430A2ABF5FB1B5DBE74D3C526657D0E8" type="description">
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<paragraph id="35345DCE0AF502A03FCECB9404170721">
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<figureCitation id="89A6D69C5B9226834A37551E63B5C34F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Manodactylus paramicola sp. nov., male holotype. A. Dorsal view; B. Lateral view; C. Ventral view; D. Aedeagus in lateral view; E. Parameres in dorsal view; F. Parameres in frontal view; G. Spiculum gastrale in dorsal view." figureDoi="10.3897/dez.71.118923.figure2" httpUri="https://binary.pensoft.net/fig/1104275">Figs 2</figureCitation>
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,
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<figureCitation id="1D4292269F8DEEB51F34C1BF113ECADC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Manodactylus paramicola sp. nov., male paratype. A. Hind wing in dorsal view; B. Epipharynx in ventral view; C. Left maxilla in ventral view; D. Left maxilla in dorsal view; E. Labium in ventral view; F. Left mandible in dorsal view; G. Left mandible in lateral internal view; H. Left mandible in ventral view." figureDoi="10.3897/dez.71.118923.figure3" httpUri="https://binary.pensoft.net/fig/1104276">3</figureCitation>
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,
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<figureCitation id="2AB867647F3A99CF063D88EFE39020DE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Manodactylus paramicola sp. nov., male holotype. A. Protibia in dorsal view; B. Detail of the downward projection of the inner angle of protibia; C. Apex of protibia; D. Apex of mesotibia; E, Apex of metatibia. Black arrows point to downward projection of the inner angle of protibia." figureDoi="10.3897/dez.71.118923.figure4" httpUri="https://binary.pensoft.net/fig/1104277">4</figureCitation>
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,
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<figureCitation id="19979F899F4C1166CF4BBC622C50547F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Manodactylus species. A, B. Macrodactylus gaujoni Ohaus, 1909, male neotype = Manodactylus gaujoni Moser, 1919, male lectotype; C, D. Manodactylus paramicola sp. nov., male holotype. A, C. Pronotum in dorsal view; B, D. Head in dorsal view." figureDoi="10.3897/dez.71.118923.figure5" httpUri="https://binary.pensoft.net/fig/1104278">5 C, D</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EHQAE" type="type material">
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||
<paragraph id="714C27D0ABA8479937F9C7051E559B0C">
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||
<heading id="D598630197926CDDA0A786BBF74D40D1" reason="title">Type material.</heading>
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||
</paragraph>
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||
<paragraph id="578B4B3FFF8A9E8ECF9281427F9C7BD2">
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<materialsCitation id="7C4E4522F1142E36D0BF8237E233CD6D" collectorName="B. Yanguatin" country="Colombia" location="Clavijo-Bustos" municipality="Instituto Humboldt" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<emphasis id="04BE5C076577ECEDED0C896D9434591F" bold="true" italics="true">
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<typeStatus id="D0FF25D3DAA74164D1BEB9D09BFD8AD6" type="holotype">Holotype</typeStatus>
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</emphasis>
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male (
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<abbrev id="ABBRID0ESQAE" xlink_title="Sección de Entomología, Colecciones Biológicas del Instituto de Investigación de Recursos Biológicos Alexander von Humboldt">IAvH-E</abbrev>
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) labeled: “ Pasto (Nar.) / 29 - X - 58 ” [white printed label, second line handwritten] // “
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<collectorName id="D240F90414C7B5806DD1EFAE8BB35F31">B. Yanguatin</collectorName>
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/
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<emphasis id="FA53D834F905FA83F8DDC65F2F69D877" italics="true">Suelo paramo</emphasis>
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/ Alt. 3500 mts ” [white handwritten label] // “
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||
<typeStatus id="F9A4E24904523CC3AF46B9BF1F9785DE" type="holotype">Holotypus</typeStatus>
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<specimenCount id="3B8DC22DDAEE02AF93705B621C918E88" count="1" type="male">♂</specimenCount>
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/
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<taxonomicName id="00536699DCF23C522F6E02E4DC35761D" authorityName="Clavijo-Bustos & Castro-Vargas & Neita Moreno" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="paramicola" status="sp. nov.">
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<emphasis id="B3B4FE02EB6F9B56B0AF2A7314645F6A" italics="true">Manodactylus paramicola</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="3109C601487F673B576E69A15D400F5B" rank="species">n. sp.</taxonomicNameLabel>
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||
/
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||
<location id="6A5BC5BB720101268EDB5FED0E095AB5" LSID="urn:lsid:plazi:treatment:FE891834304252368DBA79D5AA7866F3:6A5BC5BB720101268EDB5FED0E095AB5" country="Colombia" municipality="Instituto Humboldt" name="Clavijo-Bustos">Clavijo-Bustos</location>
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, Castro-Vargas & Neita Moreno, / 2024 ” [red printed label] // “
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<collectingMunicipality id="56F98D1F9612B060A187F31178780D79">Instituto Humboldt</collectingMunicipality>
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/
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||
<collectingCountry id="9BC9E490CBF4393C80C82675D8EBD41D" name="Colombia">Colombia</collectingCountry>
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||
/
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||
<abbrev id="ABBRID0ELRAE" xlink_title="Sección de Entomología, Colecciones Biológicas del Instituto de Investigación de Recursos Biológicos Alexander von Humboldt">IAvH-E</abbrev>
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||
- 266051 ” [white printed label with QR code]
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||
</materialsCitation>
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||
.
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||
<emphasis id="90A77398E7597BE7422C929AB0E41E10" bold="true" italics="true">
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||
<typeStatus id="9FA3AB1793B5EE80554B2CC6EBC64227" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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:
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<specimenCount id="9527EE8F99355651FA25ED23188A2C82" count="2" type="male">2 males</specimenCount>
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(
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||
<named-content id="43F60DF85470CE164EF30CBC465BBAE1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-de-nari%C3%B1o" xlink_title="Colección Entomológica de la Universidad de Nariño">
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<collectionCode id="465B2DEDE348281D9DBDFD5C9767CED6">CEUN</collectionCode>
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</named-content>
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) labeled like the
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<typeStatus id="7C21E614A131F9ABF423C62A2D1575D5" type="holotype">holotype</typeStatus>
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, except by: “
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<typeStatus id="2FA81ACEC9CBBF514E7A770DF2A93CAA" type="paratype">Paratypus</typeStatus>
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<specimenCount id="E7CC5DE598714C083A6904B92A8582A2" count="1" type="male">♂</specimenCount>
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/ … ” [yellow printed label];
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<specimenCount id="60132F56D0676F116A58AB354DEE676E" count="1" type="male">1 male</specimenCount>
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(
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<named-content id="4C9ECD383CB619686466DC55A2C229C4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">
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<collectionCode id="40533B92C60C434BB1FA1B0FCFC1D843">CMNC</collectionCode>
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</named-content>
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) labeled: “
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<collectingCountry id="B4C4C4871CBF7849DE0A070004749D4D" name="Colombia">Colombia</collectingCountry>
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,
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<collectingRegion id="99856CE0859D603FDB8CB670D101351B" country="Colombia" name="Cauca">Cauca</collectingRegion>
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, San Sebastian, / Valencia,
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4RAE" specific-use="{"type":"Point","coordinates":[-76.670081,1.900042]}">
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||
<geoCoordinate id="D6FE0774414BA18CEB3F679851EEF258" degrees="1" direction="north" minutes="54" orientation="latitude" precision="1" seconds="0.15" value="1.9000417">1 ° 54 ’ 0.15 " N</geoCoordinate>
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,
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<geoCoordinate id="389E5CDE789CAADEAB2FA9267F929E57" degrees="76" direction="west" minutes="40" orientation="longitude" precision="1" seconds="12.29" value="-76.670074">76 ° 40 ’ 12.29 " W</geoCoordinate>
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||
</named-content>
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, WGS 84,
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||
<quantity id="4012A7D658953FF7F3774DC459918B66" metricMagnitude="3" metricUnit="m" metricValue="2.95" unit="m" value="2950.0">2950 m</quantity>
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||
, Cap. manual, /
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||
<date id="A00DCB0AC976FBDB6E26FEA3863985C8" value="2018-07-09">2018-09 - 07</date>
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||
, D. E. Martínez-Revelo ” [white printed label] // “
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||
<collectingRegion id="38108E8767A8A5CB1E708417725A745A" country="Colombia" name="Cauca">Cauca</collectingRegion>
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Valencia / 7 - sep- 18 ” [white handwritten label], and the same yellow
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<typeStatus id="72026AD3A3E4BEA0710B66F9A73D51B0" type="paratype">paratype</typeStatus>
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||
label.
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||
</paragraph>
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||
<caption id="A89FD64DECB71AD3806C46C07DA6FAD2" ID-DOI="10.3897/dez.71.118923.figure2" ID-arpha="4CB6F1D8-9CD0-5EDF-86FA-BF0DE23BF5DE" httpUri="https://binary.pensoft.net/fig/1104275" startId="F2">
|
||
<paragraph id="33DF1BCFC69803620CFB6892AF77F93C">
|
||
<label id="1B139CC91DE882D345A53E9767A03918">Figure 2.</label>
|
||
</paragraph>
|
||
<paragraph id="6D9C988F0C25CF286C6F8BFBB97172B3">
|
||
<taxonomicName id="3E16641C6FA6B1EF86112F07DE447DD8" authorityName="Clavijo-Bustos & Castro-Vargas & Neita Moreno" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="paramicola" status="sp. nov.">
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<emphasis id="ADA75FEDD1C8956E8AB1622C5CFD99AD" italics="true">Manodactylus paramicola</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="77058251F20DFF995BD37A83F74D1B64" rank="species">sp. nov.</taxonomicNameLabel>
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||
, male holotype.
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||
<emphasis id="21722D9932302BEBD97280B057D35694" bold="true">A.</emphasis>
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||
Dorsal view;
|
||
<emphasis id="D69615EAF1BF5A12E610273998AF4BF3" bold="true">B.</emphasis>
|
||
Lateral view;
|
||
<emphasis id="CEFE7298D942CCCF3554682669DFD1F2" bold="true">C.</emphasis>
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||
Ventral view;
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||
<emphasis id="E4EF5043B39EB03F5B1592E59184A415" bold="true">D.</emphasis>
|
||
Aedeagus in lateral view;
|
||
<emphasis id="3CE04BD7040F2AF717BB8CF3B8692028" bold="true">E.</emphasis>
|
||
Parameres in dorsal view;
|
||
<emphasis id="7E3D594067449AEC2BB4A5EA63595289" bold="true">F.</emphasis>
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||
Parameres in frontal view;
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||
<emphasis id="D30D1DD40F68D14DCC033C60D2AA545B" bold="true">G.</emphasis>
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Spiculum gastrale in dorsal view.
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</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EITAE" type="description">
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||
<paragraph id="05FE94A56A54BBC2B260037E365D982C">
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||
<heading id="C46EB74BC2ED2BBCE989FDF4D918DA17" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="6DCC36B2D3B5CD25128977BCB3E5294F">
|
||
Male. Total length:
|
||
<quantity id="8557170622CA9C01C22D120A9F89284C" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="9.5" metricValueMin="7.9" unit="mm" value="8.7" valueMax="9.5" valueMin="7.9">7.9–9.5 mm</quantity>
|
||
(
|
||
<typeStatus id="0A4C1D0CE3F70EF6B541AE0DDF469EF5" type="holotype">holotype</typeStatus>
|
||
:
|
||
<quantity id="9BBF3A83E0AD16DBA3355B389A2D1E9D" metricMagnitude="-3" metricUnit="m" metricValue="9.3" unit="mm" value="9.3">9.3 mm</quantity>
|
||
), maximum width of pronotum: 3.0–
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||
<quantity id="BA27ED987A31A6A40E3D8F2D72153675" metricMagnitude="-3" metricUnit="m" metricValue="3.6" unit="mm" value="3.6">3.6 mm</quantity>
|
||
(
|
||
<typeStatus id="D0E49B85B16FE0EBE08DFB7325098247" type="holotype">holotype</typeStatus>
|
||
:
|
||
<quantity id="D4957E38458F85F6C92A7CC748BEF63C" metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
|
||
), width at the base of pronotum: 2.0–
|
||
<quantity id="D402B4E558B4D968DF0758AFB57AE224" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.5 mm</quantity>
|
||
(
|
||
<typeStatus id="34AA948B9926D21F330651175343FFF3" type="holotype">holotype</typeStatus>
|
||
:
|
||
<quantity id="7365F4D9C68914BBDBC7DEC14CAA5437" metricMagnitude="-3" metricUnit="m" metricValue="2.4" unit="mm" value="2.4">2.4 mm</quantity>
|
||
), width at the base of elytra:
|
||
<quantity id="3EAB378C1D5BA903CFEAE8FE629DF102" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.3" metricValueMin="2.8" unit="mm" value="3.05" valueMax="3.3" valueMin="2.8">2.8–3.3 mm</quantity>
|
||
(
|
||
<typeStatus id="654B100B8F42F37FEB621BE3C165DDC2" type="holotype">holotype</typeStatus>
|
||
:
|
||
<quantity id="6D50E21EDBCDFA7D3880B00DC0CEA97D" metricMagnitude="-3" metricUnit="m" metricValue="3.3" unit="mm" value="3.3">3.3 mm</quantity>
|
||
). Color black, shiny, with rounded micropunctures when viewed under high magnification; antennae with antennomeres 1–6 dark brown and antennomeres 7–9 (antennal club) black; apex of femora, and tibiae and tarsi, dark brown. Body elongate and slender, subparallel.
|
||
<emphasis id="48A20E3E4E853A4BE118287D4B00DFC6" bold="true" italics="true">Head</emphasis>
|
||
: Clypeus trapezoidal, anterior border emarginate medially, borders on each side of medial emargination narrowly rounded, slightly angulated and upturned; clypeus and frons forming a subpentagonal concave area, surface with small, coarse, rounded punctures densely and irregularly distributed, separated by less than one time one puncture diameter, giving a rugose appearance, and each puncture bearing a short, slender pale-yellow seta; frons with a smooth raised narrow area extending from anterior angles to the vertex, and delimiting the subpentagonal concave area. Fronto-clypeal suture not well-marked. Vertex smooth medially, laterally with punctures similar to those of the surface of clypeus and frons. Eyes small, interocular distance equal to 6.2 times the dorsal width of the eye. Ocular canthus very short and narrow, dorsally covered with small, coarse, rounded punctures densely and irregularly distributed, each bearing a moderately long, thick reddish-yellow seta. Antennae with nine antennomeres, antennal club equal to the combined length of antennomeres 2–6.
|
||
<emphasis id="7AC59EBC2E4CCBE7D66DA1AFF1F2F3C3" bold="true" italics="true">Mouthparts</emphasis>
|
||
: Mandibles triangular, sclerotized, molar lobes slightly asymmetric; long setae present in the apical half of outer edge and in the inner edge, in a membranous lobe apically setose; surface of molar lobes with ridges. Maxillae with the cardo projected horizontally; maxillary palps with four palpomeres; lacinia apically membranous, with a tuft of setae; galea with eight teeth and covered of setae apically. Labium with mentum oblong, longer than wide, slightly narrowing anteriorly, anterior edge truncate, ventral surface with large, thick setae; prementum with two labial palps, each with three palpomeres. Labrum with the anterior edge of distal epipharynx slightly emarginate medially; chaetopariae with several rows of long setae, laeotorma and dexiotorma long.
|
||
<emphasis id="7DEF8C4BE37AEFEF4ACD120F8894FA8B" bold="true" italics="true">Pronotum</emphasis>
|
||
: Pronotal disc convex, diamond shaped, strongly wider medially, and wider than long. Surface medially with few small, rounded punctures sparsely and irregularly distributed, separated by 3–5 times one puncture diameter, each puncture bearing a short, slender pale-yellow seta. Borders margined, with an irregular row of small, rounded punctures; the row has less punctures over the anterior and posterior borders than over the anterior angles; each puncture on anterior and posterior borders bearing a short, slender pale-yellow seta, otherwise, each puncture bearing a moderately long, thick reddish-yellow seta, and over the anterior angles with few punctures bearing a long, thick reddish-yellow seta. Anterior angles acute, posterior angles rounded.
|
||
<emphasis id="91E1B705EC843CA1D7A8285AAB1BA2C1" bold="true" italics="true">Scutellum</emphasis>
|
||
: Scutellar shield slightly longer than wide; surface laterally with few small, rounded punctures sparsely and irregularly distributed, separated by one to less than one times one puncture diameter, each puncture bearing a short, slender pale-yellow seta. Limit between scutellar shield and mesoscutum not evident.
|
||
<emphasis id="83F3C1208DD062E8CCA50E24B486E0D1" bold="true" italics="true">Hind wings</emphasis>
|
||
: Fully developed.
|
||
<emphasis id="CA528E5D102426E5FE709C3A8E9C4F45" bold="true" italics="true">Elytra</emphasis>
|
||
: Elytral base wider than pronotal base. Humeral and apical calluses slightly prominent, rounded. Each elytron with five striae between suture and humeral callus,
|
||
<quantity id="C126CBB0B16E4645057A3115E33EE3FB" metricMagnitude="-1" metricUnit="m" metricValue="2.54" unit="in" value="10.0">10 in</quantity>
|
||
total, but lateral striae less evident; striae disappearing over the apical declivity; each stria deep, with a regular row of small, rounded punctures distanced by 3–5 times one puncture diameter. Interstriae strongly convex. Surface of interstriae, humeral and apical calluses with scattered, small, rounded punctures. Some punctures in the surface, including some in the striae, bearing a short, slender pale-yellow setae.
|
||
<emphasis id="24EAA4FC9DFFDD59365841FC8F17125B" bold="true" italics="true">Hypomeron</emphasis>
|
||
: Hypomeral disc convex. Surface anteriorly smooth, otherwise covered with small, rounded punctures sparsely and irregularly distributed, separated by 5–8 times one puncture diameter; each puncture bearing a moderately long, pale-yellow seta.
|
||
<emphasis id="6DDC0478BE4E80C11D4BE72CA3964E70" bold="true" italics="true">Prosternum</emphasis>
|
||
: Disc with two anterior, longitudinal sulci.
|
||
<emphasis id="FBEEA89A522BAEDEF987970DEDB86F4D" bold="true" italics="true">Mesoventrite</emphasis>
|
||
: Disc flat in the middle. Surface with few small, rounded punctures separated by 1–2 times one puncture diameter, each puncture bearing a short, slender pale-yellow seta.
|
||
<emphasis id="8CCA66A153755D57B9854CA1858002A9" bold="true" italics="true">Mesepimerum and metepisternum</emphasis>
|
||
: Surfaces with scattered small, rounded punctures, each puncture bearing a short, slender pale-yellow seta.
|
||
<emphasis id="890785051E26704196415B2CC8F05505" bold="true" italics="true">Metaventrite</emphasis>
|
||
: Disc flat and densely setose in the central area. Surface of central area covered with small, rounded punctures densely and irregularly distributed, separated by less than one time one puncture diameter, each bearing a moderately long, slender pale-yellow seta. Surface of lateral arms similar to that of central area, but with evidently sparser punctures, thus less setose, and smooth externally.
|
||
<emphasis id="CC686EA7B730CDF5076A176368F5F8F8" bold="true" italics="true">Hind wings</emphasis>
|
||
: Apical edge rounded; RP
|
||
<subScript id="1D38FE573C01A258B48D9811B1B08B09">3</subScript>
|
||
y RA
|
||
<subScript id="5C2F3B54C73A73F2F6666047629311AB">4</subScript>
|
||
+ RP
|
||
<subScript id="A76F73A2226E4477D9399FADE7865E35">1</subScript>
|
||
apically parallel; MSP projected toward the wing margin;
|
||
<collectionCode id="B7C4A5C4397ED041DE61566F25F7DD31" country="0" name="Mohonk Preserve, Inc.">MP</collectionCode>
|
||
<subScript id="F7DA698ABE8DDE5DA2FDCBBBC0ECE3E7">3 + 4</subScript>
|
||
short, not projected apically.
|
||
<emphasis id="92FA5DBAA8343A719E6F3608747A8F4E" bold="true" italics="true">Legs</emphasis>
|
||
: Femora subequal in length to their respective tibiae; surface almost completely smooth, with scattered small, rounded punctures, each bearing a short, slender pale-yellow seta; anterior and posterior border moderately setose in the basal third. Tibiae shorter than their respective tarsi. Tarsal claws bifid, ventral part slightly thicker, shorter, and more downward curved than dorsal part; each claw over the basal third with an acute triangular tooth. Protibia along the outer edge with two teeth distributed in the apical third, apical tooth larger than basal; inner angle slightly projected downward; protibia gradually widening from base to apex but narrowing from the inner edge in the apical third; surface smooth except for a few scattered punctures and a regular row of small, rounded punctures separated by 2–4 times one puncture diameter, each puncture bearing a short, slender pale-yellow seta; anterior and posterior borders with scattered moderately long, slender pale-yellow setae; lacking apical spur. Protarsomere 1 subequal to the combined length of protarsomeres 2–3, protarsomere 2–4 subequal in length, and protarsomere 5 subequal to the combined length of protarsomeres 3–4; internal face of protarsomeres with few scattered short to moderately long, thick reddish-yellow seta. Mesotibiae with a more or less evident ventral transverse carina near the apical third of its length; surface with few small, rounded punctures sparsely distributed, each bearing a short or moderately long, thick reddish-yellow seta; apex over the ventral surface with intercalate extremely short and short, thick reddish-yellow spinules; two subequal apical spurs present, the dorsal spur slightly longer than the ventral, both about three quarters of the length of the mesotarsomere 1. Mesotarsomere 1 slightly longer than mesotarsomere 2, mesotarsomere 2 slightly longer than mesotarsomere 3, mesotarsomeres 3 and 4 subequal in length, and mesotarsomere 5 slightly shorter than the combined length of the mesotarsomere 3–4; internal face of mesotarsomeres with a few scattered short to moderately long, thick reddish-yellow seta. Metatibiae surface with a few small, rounded punctures sparsely distributed, each bearing a short or moderately long, thick reddish-yellow seta; apex over the ventral surface with intercalate extremely short and short, thick reddish-yellow spinules; one apical spur present, about a half of the length of the metatarsomere 1. Metatarsomere 1 slightly longer than metatarsomere 2, metatarsomere 2 slightly longer than metatarsomere 3, metatarsomeres 3 slightly longer than metatarsomere 4, and metatarsomere 5 slightly shorter than the combined length of the metatarsomere 3–4; internal face of metatarsomeres with few scattered short or moderately long, thick reddish-yellow seta (thicker than the slender setae mentioned, but not as spine-like setae).
|
||
<emphasis id="ECA8F441A12E441EA9AF814338D6D967" bold="true" italics="true">Abdomen</emphasis>
|
||
: Abdominal ventrites 2–5 densely setose medially, surface covered with small, rounded punctures densely and irregularly distributed, separated by less than 1 times one puncture diameter, each bearing a moderately long, slender pale-yellow seta, some few bearing longer setae; surface laterally similar, but with evidently sparser punctures, thus sparsely setose; anterior border of each of these ventrites densely setose. Ventrite 5 longer than the others. Ventrite 6 almost smooth, with few small, rounded punctures bearing a long, slender pale-yellow seta; anterior border with moderately long, thick reddish-yellow setae. Pygidium vertical, strongly convex; surface with few small, rounded punctures sparsely and irregularly distributed, separated by 2–5 times one puncture diameter, each bearing a short, slender pale-yellow seta; setae convergent over the longitudinal central area, and punctures sparser externally.
|
||
<emphasis id="3C21C90D3CF8DDCD45E83F6386E47F07" bold="true" italics="true">Genitalia</emphasis>
|
||
: Hemisternite 9 covered with small, rounded punctures densely and irregularly distributed, separated by one time one puncture diameter; anterior edge truncate and slightly emarginate medially. Spiculum lateral arms short, the spiculum gastrale around two times the length of a spiculum lateral arm, and the cranial part with apex rounded. Phallobase cylindrical, 2.5 times the length of parameres. Parameres fused at the base, short, apically curved and slightly downward, with the apex rounded; the inner edge of each paramere sinuous, with an evident, strong indentation in the basal third.
|
||
</paragraph>
|
||
<caption id="8F90C138EA91DB1761E79B1A99E81188" ID-DOI="10.3897/dez.71.118923.figure3" ID-arpha="EB18B956-0325-50EB-90DD-74D7AAA49897" httpUri="https://binary.pensoft.net/fig/1104276" startId="F3">
|
||
<paragraph id="1B093C59321725133EDC022AE3108F31">
|
||
<label id="650615399321CB22C4685CA9AD072C39">Figure 3.</label>
|
||
</paragraph>
|
||
<paragraph id="850DFD34836966FB75F58421E1CE9546">
|
||
<taxonomicName id="CF6271AE82CBD0B8A66197CA443848C5" authorityName="Clavijo-Bustos & Castro-Vargas & Neita Moreno" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="paramicola" status="sp. nov.">
|
||
<emphasis id="FCADAE4800FA2782DCA71D31CAFCD801" italics="true">Manodactylus paramicola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2432839EEED99F3CEC3E9F10C43CA874" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male paratype.
|
||
<emphasis id="E6C5F6EDE2022E21DE2AFD4C2A8EEFFD" bold="true">A.</emphasis>
|
||
Hind wing in dorsal view;
|
||
<emphasis id="4C4A5E037F8CCB1CC734FE31B257A69E" bold="true">B.</emphasis>
|
||
Epipharynx in ventral view;
|
||
<emphasis id="E46309388B1477639320476B6BBC8CA6" bold="true">C.</emphasis>
|
||
Left maxilla in ventral view;
|
||
<emphasis id="4743B626FCD32FAB4EDBC795E3FDE45F" bold="true">D.</emphasis>
|
||
Left maxilla in dorsal view;
|
||
<emphasis id="3C9727C82D7DC666CD7E965BD7A2C4E9" bold="true">E.</emphasis>
|
||
Labium in ventral view;
|
||
<emphasis id="BB4A5A4938572B0AA1E2C5C15A8420DA" bold="true">F.</emphasis>
|
||
Left mandible in dorsal view;
|
||
<emphasis id="3108C3B2C0797D5C1B99E40280802372" bold="true">G.</emphasis>
|
||
Left mandible in lateral internal view;
|
||
<emphasis id="949E5924698A9BA3B7D2F68F1AA09D59" bold="true">H.</emphasis>
|
||
Left mandible in ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4AA48246B86AA1131970E874B1CC5C9C">
|
||
<emphasis id="7C671495BB1494E6EE2A28F0BBA16D91" bold="true">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EUWAE" type="diagnosis">
|
||
<paragraph id="8A1EC2019450D9D8A36065C93E0077AF">
|
||
<heading id="128C7CA22345CA773AAF11FC85C04412" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="EE70CAF840B2D315E575E6220647C36B">
|
||
<taxonomicName id="084A920DC1128023C0365C2782C825E8" authorityName="Clavijo-Bustos & Castro-Vargas & Neita Moreno" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="paramicola">
|
||
<emphasis id="6349B20AEA7B4493359C846065882084" italics="true">Manodactylus paramicola</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="683F29EEFEEE2D5D375313F79CC6B98E" authorityName="Ohaus" authorityYear="1909" class="Insecta" family="Scarabaeidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gaujoni">
|
||
<emphasis id="DE47E952E01A0937A53283F55B9196C7" italics="true">M. gaujoni</emphasis>
|
||
</taxonomicName>
|
||
the overall appearance (see Diagnosis for
|
||
<taxonomicName id="60639745AF575EC67DA27C05F8DFD807" authorityName="Ohaus" authorityYear="1909" class="Insecta" family="Scarabaeidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gaujoni">
|
||
<emphasis id="F12B7E3409B0F5CD4EC27E6EC79C32C3" italics="true">M. gaujoni</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="9058D167A0EF26EE48EE068F1CA5B80B" authorityName="Clavijo-Bustos & Castro-Vargas & Neita Moreno" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="paramicola">
|
||
<emphasis id="11A79EFE9005658244164B1B3B680AB8" italics="true">Manodactylus paramicola</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="7C6A58396EDE613DD89C990620E3DA87" authorityName="Ohaus" authorityYear="1909" class="Insecta" family="Scarabaeidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gaujoni">
|
||
<emphasis id="7F3B8CFCF6D94DFBEFFBA5D22B4AC2FC" italics="true">M. gaujoni</emphasis>
|
||
</taxonomicName>
|
||
by having the femora, tibiae, and tarsi very dark brown; clypeus emarginate medially, with borders on each side of medial emargination narrowly rounded, slightly angulated and upturned; the subpentagonal area formed by clypeus and frons concave; the pronotum also with small, rounded, setigerous punctures on the borders but few also present medially in the disc, separated by 3–5 times one puncture diameter; each elytral stria deep, with a regular row of small, rounded punctures distanced by 3–5 times one puncture diameter; and interstriae strongly convex.
|
||
</paragraph>
|
||
<caption id="400A2EEECACD9BD352B2184B619E1AC0" ID-DOI="10.3897/dez.71.118923.figure4" ID-arpha="017F09DA-2493-5F1A-AE74-D12460703A73" httpUri="https://binary.pensoft.net/fig/1104277" startId="F4">
|
||
<paragraph id="025765014DFCEE5DD580B5D988C2870D">
|
||
<label id="A18295E71F0EF58525ECA7D9CAA9E4F4">Figure 4.</label>
|
||
</paragraph>
|
||
<paragraph id="6529D0AF35B4E93D2D8AD15E7804B3E5">
|
||
<taxonomicName id="77919105FF4162DB77B2E9C49142207C" authorityName="Clavijo-Bustos & Castro-Vargas & Neita Moreno" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="paramicola" status="sp. nov.">
|
||
<emphasis id="30B7755299CB693907969FA84086B8FC" italics="true">Manodactylus paramicola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3E4A1F406486ED1D50E1090D5384736" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male holotype.
|
||
<emphasis id="3AE65C2620D77E2EF36C2C82D62ECEE6" bold="true">A.</emphasis>
|
||
Protibia in dorsal view;
|
||
<emphasis id="524EC8B8D3220F71932847A76A8CEAA2" bold="true">B.</emphasis>
|
||
Detail of the downward projection of the inner angle of protibia;
|
||
<emphasis id="AE4A761F42E91277811827C2F0B2EAA2" bold="true">C.</emphasis>
|
||
Apex of protibia;
|
||
<emphasis id="AC2DD0EE2E2F3A65E0635758956A98C1" bold="true">D.</emphasis>
|
||
Apex of mesotibia;
|
||
<emphasis id="E425966257480311D6BF6B0B065CFF39" bold="true">E</emphasis>
|
||
, Apex of metatibia. Black arrows point to downward projection of the inner angle of protibia.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ETZAE" type="etymology">
|
||
<paragraph id="934BA2CED33FE4A4B8D08C87F1E3F8C3">
|
||
<heading id="124301283B8BF7C2FC338235475E43D6" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="CE10B913F3A145B4D1F9C509A53D6F15">
|
||
Epithet compound, from the Spanish noun
|
||
<emphasis id="73A1FBDE5CD7B8C11AD0310EFB39FE39" italics="true">Páramo</emphasis>
|
||
for Paramo ecosystem, and the Latin suffix –
|
||
<emphasis id="2896279A2899E2DD2C9318108134E8FE" italics="true">icola</emphasis>
|
||
meaning inhabitant. The specific name means ‘ inhabitant of Paramo’, in allusion to the ecosystem where the specimens were collected.
|
||
</paragraph>
|
||
<caption id="AAF9C39FE16B4D8592FE54E147B8FAC2" ID-DOI="10.3897/dez.71.118923.figure5" ID-arpha="998E23A6-B664-5B19-86F3-50D217BDDA40" httpUri="https://binary.pensoft.net/fig/1104278" startId="F5">
|
||
<paragraph id="66822E40B440682CF1087666EE937663">
|
||
<label id="0509DDD62EF105D33C2F7373C8345740">Figure 5.</label>
|
||
</paragraph>
|
||
<paragraph id="D2AB64567433A45336A6EE10216D0B1F">
|
||
<taxonomicName id="163423C05043B440A8FCC297233A2D39" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined-A">
|
||
<emphasis id="42A74DAE7E76B01C03C25CAA00A2CC96" italics="true">Manodactylus</emphasis>
|
||
species.
|
||
<emphasis id="CFF60B0AB2911DF2BE90406768ECCB65" bold="true">A, B.</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="E54FC4B3F18992F036A8903ACDCF84F9" authority="Ohaus, 1909" authorityName="Ohaus" authorityYear="1909" class="Insecta" family="Scarabaeidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gaujoni">
|
||
<emphasis id="FB8D81C653BB4BCD9F3F18BE5A868472" italics="true">Macrodactylus gaujoni</emphasis>
|
||
<bibRefCitation id="CA3CEF2A7FF192C5D836732DF0C5798D" DOI="10.1002/mmnd.48019090312" author="Ohaus F" journalOrPublisher="Stettiner Entomologische Zeitung" pagination="3-139" refId="B4" refString="Ohaus F (1909) Bericht uber eine entomologische Studienreise in Sudamerika. Stettiner Entomologische Zeitung 70: 3 - 139. https: // doi. org / 10.1002 / mmnd. 48019090312" title="Bericht über eine entomologische Studienreise in Südamerika." volume="70" year="1909">Ohaus, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
, male neotype =
|
||
<taxonomicName id="66D3593EF1F2713038E3C5B11764C3CC" authority="Moser, 1919" authorityName="sensu Moser" authorityYear="1919" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gaujoni">
|
||
<emphasis id="ACD13F6DAAEE26A9A0C4D737167398ED" italics="true">Manodactylus gaujoni</emphasis>
|
||
<bibRefCitation id="531732855FA694CE84C3B7AC281D678A" author="Moser J" journalOrPublisher="Stettiner Entomologische Zeitung" pagination="3-64" refId="B3" refString="Moser J (1919) Beitrag zur Kenntnis der Melolonthiden (Col.). (IX). Stettiner Entomologische Zeitung 80: 3 - 64." title="Beitrag zur Kenntnis der Melolonthiden (Col.). (IX)." volume="80" year="1919">Moser, 1919</bibRefCitation>
|
||
</taxonomicName>
|
||
, male lectotype;
|
||
<emphasis id="208AD08A10820D244CE20AA7DCCBD9CA" bold="true">C, D.</emphasis>
|
||
<taxonomicName id="48D17E97D289076600C5A32F9FC45646" authorityName="Clavijo-Bustos & Castro-Vargas & Neita Moreno" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="paramicola" status="sp. nov.">
|
||
<emphasis id="993DCC1498C1D6CF2E988793593B9F58" italics="true">Manodactylus paramicola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8D4BC9957A1892F70C7B029F645C68CB" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male holotype.
|
||
<emphasis id="3B8D782D2EC2F3F8DCCC5DEEE01EC4DA" bold="true">A, C.</emphasis>
|
||
Pronotum in dorsal view;
|
||
<emphasis id="CE212AA8312F59685F5CB339021C2590" bold="true">B, D.</emphasis>
|
||
Head in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E12AE" type="distribution">
|
||
<paragraph id="345C623BF29C6B2D0BF199D760386523">
|
||
<heading id="AF7B3944C63283B7930E3884776F3764" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="9A51ED5300FCA50481FFA1C47BBDE6F6">
|
||
<collectingCountry id="401AB68BEB03E37B5469D0BC93629F4E" name="Colombia">Colombia</collectingCountry>
|
||
, departments of
|
||
<collectingRegion id="17F3844E6D5564DEA3BBCA63BC1BB13F" country="Colombia" name="Cauca">Cauca</collectingRegion>
|
||
and
|
||
<collectingRegion id="082E751D1C5F0F4974751D10425E59F6" country="Colombia" name="Narino">Nariño</collectingRegion>
|
||
(Fig.
|
||
<figureCitation id="605373B605D40C7C4E0529A453EDF5E9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution map of Manodactylus. Macrodactylus gaujoni Ohaus, 1909, male neotype = Manodactylus gaujoni Moser, 1919, male lectotype. A star is used for approximate holotype locality when possible; ‘? ’ to denote unverified records; red color is used for Manodactylus gaujoni Moser, 1919; orange color is used for Macrodactylus gaujoni Ohaus, 1909; yellow color is used for Manodactylus paramicola sp. nov." figureDoi="10.3897/dez.71.118923.figure6" httpUri="https://binary.pensoft.net/fig/1104279">6</figureCitation>
|
||
). Three of the specimens were collected in Pasto, department of
|
||
<collectingRegion id="584C4C7EC5786CAF8D55991D87228CF6" country="Colombia" name="Narino">Nariño</collectingRegion>
|
||
, with the information ‘ Paramo ground’ (
|
||
<emphasis id="7058E977237ACE67965E0B3D502D0382" italics="true">Suelo paramo</emphasis>
|
||
) on the label, referring to the ecosystem in which the specimens were found. The fourth specimen was collected in San Sebastián, a small, populated place located less than 10 kilometers from the Páramo de las Papas in the department of
|
||
<collectingRegion id="596B7369EEF75ADE5BC18247B957C512" country="Colombia" name="Cauca">Cauca</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<caption id="D8EBB28445EBBABF780C181E89365CD2" ID-DOI="10.3897/dez.71.118923.figure6" ID-arpha="309D773D-A54F-5738-ACF6-7F6CB0DC36A9" httpUri="https://binary.pensoft.net/fig/1104279" startId="F6">
|
||
<paragraph id="DD2D75638413C30C518DB21EDE989765">
|
||
<label id="8220CC41F39FC1A65E3D9B98EAFC5362">Figure 6.</label>
|
||
</paragraph>
|
||
<paragraph id="DB7CBD9207789F84A2FDFF6729E99695">
|
||
Distribution map of
|
||
<taxonomicName id="5DD857739465029F118EB35652DE56AC" authorityName="Moser" authorityYear="1919" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="45B1B817495DE7F9083247F91CEDB4C7" italics="true">Manodactylus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="8443D558CEEDB0866C48D75F7E5F55AC" authority="Ohaus, 1909" authorityName="Ohaus" authorityYear="1909" class="Insecta" family="Scarabaeidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gaujoni">
|
||
<emphasis id="20F8AA6A2E5DB7DCDA62DCC632BFE677" italics="true">Macrodactylus gaujoni</emphasis>
|
||
<bibRefCitation id="2BEA550C692D707737367B06936C7E5C" DOI="10.1002/mmnd.48019090312" author="Ohaus F" journalOrPublisher="Stettiner Entomologische Zeitung" pagination="3-139" refId="B4" refString="Ohaus F (1909) Bericht uber eine entomologische Studienreise in Sudamerika. Stettiner Entomologische Zeitung 70: 3 - 139. https: // doi. org / 10.1002 / mmnd. 48019090312" title="Bericht über eine entomologische Studienreise in Südamerika." volume="70" year="1909">Ohaus, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
, male neotype =
|
||
<taxonomicName id="CDCDB894FDF86F87B2783A53429F6612" authority="Moser, 1919" authorityName="sensu Moser" authorityYear="1919" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gaujoni">
|
||
<emphasis id="9AEC73C691454B8F803859BA08D1B5FB" italics="true">Manodactylus gaujoni</emphasis>
|
||
<bibRefCitation id="861074F30B2DC29644B93C92E721C7E7" author="Moser J" journalOrPublisher="Stettiner Entomologische Zeitung" pagination="3-64" refId="B3" refString="Moser J (1919) Beitrag zur Kenntnis der Melolonthiden (Col.). (IX). Stettiner Entomologische Zeitung 80: 3 - 64." title="Beitrag zur Kenntnis der Melolonthiden (Col.). (IX)." volume="80" year="1919">Moser, 1919</bibRefCitation>
|
||
</taxonomicName>
|
||
, male lectotype. A star is used for approximate holotype locality when possible; ‘? ’ to denote unverified records; red color is used for
|
||
<taxonomicName id="C4545EFEDCF2182C11301AD0585AAE41" authority="Moser, 1919" authorityName="sensu Moser" authorityYear="1919" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gaujoni">
|
||
<emphasis id="E55977E2DEA316F56FCE56D4527ED67A" italics="true">Manodactylus gaujoni</emphasis>
|
||
<bibRefCitation id="F490911AD2BEC0898C3A9860B208E10E" author="Moser J" journalOrPublisher="Stettiner Entomologische Zeitung" pagination="3-64" refId="B3" refString="Moser J (1919) Beitrag zur Kenntnis der Melolonthiden (Col.). (IX). Stettiner Entomologische Zeitung 80: 3 - 64." title="Beitrag zur Kenntnis der Melolonthiden (Col.). (IX)." volume="80" year="1919">Moser, 1919</bibRefCitation>
|
||
</taxonomicName>
|
||
; orange color is used for
|
||
<taxonomicName id="AE27445FF70D64003C4E883605A8B6C1" authority="Ohaus, 1909" authorityName="Ohaus" authorityYear="1909" class="Insecta" family="Scarabaeidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gaujoni">
|
||
<emphasis id="149B0F97564978F876D1F191086719FE" italics="true">Macrodactylus gaujoni</emphasis>
|
||
<bibRefCitation id="019F5EFAD27CE8F954ACBF89BA73EEB8" DOI="10.1002/mmnd.48019090312" author="Ohaus F" journalOrPublisher="Stettiner Entomologische Zeitung" pagination="3-139" refId="B4" refString="Ohaus F (1909) Bericht uber eine entomologische Studienreise in Sudamerika. Stettiner Entomologische Zeitung 70: 3 - 139. https: // doi. org / 10.1002 / mmnd. 48019090312" title="Bericht über eine entomologische Studienreise in Südamerika." volume="70" year="1909">Ohaus, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
; yellow color is used for
|
||
<taxonomicName id="B828449CD76CE95EB8F6234B3F4E8A0D" authorityName="Clavijo-Bustos & Castro-Vargas & Neita Moreno" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Manodactylus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="paramicola" status="sp. nov.">
|
||
<emphasis id="92F8B70847C5F3F46E838DCF3E90D0CC" italics="true">Manodactylus paramicola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="15A8A949C7FB58DE853C8373BCD03445" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |