278 lines
40 KiB
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278 lines
40 KiB
XML
<document id="929BC47586DC9F639DED7F401228843B" ID="10.11646/zootaxa.4147.4.4" ID-DOI="10.11646/zootaxa.4147.4.4" ID-GBIF-Dataset="db022185-3c2d-4978-8951-d0c23ccdb47a" ID-ISSN="1175-5326" ID-Zenodo-Dep="264988" ID-ZooBank="8B25DA4D-0B7A-4568-8E7C-642EB932F2D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470379029729" checkinUser="plazi" docAuthor="Arango, Stephania Sandoval, Obando, Ranulfo González & Aldrete, Alfonso Neri García" docDate="2016" docId="03FFFA7A6D0DFFBF269A2385FCA554BE" docLanguage="en" docName="zootaxa.4147.4.4.pdf" docOrigin="Zootaxa 4147 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Graphocaecilius mockfordi Arango, Obando & Aldrete, 2016, n. sp." docType="treatment" docVersion="7" lastPageNumber="423" masterDocId="FFC682026D0FFFBD260D2312FFB1534F" masterDocTitle="New species of Graphocaecilius Enderlein (Psocodea: ‘ Psocoptera’: Lachesillidae) from Colombia" masterLastPageNumber="432" masterPageNumber="421" pageNumber="423" updateTime="1698367001650" updateUser="plazi">
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<mods:titleInfo id="945581D0EE4A85BC3D3FC1C32054E364">
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<mods:title id="F834A208A91F0B4F675AED77633D2759">New species of Graphocaecilius Enderlein (Psocodea: ‘ Psocoptera’: Lachesillidae) from Colombia</mods:title>
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<mods:name id="B230FAD5FB07504DDB93CF1A2D66F423" type="personal">
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<mods:namePart id="3408E9DC0C1CDCC7DECCC59285E1FA69">Arango, Stephania Sandoval</mods:namePart>
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<mods:roleTerm id="C1F003D0E801162E0B2BBD82D54838E9">Author</mods:roleTerm>
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<mods:namePart id="E632D59A45E490DC70C778738E7C5849">Obando, Ranulfo González</mods:namePart>
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<mods:name id="6A392FC16E086346E2056C23ACF30A5A" type="personal">
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<mods:roleTerm id="C94163111585971AE465B1BD3C621DF5">Author</mods:roleTerm>
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<mods:namePart id="EB25730737C77801680B822115B00540">Aldrete, Alfonso Neri García</mods:namePart>
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<mods:typeOfResource id="BB873BB9F223C2120FCDED00824A3EA9">text</mods:typeOfResource>
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<mods:title id="D1D272718762646C581A9A71A5B3F875">Zootaxa</mods:title>
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<mods:date id="9C872ECDDA0BC68445B273EF9389A222">2016</mods:date>
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<mods:number id="FAE88E5EB1762613B612A5DFEFC05617">4147</mods:number>
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<mods:classification id="59495EB54D1B2D0589B8A6A1B86788B7">journal article</mods:classification>
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<mods:identifier id="1CDCE3D6F92ED83E6AEF1FE0AE7B63AB" type="DOI">10.11646/zootaxa.4147.4.4</mods:identifier>
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<treatment id="03FFFA7A6D0DFFBF269A2385FCA554BE" ID-DOI="http://doi.org/10.5281/zenodo.6085904" ID-GBIF-Taxon="123514463" ID-Zenodo-Dep="6085904" LSID="urn:lsid:plazi:treatment:03FFFA7A6D0DFFBF269A2385FCA554BE" httpUri="http://treatment.plazi.org/id/03FFFA7A6D0DFFBF269A2385FCA554BE" lastPageNumber="423" pageId="2" pageNumber="423">
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<subSubSection id="C34C18E76D0DFFBF269A2385FEA8539C" pageId="2" pageNumber="423" type="nomenclature">
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<paragraph id="8BE94B6C6D0DFFBF269A2385FD9B53FD" blockId="2.[151,554,151,211]" box="[151,554,151,178]" pageId="2" pageNumber="423">
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<heading id="D0A1FC006D0DFFBF269A2385FD9B53FD" bold="true" box="[151,554,151,178]" fontSize="11" level="1" pageId="2" pageNumber="423" reason="1">
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<emphasis id="B922977E6D0DFFBF269A2385FD9B53FD" bold="true" box="[151,554,151,178]" pageId="2" pageNumber="423">
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<taxonomicName id="4C5630EF6D0DFFBF269A2385FE5053FE" ID-CoL="6KRCP" box="[151,481,151,177]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mockfordi" status="sp. nov.">
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<emphasis id="B922977E6D0DFFBF269A2385FE5053FE" bold="true" box="[151,481,151,177]" italics="true" pageId="2" pageNumber="423">Graphocaecilius mockfordi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2112A056D0DFFBF27E5238AFD9B53FD" box="[488,554,152,178]" pageId="2" pageNumber="423" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE94B6C6D0DFFBF269A23A8FEA8539C" blockId="2.[151,554,151,211]" box="[151,281,186,211]" pageId="2" pageNumber="423">
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(
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||
<figureCitation id="136D57E96D0DFFBF269223A8FEA1539C" box="[159,272,186,211]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 10" captionStartId-0="3.[151,264,1781,1804]" captionStartId-1="4.[151,264,1725,1748]" captionTargetBox-0="[232,1356,214,1729]" captionTargetBox-1="[241,1351,196,1692]" captionTargetId-0="figure@3.[227,1359,188,1729]" captionTargetId-1="figure@4.[239,1353,193,1692]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." captionText-1="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri-0="https://zenodo.org/record/264989/files/figure.png" httpUri-1="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Figs 1–10</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34C18E76D0DFFBF269A2210FD485110" pageId="2" pageNumber="423" type="diagnosis">
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<paragraph id="8BE94B6C6D0DFFBF269A2210FD485110" blockId="2.[151,1437,258,2033]" pageId="2" pageNumber="423">
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<emphasis id="B922977E6D0DFFBF269A2210FEA05254" bold="true" box="[151,273,258,283]" pageId="2" pageNumber="423">Diagnosis.</emphasis>
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Differing from
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<taxonomicName id="4C5630EF6D0DFFBF27D62211FCE05254" authority="Roesler (1940)" authorityName="Roesler" authorityYear="1940" box="[475,849,258,283]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="interpretatus">
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<emphasis id="B922977E6D0DFFBF27D62211FD235255" box="[475,658,259,282]" italics="true" pageId="2" pageNumber="423">G. interpretatus</emphasis>
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Roesler (1940)
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</taxonomicName>
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,
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<emphasis id="B922977E6D0DFFBF25682211FC7B5255" box="[869,970,259,282]" italics="true" pageId="2" pageNumber="423">G. p i c t u s</emphasis>
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<bibRefCitation id="EFC7369D6D0DFFBF25D02210FA885254" author="Mockford" box="[989,1337,258,283]" pageId="2" pageNumber="423" refString="Mockford, E. L. & Sullivan, D. M. (1986) Systematics of the graphocaeciliine psocids with a proposed higher classification of the family Lachesillidae (Psocoptera). Transactions of the American Entomological Society, 112 (1), 1 - 80." type="journal article" year="1986">Mockford & Sullivan (1986)</bibRefCitation>
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, and
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<taxonomicName id="4C5630EF6D0DFFBF23892211FE5B5270" authority="Enderlein (1900)" authorityName="Enderlein" authorityYear="1900" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="trypetoides">
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<emphasis id="B922977E6D0DFFBF23892211FEA55271" italics="true" pageId="2" pageNumber="423">G. trypetoides</emphasis>
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<bibRefCitation id="EFC7369D6D0DFFBF272F2234FE5B5270" author="Enderlein" box="[290,490,294,319]" pageId="2" pageNumber="423" refString="Enderlein, G. (1900) Die Psocidenfauna Perus. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Thiere, 14, 133 - 160." type="journal article" year="1900">Enderlein (1900)</bibRefCitation>
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</taxonomicName>
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, in having the forewings hyaline, lacking pigmented areas. It differs from
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<taxonomicName id="4C5630EF6D0DFFBF23892235FE73522C" authority="Williner (1949)" authorityName="Williner" authorityYear="1949" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="enderleini">
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<emphasis id="B922977E6D0DFFBF23892235FEB8522C" italics="true" pageId="2" pageNumber="423">G. enderleini</emphasis>
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<bibRefCitation id="EFC7369D6D0DFFBF271C2258FE73522C" author="Williner" box="[273,450,330,355]" pageId="2" pageNumber="423" refString="Williner, G. J. (1949) Corrodentios de Bolivia. Revista del Instituto Nacional de Investigacion de las Ciencias Naturales, 1 (4), 95 - 126." type="journal article" year="1949">Williner (1949)</bibRefCitation>
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</taxonomicName>
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, in lacking a brown spot at the fused Rs-M, and in having the areola postica taller and narrower. It differs from
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<taxonomicName id="4C5630EF6D0DFFBF27BA227DFD5B52C8" authority="Williner, 1949" authorityName="Williner" authorityYear="1949" box="[439,746,366,391]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="lacteus">
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<emphasis id="B922977E6D0DFFBF27BA227DFD9652C9" box="[439,551,366,390]" italics="true" pageId="2" pageNumber="423">G. lacteus</emphasis>
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(
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<bibRefCitation id="EFC7369D6D0DFFBF2434227CFD5352C8" author="Williner" box="[569,738,366,391]" pageId="2" pageNumber="423" refString="Williner, G. J. (1949) Corrodentios de Bolivia. Revista del Instituto Nacional de Investigacion de las Ciencias Naturales, 1 (4), 95 - 126." type="journal article" year="1949">Williner, 1949</bibRefCitation>
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)
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</taxonomicName>
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, in having the fused Rs-M shorter, and in having the areola postica distinctly narrower. It differs from
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<taxonomicName id="4C5630EF6D0DFFBF24712281FBDB52E4" authority="Mockford & Sullivan (1986)" authorityName="Mockford & Sullivan" authorityYear="1986" box="[636,1130,402,427]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="peruvianus">
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<emphasis id="B922977E6D0DFFBF24712281FCA852E5" box="[636,793,403,426]" italics="true" pageId="2" pageNumber="423">G. peruvianus</emphasis>
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<bibRefCitation id="EFC7369D6D0DFFBF252C2280FBDB52E4" author="Mockford" box="[801,1130,402,427]" pageId="2" pageNumber="423" refString="Mockford, E. L. & Sullivan, D. M. (1986) Systematics of the graphocaeciliine psocids with a proposed higher classification of the family Lachesillidae (Psocoptera). Transactions of the American Entomological Society, 112 (1), 1 - 80." type="journal article" year="1986">Mockford & Sullivan (1986)</bibRefCitation>
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</taxonomicName>
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, in having the hypandrium with a well defined posterior projection, in having the phallosome anteriorly unstemmed, in having the endophallus distinctly different (compare
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||
<figureCitation id="136D57E96D0DFFBF27EE22C8FD9752BC" box="[483,550,474,499]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1781,1804]" captionTargetBox="[232,1356,214,1729]" captionTargetId="figure@3.[227,1359,188,1729]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264989/files/figure.png" pageId="2" pageNumber="423">
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Fig.
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<quantity id="4CAEE6896D0DFFBF241522C9FD9752BC" box="[536,550,475,499]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="423" unit="in" value="4.0">4</quantity>
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</figureCitation>
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in this paper with
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<figureCitation id="136D57E96D0DFFBF250D22C8FCE352BC" box="[768,850,474,499]" captionStart="FIGURES 21 – 25" captionStartId="9.[151,264,1781,1804]" captionTargetBox="[247,1374,262,1764]" captionTargetId="figure@9.[225,1376,194,1764]" captionTargetPageId="9" captionText="FIGURES 21 – 25. Graphocaecilius turneri (Male). 21. Fore- and hind wings. 22. Front view of head. 23. Clunium, paraprocts and epiproct. 24. Phallosome. 25. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264993/files/figure.png" pageId="2" pageNumber="423">
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Fig.
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<quantity id="4CAEE6896D0DFFBF253B22C9FCE352BC" box="[822,850,475,499]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="2" pageNumber="423" unit="in" value="21.0">21</quantity>
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</figureCitation>
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in
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<bibRefCitation id="EFC7369D6D0DFFBF257422C8FB0B52BC" author="Mockford" box="[889,1210,474,499]" pageId="2" pageNumber="423" refString="Mockford, E. L. & Sullivan, D. M. (1986) Systematics of the graphocaeciliine psocids with a proposed higher classification of the family Lachesillidae (Psocoptera). Transactions of the American Entomological Society, 112 (1), 1 - 80." type="journal article" year="1986">Mockford & Sullivan, 1986</bibRefCitation>
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), and in lacking the broad, rugose, clunial projections over the area of the paraprocts. Differing from
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<emphasis id="B922977E6D0DFFBF221022EDFBC95159" box="[1053,1144,511,534]" italics="true" pageId="2" pageNumber="423">G. v a r t yi</emphasis>
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<bibRefCitation id="EFC7369D6D0DFFBF228E22EDFA935158" author="Turner" box="[1155,1314,511,535]" pageId="2" pageNumber="423" refString="Turner, B. D. (1984) Two new Psocoptera from Colombia. Entomologist's Monthly Magazine, 120, 211 - 215." type="journal article" year="1984">Turner (1984)</bibRefCitation>
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, in lacking a crescent shaped, clear membranous area in the mid clunium and projections over the area of the paraprocts, and in the shape of the hypandrium and endophallic sclerites.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C34C18E76D0DFFBF26CA2178FE3255C1" pageId="2" pageNumber="423" type="description">
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||
<paragraph id="8BE94B6C6D0DFFBF26CA2178FA2B51E8" blockId="2.[151,1437,258,2033]" pageId="2" pageNumber="423">
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<emphasis id="B922977E6D0DFFBF26CA2178FEEF51CC" bold="true" box="[199,350,618,643]" pageId="2" pageNumber="423">Male. Color.</emphasis>
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Body tawny brown, compound eyes black, legs yellow, forewings pale yellowish; hindwings almost transparent. Tergal lobes of meso- and metathorax pale brown, abdomen creamy. Clunial projection brown.
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</paragraph>
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<paragraph id="8BE94B6C6D0DFFBF26CA21A0FD645742" blockId="2.[151,1437,258,2033]" pageId="2" pageNumber="423">
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<emphasis id="B922977E6D0DFFBF26CA21A0FED05184" bold="true" box="[199,353,690,715]" pageId="2" pageNumber="423">Morphology.</emphasis>
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As in diagnosis, plus the following: head vertex almost straight; upper border of compound eyes below the level of the vertex, with interommatidial setae (
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<figureCitation id="136D57E96D0DFFBF253621C4FC3351A0" box="[827,898,726,751]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1781,1804]" captionTargetBox="[232,1356,214,1729]" captionTargetId="figure@3.[227,1359,188,1729]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264989/files/figure.png" pageId="2" pageNumber="423">Fig. 2</figureCitation>
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). Forewing L/W: 2.49, with setae in the veins except for Cu2; lp/wp: 3.05, pterostigma without setae in the membrane, areola postica triangular (
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<figureCitation id="136D57E96D0DFFBF234F21EBFA3A505D" box="[1346,1419,761,786]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1781,1804]" captionTargetBox="[232,1356,214,1729]" captionTargetId="figure@3.[227,1359,188,1729]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264989/files/figure.png" pageId="2" pageNumber="423">Fig. 1</figureCitation>
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). Hindwing (
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<figureCitation id="136D57E96D0DFFBF2716200DFED05077" box="[283,353,799,824]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1781,1804]" captionTargetBox="[232,1356,214,1729]" captionTargetId="figure@3.[227,1359,188,1729]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264989/files/figure.png" pageId="2" pageNumber="423">Fig. 1</figureCitation>
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), L/W: 2.67. Hypandrium (
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<figureCitation id="136D57E96D0DFFBF24A7200DFD435077" box="[682,754,799,824]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1781,1804]" captionTargetBox="[232,1356,214,1729]" captionTargetId="figure@3.[227,1359,188,1729]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264989/files/figure.png" pageId="2" pageNumber="423">Fig. 5</figureCitation>
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), distinctly projected posteriorly. Arms of the phallosome aedeagal arch proximally wide; endophallus with minute denticles and two groups of sclerites: the anterior one with two denticles, and the posterior with seven, disposed as illustrated (
|
||
<figureCitation id="136D57E96D0DFFBF25C52074FBBF5030" box="[968,1038,870,895]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1781,1804]" captionTargetBox="[232,1356,214,1729]" captionTargetId="figure@3.[227,1359,188,1729]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264989/files/figure.png" pageId="2" pageNumber="423">Fig. 4</figureCitation>
|
||
). External parameres bearing pores distally (
|
||
<figureCitation id="136D57E96D0DFFBF26F42098FE8E50ED" box="[249,319,906,931]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1781,1804]" captionTargetBox="[232,1356,214,1729]" captionTargetId="figure@3.[227,1359,188,1729]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264989/files/figure.png" pageId="2" pageNumber="423">Fig. 4</figureCitation>
|
||
). Posterior border of clunium projected over the area of the epiproct, covering about one fifth of its length (
|
||
<figureCitation id="136D57E96D0DFFBF26E020BFFE835089" box="[237,306,941,966]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1781,1804]" captionTargetBox="[232,1356,214,1729]" captionTargetId="figure@3.[227,1359,188,1729]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264989/files/figure.png" pageId="2" pageNumber="423">Fig. 3</figureCitation>
|
||
). Clunium not projected over the area of the paraprocts (
|
||
<figureCitation id="136D57E96D0DFFBF25B920BFFC495089" box="[948,1016,941,966]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1781,1804]" captionTargetBox="[232,1356,214,1729]" captionTargetId="figure@3.[227,1359,188,1729]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264989/files/figure.png" pageId="2" pageNumber="423">Fig. 3</figureCitation>
|
||
), these broad, elongate, with setae as illustrated; sensory fields with 16–19 trichobothria in basal rosettes. Epiproct (
|
||
<figureCitation id="136D57E96D0DFFBF226D20C3FB1A50A6" box="[1120,1195,977,1002]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1781,1804]" captionTargetBox="[232,1356,214,1729]" captionTargetId="figure@3.[227,1359,188,1729]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Graphocaecilius mockfordi (Male). 1. Fore- and hind wings. 2. Front view of head. 3. Clunium, paraprocts and epiproct. 4. Phallosome. 5. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264989/files/figure.png" pageId="2" pageNumber="423">Fig. 3</figureCitation>
|
||
) anteriorly straight, posteriorly rounded, with macrosetae as illustrated.
|
||
</paragraph>
|
||
<paragraph id="8BE94B6C6D0DFFBF26CA270AFE325738" blockId="2.[151,1437,258,2033]" pageId="2" pageNumber="423">
|
||
<emphasis id="B922977E6D0DFFBF26CA270AFEC7577E" bold="true" box="[199,374,1048,1073]" pageId="2" pageNumber="423">Measurements</emphasis>
|
||
(in microns). FW: 2425, HW: 1800, F: 525, T: 900, t1: 300, t2: 100, ctt1: 22,
|
||
<collectingCountry id="F3410BFC6D0DFFBF22F3270AFA99577F" box="[1278,1320,1048,1072]" name="Mexico" pageId="2" pageNumber="423">Mx</collectingCountry>
|
||
4: 150, f1: 490, f2: 330, f3: 280, f4: 240, f5: 160, f6: 160, f7: 150, f8: 130, f9: 110, f10: 100, f11: 110, IO: 460, D: 180, d: 120, IO/d: 3.83, PO: 0.66.
|
||
</paragraph>
|
||
<paragraph id="8BE94B6C6D0DFFBF26CA2790FD0B57AD" blockId="2.[151,1437,258,2033]" pageId="2" pageNumber="423">
|
||
<emphasis id="B922977E6D0DFFBF26CA2790FEC457D4" bold="true" box="[199,373,1154,1179]" pageId="2" pageNumber="423">Female. Color.</emphasis>
|
||
Same as the male (
|
||
<figureCitation id="136D57E96D0DFFBF245D2790FD1557D4" box="[592,676,1154,1179]" captionStart="FIGURES 26 – 28. 26" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[152,1433,193,1832]" captionTargetId="figure@10.[152,1435,193,1834]" captionTargetPageId="10" captionText="FIGURES 26 – 28. 26. Graphocaecilius mockfordi (Female). 27. Graphocaecilius sullivani (Male). 28. Graphocaecilius sullivani (Female). Scales in mm." httpUri="https://zenodo.org/record/264994/files/figure.png" pageId="2" pageNumber="423">Fig. 26</figureCitation>
|
||
); abdomen creamy, with transverse, subcuticular pale brown bands. Clunium brown; epiproct pale brown, paraprocts creamy with pale brown spots. Subgenital plate pale brown with a median apical creamy longitudinal band (
|
||
<figureCitation id="136D57E96D0DFFBF246527DBFD1C57AD" box="[616,685,1225,1250]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1725,1748]" captionTargetBox="[241,1351,196,1692]" captionTargetId="figure@4.[239,1353,193,1692]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE94B6C6D0DFFBF26CA27FFFF40556B" blockId="2.[151,1437,258,2033]" pageId="2" pageNumber="423">
|
||
<emphasis id="B922977E6D0DFFBF26CA27FFFED05649" bold="true" box="[199,353,1261,1286]" pageId="2" pageNumber="423">Morphology.</emphasis>
|
||
Head as in the male, compound eyes smaller than in male, below the level of the vertex (
|
||
<figureCitation id="136D57E96D0DFFBF234727FFFA3F564A" box="[1354,1422,1261,1286]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1725,1748]" captionTargetBox="[241,1351,196,1692]" captionTargetId="figure@4.[239,1353,193,1692]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Fig. 7</figureCitation>
|
||
). Forewing L/W: 2.59. Pterostigma, lp/wp: 3.33. Areola postica triangular, not slanted posteriorly (
|
||
<figureCitation id="136D57E96D0DFFBF22DE2602FAA65666" box="[1235,1303,1296,1321]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1725,1748]" captionTargetBox="[241,1351,196,1692]" captionTargetId="figure@4.[239,1353,193,1692]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Fig. 6</figureCitation>
|
||
). Hindwing L/W: 2.74 (
|
||
<figureCitation id="136D57E96D0DFFBF27142626FEEF5602" box="[281,350,1332,1357]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1725,1748]" captionTargetBox="[241,1351,196,1692]" captionTargetId="figure@4.[239,1353,193,1692]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Fig. 6</figureCitation>
|
||
). Subgenital plate (
|
||
<figureCitation id="136D57E96D0DFFBF24342626FDCC5603" box="[569,637,1332,1357]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1725,1748]" captionTargetBox="[241,1351,196,1692]" captionTargetId="figure@4.[239,1353,193,1692]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Fig. 9</figureCitation>
|
||
), densely setose, with sides converging to a straight posterior border; a longitudinal slender band next to posterior border; IX sternum (
|
||
<figureCitation id="136D57E96D0DFFBF25522645FC015620" box="[863,944,1367,1392]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1725,1748]" captionTargetBox="[241,1351,196,1692]" captionTargetId="figure@4.[239,1353,193,1692]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Fig. 10</figureCitation>
|
||
) well developed, slightly pigmented, with a rectangular anterior projection. Gonapophyses (
|
||
<figureCitation id="136D57E96D0DFFBF24B32668FCA556DC" box="[702,788,1402,1427]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1725,1748]" captionTargetBox="[241,1351,196,1692]" captionTargetId="figure@4.[239,1353,193,1692]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Fig. 10</figureCitation>
|
||
), with
|
||
<date id="FFE86DAC6D0DFFBF25672669FC3156D8" box="[874,896,1403,1431]" pageId="2" pageNumber="423">v1</date>
|
||
elongate, slender, joined to clunium;
|
||
<date id="FFE86DAC6D0DFFBF23332669FAE556D8" box="[1342,1364,1403,1431]" pageId="2" pageNumber="423">v2</date>
|
||
+3 with anterior half hyaline, posterior half pigmented, slightly projected distally, bearing a row of setae along posterior border. Paraprocts (
|
||
<figureCitation id="136D57E96D0DFFBF277126D6FE725692" box="[380,451,1476,1501]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1725,1748]" captionTargetBox="[241,1351,196,1692]" captionTargetId="figure@4.[239,1353,193,1692]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Fig. 8</figureCitation>
|
||
) twice as long as wide, rhomboid, with setae as illustrated; sensory fields with 14 trichobothria issuing from basal rosettes (
|
||
<figureCitation id="136D57E96D0DFFBF246B26F5FD1A554F" box="[614,683,1511,1536]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1725,1748]" captionTargetBox="[241,1351,196,1692]" captionTargetId="figure@4.[239,1353,193,1692]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Fig. 8</figureCitation>
|
||
). Epiproct broadly triangular, anteriorly convex, setae as illustrated (
|
||
<figureCitation id="136D57E96D0DFFBF26922519FF55556B" box="[159,228,1547,1572]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1725,1748]" captionTargetBox="[241,1351,196,1692]" captionTargetId="figure@4.[239,1353,193,1692]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Graphocaecilius mockfordi (Female). 6. Fore- and hind wings. 7. Front view of head. 8. Clunium, paraprocts and epiproct. 9. Subgenital plate. 10. Gonapophyses and ninth sternum. Scales in mm." httpUri="https://zenodo.org/record/264990/files/figure.png" pageId="2" pageNumber="423">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE94B6C6D0DFFBF26CA253CFE3255C1" blockId="2.[151,1437,258,2033]" pageId="2" pageNumber="423">
|
||
<emphasis id="B922977E6D0DFFBF26CA253CFEC75508" bold="true" box="[199,374,1582,1607]" pageId="2" pageNumber="423">Measurements</emphasis>
|
||
(in microns). FW: 2850, HW: 2195, F: 650, T: 1100, t1: 350, t2: 125, ctt1: 17,
|
||
<collectingCountry id="F3410BFC6D0DFFBF22F2253DFA9B5508" box="[1279,1322,1583,1607]" name="Mexico" pageId="2" pageNumber="423">Mx</collectingCountry>
|
||
4: 160, f1: 580, f2: 360, f3: 300, f4: 260, f5: 170, f6: 160, f7: 135, f8: 120, f9: 105, f10: 100, f11: 110, IO: 560, D: 180, d: 100, IO/d: 5.60, PO: 0.55.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C18E76D0DFFBF26CA258BFCA554BE" pageId="2" pageNumber="423" type="etymology">
|
||
<paragraph id="8BE94B6C6D0DFFBF26CA258BFEA25453" blockId="2.[151,1437,258,2033]" pageId="2" pageNumber="423">
|
||
<emphasis id="B922977E6D0DFFBF26CA258BFEFF55FD" bold="true" box="[199,334,1689,1714]" pageId="2" pageNumber="423">Etymology.</emphasis>
|
||
This species is dedicated to Dr. Edward L. Mockford, of Illinois State University, in recognition to his many and fundamental studies on the taxonomy of ‘Psocoptera’, including his foundational work on the taxonomy of the Graphocaeciliine psocids, in collaboration with Mr. D. M. Sullivan, formerly of Illinois State University.
|
||
</paragraph>
|
||
<paragraph id="8BE94B6C6D0DFFBF26CA2435FCA554BE" blockId="2.[151,1437,258,2033]" pageId="2" pageNumber="423">
|
||
<materialsCitation id="3B3E41316D0DFFBF26CA2435FA295470" ID-GBIF-Occurrence="1290996067" box="[199,1432,1831,1856]" collectorName="La Cumbre" country="Colombia" county="Specimens" latitude="3.6364164" location="Montanitas" municipality="La Cumbre" pageId="2" pageNumber="423" specimenCount="1" stateProvince="Valle del Cauca" typeStatus="holotype">
|
||
<emphasis id="B922977E6D0DFFBF26CA2435FE17540F" bold="true" box="[199,422,1831,1856]" pageId="2" pageNumber="423">
|
||
<collectingCounty id="628833E06D0DFFBF26CA2435FEF4540F" box="[199,325,1831,1856]" pageId="2" pageNumber="423">Specimens</collectingCounty>
|
||
studied
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="54EDF5CE6D0DFFBF27BA2435FDAE540F" box="[439,543,1831,1856]" pageId="2" pageNumber="423" type="holotype">Holotype</typeStatus>
|
||
male.
|
||
<collectingCountry id="F3410BFC6D0DFFBF247C2435FCB85470" box="[625,777,1831,1855]" name="Colombia" pageId="2" pageNumber="423">COLOMBIA</collectingCountry>
|
||
,
|
||
<collectingRegion id="4992858E6D0DFFBF25152435FC605470" box="[792,977,1831,1856]" country="Colombia" name="Valle del Cauca" pageId="2" pageNumber="423">Valle del Cauca</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B8DD1166D0DFFBF25D2243AFBD5540F" box="[991,1124,1831,1856]" pageId="2" pageNumber="423">
|
||
<collectorName id="26A32EBA6D0DFFBF25D2243AFBD5540F" box="[991,1124,1831,1856]" pageId="2" pageNumber="423">La Cumbre</collectorName>
|
||
</collectingMunicipality>
|
||
,
|
||
<location id="8E891DB76D0DFFBF227F2435FB42540F" LSID="urn:lsid:plazi:treatment:03FFFA7A6D0DFFBF269A2385FCA554BE:8E891DB76D0DFFBF227F2435FB42540F" box="[1138,1267,1831,1856]" country="Colombia" county="Specimens" latitude="3.6364164" municipality="La Cumbre" name="Montanitas" pageId="2" pageNumber="423" stateProvince="Valle del Cauca">Montañitas</location>
|
||
,
|
||
<geoCoordinate id="EE622DAB6D0DFFBF230C2435FA295470" box="[1281,1432,1831,1855]" direction="north" orientation="latitude" pageId="2" pageNumber="423" precision="1" value="3.6364164">3°38’11.1’’N</geoCoordinate>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="3B3E41316D0DFFBF269A2458FDC554C8" ID-GBIF-Occurrence="1290996062" collectingDate="2012-04-26" collectorName="La Cumbre" country="Colombia" county="Specimens" elevation="1665" latitude="3.6364164" location="Montanitas" longitude="-76.53148" municipality="La Cumbre" pageId="2" pageNumber="423" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="paratype">
|
||
<geoCoordinate id="EE622DAB6D0DFFBF269A2458FE8E542C" box="[151,319,1866,1891]" direction="west" orientation="longitude" pageId="2" pageNumber="423" precision="1" value="-76.53148">76°31’53.3’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4CAEE6896D0DFFBF27412458FE15542D" box="[332,420,1866,1891]" metricMagnitude="3" metricUnit="m" metricValue="1.6649999999999998" pageId="2" pageNumber="423" unit="m" value="1665.0">
|
||
<elevation id="007BAC5F6D0DFFBF27412458FE15542D" box="[332,420,1866,1891]" metricMagnitude="3" metricUnit="m" metricValue="1.6649999999999998" pageId="2" pageNumber="423" unit="m" value="1665.0">1665 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFE86DAC6D0DFFBF27BD2458FD9E542C" box="[432,559,1866,1891]" pageId="2" pageNumber="423" value="2012-04-26">
|
||
<collectingDate id="EFAC94446D0DFFBF27BD2458FD9E542C" box="[432,559,1866,1891]" pageId="2" pageNumber="423" value="2012-04-26">26.IV.2012</collectingDate>
|
||
</date>
|
||
, MUSENUV slide code 26161, R. González.
|
||
<typeStatus id="54EDF5CE6D0DFFBF22382459FB16542C" box="[1077,1191,1867,1891]" pageId="2" pageNumber="423" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D5080E56D0DFFBF22BF2459FAB0542C" box="[1202,1281,1866,1891]" pageId="2" pageNumber="423" type="male">1 male</specimenCount>
|
||
, same data as holotype, MUSENUV slide code 26162
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3E41316D0DFFBF248B247DFE9E54E5" ID-GBIF-Occurrence="1290996066" collectorName="El Porvenir" country="Colombia" county="El Porvenir" latitude="3.4498334" location="Los Yarumos Nature Reserve" municipality="Felidia" pageId="2" pageNumber="423" specimenCount="2" specimenCount-female="2" stateProvince="Valle del Cauca" typeStatus="paratype">
|
||
<specimenCount id="9D5080E56D0DFFBF248B247DFD4A54C8" box="[646,763,1902,1927]" pageId="2" pageNumber="423" type="female">2 females</specimenCount>
|
||
,
|
||
<collectingCounty id="628833E06D0DFFBF2500247CFC2754C8" box="[781,918,1902,1927]" pageId="2" pageNumber="423">
|
||
<collectorName id="26A32EBA6D0DFFBF2500247CFC2754C8" box="[781,918,1902,1927]" pageId="2" pageNumber="423">El Porvenir</collectorName>
|
||
</collectingCounty>
|
||
, ca.
|
||
<collectingMunicipality id="6B8DD1166D0DFFBF25D8247CFB9754C8" box="[981,1062,1902,1927]" pageId="2" pageNumber="423">Felidia</collectingMunicipality>
|
||
,
|
||
<location id="8E891DB76D0DFFBF2235247DFA2954C9" LSID="urn:lsid:plazi:treatment:03FFFA7A6D0DFFBF269A2385FCA554BE:8E891DB76D0DFFBF2235247DFA2954C9" box="[1080,1432,1903,1926]" country="Colombia" county="El Porvenir" latitude="3.4498334" municipality="Felidia" name="Los Yarumos Nature Reserve" pageId="2" pageNumber="423" stateProvince="Valle del Cauca">Los Yarumos Nature Reserve</location>
|
||
,
|
||
<geoCoordinate id="EE622DAB6D0DFFBF269A2483FE9E54E5" box="[151,303,1937,1962]" direction="north" orientation="latitude" pageId="2" pageNumber="423" precision="1" value="3.4498334">3°26’59.4’’N</geoCoordinate>
|
||
</materialsCitation>
|
||
: 76°39’49.5’’W,
|
||
<quantity id="4CAEE6896D0DFFBF27F12480FDEA54E6" box="[508,603,1938,1962]" metricMagnitude="3" metricUnit="m" metricValue="2.328" pageId="2" pageNumber="423" unit="m" value="2328.0">2328 m</quantity>
|
||
,
|
||
<date id="FFE86DAC6D0DFFBF24632480FD5D54E5" box="[622,748,1938,1962]" pageId="2" pageNumber="423" value="2012-04-27">27.IV.2012</date>
|
||
, MUSENUV slide code 26163, R. González.
|
||
<materialsCitation id="3B3E41316D0DFFBF232C2480FBD15481" ID-GBIF-Occurrence="1290996061" collectorName="Planes de San Rafael" country="Colombia" county="Santuario" latitude="5.1395555" location="Laguna Encantada" municipality="Planes de San Rafael" pageId="2" pageNumber="423" specimenCount="2" specimenCount-female="2" stateProvince="Risaralda" typeStatus="paratype">
|
||
<specimenCount id="9D5080E56D0DFFBF232C2480FA2954E5" box="[1313,1432,1937,1962]" pageId="2" pageNumber="423" type="female">2 females</specimenCount>
|
||
,
|
||
<collectingRegion id="4992858E6D0DFFBF269A24A7FEB55481" box="[151,260,1973,1998]" country="Colombia" name="Risaralda" pageId="2" pageNumber="423">Risaralda</collectingRegion>
|
||
,
|
||
<collectingCounty id="628833E06D0DFFBF271724A7FE395481" box="[282,392,1973,1998]" pageId="2" pageNumber="423">Santuario</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6B8DD1166D0DFFBF279324A7FD1E5481" box="[414,687,1973,1998]" pageId="2" pageNumber="423">
|
||
<collectorName id="26A32EBA6D0DFFBF279324A7FD1E5481" box="[414,687,1973,1998]" pageId="2" pageNumber="423">Planes de San Rafael</collectorName>
|
||
</collectingMunicipality>
|
||
,
|
||
<location id="8E891DB76D0DFFBF24C824A4FC155481" LSID="urn:lsid:plazi:treatment:03FFFA7A6D0DFFBF269A2385FCA554BE:8E891DB76D0DFFBF24C824A4FC155481" box="[709,932,1973,1998]" country="Colombia" county="Santuario" latitude="5.1395555" municipality="Planes de San Rafael" name="Laguna Encantada" pageId="2" pageNumber="423" stateProvince="Risaralda">Laguna Encantada</location>
|
||
,
|
||
<geoCoordinate id="EE622DAB6D0DFFBF25B724A7FBD15481" box="[954,1120,1973,1998]" direction="north" orientation="latitude" pageId="2" pageNumber="423" precision="1" value="5.1395555">05°08’22.4’’N</geoCoordinate>
|
||
</materialsCitation>
|
||
: 75°59’42.4’’W,
|
||
<quantity id="4CAEE6896D0DFFBF233824A7FA265482" box="[1333,1431,1973,1997]" metricMagnitude="3" metricUnit="m" metricValue="2.27" pageId="2" pageNumber="423" unit="m" value="2270.0">2270 m</quantity>
|
||
,
|
||
<date id="FFE86DAC6D0DFFBF269A24CBFEA854BE" box="[151,281,2009,2033]" pageId="2" pageNumber="423" value="2014-11-09">09.XI.2014</date>
|
||
, MUSENUV slide code 26164, R. González.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |