304 lines
33 KiB
XML
304 lines
33 KiB
XML
<document ID-DOI="10.1206/0003-0082(2003)421<0001:ANSOTR>2.0.CO;2" ID-GBIF-Dataset="f4c2f80e-4917-4b5b-9415-0ee6377973ce" ID-ISSN="0003-0082" ID-Zenodo-Dep="4734917" approvalRequired="4" approvalRequired_for_taxonomicNames="4" checkinTime="1620068002453" checkinUser="felipe" docAuthor="VOSS, ROBERT S." docDate="2003" docId="03FAB267FFAAFFD6FF33FB27D9D5DD20" docLanguage="en" docName="AmMusNovit.2003.3421.1-47.pdf" docOrigin="American Museum Novitates 3421" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0082%282003%29421%3C0001%3AANSOTR%3E2.0.CO%3B2" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.1:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="1" docTitle="Thomasomys cinnameus Anthony 1924" docType="treatment" docVersion="5" lastPageNumber="30" masterDocId="FFC3CA1FFFB6FFCBFF8AFF92DB29DF75" masterDocTitle="A New Species of Thomasomys (Rodentia: Muridae) from Eastern Ecuador, with Remarks on Mammalian Diversity and Biogeography in the Cordillera Oriental" masterLastPageNumber="48" masterPageNumber="1" pageNumber="29" updateTime="1671043263584" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A New Species of Thomasomys (Rodentia: Muridae) from Eastern Ecuador, with Remarks on Mammalian Diversity and Biogeography in the Cordillera Oriental</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>VOSS, ROBERT S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>American Museum Novitates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2003</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2003-12-09</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>3421</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>48</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://www.bioone.org/doi/abs/10.1206/0003-0082%282003%29421%3C0001%3AANSOTR%3E2.0.CO%3B2</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1206/0003-0082(2003)421<0001:ANSOTR>2.0.CO;2</mods:identifier>
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<mods:identifier type="GBIF-Dataset">f4c2f80e-4917-4b5b-9415-0ee6377973ce</mods:identifier>
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<mods:identifier type="ISSN">0003-0082</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4734917</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5607554" ID-GBIF-Taxon="189547101" ID-Zenodo-Dep="5607554" LSID="urn:lsid:plazi:treatment:03FAB267FFAAFFD6FF33FB27D9D5DD20" httpUri="http://treatment.plazi.org/id/03FAB267FFAAFFD6FF33FB27D9D5DD20" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
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<subSubSection box="[185,582,1205,1228]" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph blockId="28.[185,582,1205,1228]" box="[185,582,1205,1228]" pageId="28" pageNumber="29">
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<taxonomicName authority="Anthony" authorityName="Anthony" authorityYear="1924" box="[185,582,1205,1228]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="cinnameus">
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<emphasis box="[185,470,1205,1228]" italics="true" pageId="28" pageNumber="29">Thomasomys cinnameus</emphasis>
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Anthony
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
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<paragraph blockId="28.[321,456,1238,1257]" box="[321,456,1238,1257]" pageId="28" pageNumber="29">
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<heading box="[321,456,1238,1257]" centered="true" fontSize="36" level="3" pageId="28" pageNumber="29" reason="7">
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<figureCitation box="[321,456,1238,1257]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="24.[150,188,1394,1415]" captionStartId-1="25.[149,187,1392,1413]" captionStartId-2="26.[150,188,1588,1609]" captionTargetBox-0="[155,1216,195,1359]" captionTargetBox-1="[135,1215,203,1362]" captionTargetBox-2="[128,1226,194,1565]" captionTargetId-0="figure-12@24.[130,1225,188,1374]" captionTargetId-1="figure-13@25.[128,1222,186,1371]" captionTargetId-2="figure-10@26.[125,1229,187,1568]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionText-0="Fig. 11. Dorsal cranial views of eight Thomasomys species that occur in the Cordillera Oriental of northern Ecuador. Top row (left to right): T. aureus (UMMZ 127114), T. baeops (UMMZ 155708), T. cinnameus (UMMZ 155671), T. erro (UMMZ 155713). Bottom row (left to right): T. paramorum (UMMZ 155737), T. rhoadsi (AMNH 66256), T. silvestris (USNM 513592), T. ucucha (UMMZ 155644)." captionText-1="Fig. 12. Ventral cranial views of eight Thomasomys species from the Cordillera Oriental of northern Ecuador. Top row (left to right): T. aureus (UMMZ 127114), T. baeops (UMMZ 155708), T. cinnameus (UMMZ 155671), T. erro (UMMZ 155713). Bottom row (left to right): T. paramorum (UMMZ 155737), T. rhoadsi (AMNH 66256), T. silvestris (USNM 513592), T. ucucha (UMMZ 155644)." captionText-2="Fig. 13. Lateral cranial and mandibular views of eight Thomasomys species from the Cordillera Oriental of northern Ecuador. Left side (top to bottom): T. aureus (UMMZ 127114), T. baeops (UMMZ 155708), T. cinnameus (UMMZ 155671), T. erro (UMMZ 155713). Right column (top to bottom): T. paramorum (UMMZ 155737), T. rhoadsi (AMNH 66256), T. silvestris (USNM 513592), T. ucucha (UMMZ 155644)." figureDoi-0="http://doi.org/10.5281/zenodo.4734946" figureDoi-1="http://doi.org/10.5281/zenodo.4734948" figureDoi-2="http://doi.org/10.5281/zenodo.4734950" httpUri-0="https://zenodo.org/record/4734946/files/figure.png" httpUri-1="https://zenodo.org/record/4734948/files/figure.png" httpUri-2="https://zenodo.org/record/4734950/files/figure.png" pageId="28" pageNumber="29">Figures 11–13</figureCitation>
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</heading>
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</paragraph>
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<paragraph blockId="28.[124,654,1278,1741]" pageId="28" pageNumber="29">
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SPECIMENS COLLECTED: 1.6 km (by road) W Papallacta,
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<quantity box="[308,421,1308,1331]" metricMagnitude="0" metricUnit="m" metricValue="3.2003999999999997" pageId="28" pageNumber="29" unit="ft" value="10.5">10,500 ft</quantity>
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(UMMZ 155668– 155670); Río Papallacta valley [
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<quantity box="[517,611,1337,1360]" metricMagnitude="3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="28" pageNumber="29" unit="km" value="4.0" valueMax="5.0" valueMin="3.0">3–5 km</quantity>
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by trail NNW Papallacta],
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<quantity box="[425,539,1366,1389]" metricMagnitude="0" metricUnit="m" metricValue="3.38328" pageId="28" pageNumber="29" unit="ft" value="11.1">11,100 ft</quantity>
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(AMNH 155671, 155800).
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</paragraph>
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<paragraph blockId="28.[124,654,1278,1741]" box="[153,445,1425,1448]" pageId="28" pageNumber="29">OTHER MATERIAL: None.</paragraph>
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<paragraph blockId="28.[124,654,1278,1741]" lastBlockId="28.[700,1229,193,1536]" pageId="28" pageNumber="29">
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TAXONOMY: This material of
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<taxonomicName authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="cinnameus">
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<emphasis italics="true" pageId="28" pageNumber="29">Thomasomys cinnameus</emphasis>
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</taxonomicName>
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was the first to be reported (by
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<bibRefCitation author="Voss, R. S." box="[124,261,1513,1536]" pageId="28" pageNumber="29" pagination="259 - 493" refId="ref25650" refString="Voss, R. S. 1988. Systematics and ecology of ichthyomyine rodents (Muroidea): patterns of morphological evolution in a small adaptive radiation. Bulletin of the American Museum of Natural History 188: 259 - 493." type="journal article" year="1988">Voss, 1988</bibRefCitation>
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: table 43) since the original description based on a single specimen collected at
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<quantity box="[198,287,1572,1595]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="28" pageNumber="29" unit="ft" value="8000.0">8000 ft</quantity>
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(
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<quantity box="[307,403,1572,1595]" metricMagnitude="3" metricUnit="m" metricValue="2.438" pageId="28" pageNumber="29" unit="m" value="2438.0">2438 m</quantity>
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) near Hacienda San Francisco, Provincia Tunguragua,
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<collectingCountry box="[556,653,1601,1624]" name="Ecuador" pageId="28" pageNumber="29">Ecuador</collectingCountry>
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(
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<bibRefCitation author="Anthony, H. E." box="[132,328,1630,1653]" pageId="28" pageNumber="29" pagination="1 - 9" refId="ref21372" refString="Anthony, H. E. 1924 b. Preliminary report on Ecuadorean mammals. No. 6. American Museum Novitates 139: 1 - 9." type="journal article" year="1924">Anthony, 1924b</bibRefCitation>
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). Sidebyside comparisons indicate that the Papallacta series closely resembles the
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<typeStatus box="[275,377,1689,1712]" pageId="28" pageNumber="29">holotype</typeStatus>
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(AMNH 67401) in all qualitative and most quantitative characters. Although the
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<typeStatus box="[872,974,193,216]" pageId="28" pageNumber="29">holotype</typeStatus>
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slightly exceeds the Papallacta specimens in several external and craniodental measurements (
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<tableCitation box="[1036,1119,252,275]" captionStart="TABLE 5" captionStartId="29.[336,410,194,213]" captionTargetId="figure-23@29.[121,653,285,979]" captionTargetPageId="29" captionText="TABLE 5 Measurements (mm) and Weights (g) of Thomasomys cinnameus" pageId="28" pageNumber="29">table 5</tableCitation>
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), the differences are so small and the number of Papallacta specimens (five) is so few that these discrepancies do not seem taxonomically significant.
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</paragraph>
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<paragraph blockId="28.[700,1229,193,1536]" pageId="28" pageNumber="29">
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One of the smallest known species of the genus,
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<taxonomicName authorityName="Anthony" authorityYear="1924" box="[789,1078,428,451]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="cinnameus">
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<emphasis box="[789,1078,428,451]" italics="true" pageId="28" pageNumber="29">Thomasomys cinnameus</emphasis>
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</taxonomicName>
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is currently regarded as a subspecies or junior synonym of
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<taxonomicName authority="Thomas (1917)" authorityName="Thomas" authorityYear="1917" box="[735,1053,486,510]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[735,858,487,510]" italics="true" pageId="28" pageNumber="29">T. gracilis</emphasis>
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<bibRefCitation author="Thomas, O." box="[869,1053,486,509]" pageId="28" pageNumber="29" pagination="1 - 3" refId="ref25334" refString="Thomas, O. 1917. Preliminary diagnoses of new mammals obtained by the Yale - National Geographic Society Peruvian Expedition. Smithsonian Miscellaneous Collections 68 (4): 1 - 3." type="journal article" year="1917">Thomas (1917)</bibRefCitation>
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</taxonomicName>
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, originally described from a specimen collected at Machu Picchu in southern
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<collectingCountry box="[961,1014,545,568]" name="Peru" pageId="28" pageNumber="29">Peru</collectingCountry>
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(
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<bibRefCitation author="Cabrera, A." box="[1042,1223,545,568]" pageId="28" pageNumber="29" pagination="309 - 732" refId="ref21669" refString="Cabrera, A. 1961. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' (Ciencias Zoologicas) 4 (2): 309 - 732." type="journal article" year="1961">Cabrera, 1961</bibRefCitation>
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;
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<bibRefCitation author="Musser, G. G. & M. D. Carleton" box="[700,1040,574,597]" pageId="28" pageNumber="29" pagination="501 - 755" refId="ref23955" refString="Musser, G. G., and M. D. Carleton. 1993. Family Muridae. In D. E. Wilson and D. M. Reeder (editors), Mammal species of the world, 2 nd ed.: 501 - 755. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1993">Musser and Carleton, 1993</bibRefCitation>
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). However, the hypothesis that
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<taxonomicName authorityName="Anthony" authorityYear="1924" box="[886,1010,604,627]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="cinnameus">
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<emphasis box="[886,1010,604,627]" italics="true" pageId="28" pageNumber="29">cinnameus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Thomas" authorityYear="1917" box="[1070,1159,604,627]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[1070,1159,604,627]" italics="true" pageId="28" pageNumber="29">gracilis</emphasis>
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</taxonomicName>
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(with type localities separated by
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<quantity box="[1025,1128,633,656]" metricMagnitude="6" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="29" unit="km" value="1500.0">1500 km</quantity>
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of highly dissected mountainous terrain) are conspecific is unsupported by any published analysis or discussion of character data. Rather, my examination of both
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<typeStatus box="[995,1108,750,773]" pageId="28" pageNumber="29">holotypes</typeStatus>
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and other representative material
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<superScript attach="left" box="[973,982,779,793]" fontSize="6" pageId="28" pageNumber="29">6</superScript>
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indicates that these are unambiguously diagnosable taxa that should be recognized as distinct species.
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</paragraph>
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<paragraph blockId="28.[700,1229,193,1536]" pageId="28" pageNumber="29">
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Although similar to
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<taxonomicName authorityName="Thomas" authorityYear="1917" box="[977,1229,868,891]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[977,1229,868,891]" italics="true" pageId="28" pageNumber="29">Thomasomys gracilis</emphasis>
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</taxonomicName>
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in size, external proportions, and pelage coloration,
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<taxonomicName authorityName="Anthony" authorityYear="1924" box="[801,960,927,950]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="cinnameus">
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<emphasis box="[801,960,927,950]" italics="true" pageId="28" pageNumber="29">T. cinnameus</emphasis>
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</taxonomicName>
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consistently lacks genal vibrissae, long black tactile hairs of the upper cheek that are consistently present in
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<taxonomicName authorityName="Thomas" authorityYear="1917" box="[700,789,1015,1038]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[700,789,1015,1038]" italics="true" pageId="28" pageNumber="29">gracilis</emphasis>
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</taxonomicName>
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. Among other trenchant craniodental comparisons,
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<taxonomicName authorityName="Anthony" authorityYear="1924" box="[866,990,1044,1067]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="cinnameus">
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<emphasis box="[866,990,1044,1067]" italics="true" pageId="28" pageNumber="29">cinnameus</emphasis>
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</taxonomicName>
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is distinguished by (1) incisive foramina that are widest posteriorly, behind the maxillary/premaxillary suture; (2) absence of sphenopalatine vacuities; (3) smaller and less inflated auditory bullae; and (4) larger and more hypsodont molars with weakly developed cingula and stylar cusps. By contrast,
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<taxonomicName authorityName="Thomas" authorityYear="1917" box="[944,1033,1249,1272]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[944,1033,1249,1272]" italics="true" pageId="28" pageNumber="29">gracilis</emphasis>
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</taxonomicName>
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exhibits (1) incisive foramina that are widest anteriorly, at or near the maxillary/premaxillary suture; (2) large sphenopalatine vacuities that perforate the bony roof of the mesopterygoid fossa on each side of the basisphenoid/presphenoid suture; (3) larger and more inflated auditory bullae; and (4) smaller, brachydont molars with better developed cingula and stylar cusps.
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</paragraph>
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<paragraph blockId="28.[700,1229,1577,1741]" pageId="28" pageNumber="29">
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<materialsCitation ID-GBIF-Occurrence="3395013305" collectionCode="USNM, AMNH" elevation="4267" location="Peruvian" pageId="28" pageNumber="29" specimenCode="USNM 194816, USNM 194799, 194801, AMNH 95206, 95207, USNM 194785- 194787, 194790, 194807, 194808, 194811-194813" specimenCount="2" stateProvince="Cusco" typeStatus="holotype">
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<superScript attach="right" box="[723,730,1577,1588]" fontSize="5" pageId="28" pageNumber="29">6</superScript>
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The comparative material of
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<taxonomicName authorityName="Thomas" authorityYear="1917" box="[1026,1229,1578,1597]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[1026,1229,1578,1597]" italics="true" pageId="28" pageNumber="29">Thomasomys gracilis</emphasis>
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</taxonomicName>
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that I examined included the
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<typeStatus box="[976,1057,1602,1621]" pageId="28" pageNumber="29">holotype</typeStatus>
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(
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<specimenCode box="[1071,1218,1602,1621]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">USNM 194816</specimenCode>
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), two topotypes (
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<specimenCode box="[850,1000,1626,1645]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">USNM 194799</specimenCode>
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,
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<specimenCode box="[1013,1086,1626,1645]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">194801</specimenCode>
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), and 12
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<typeStatus isOtherTypeStatus="true" pageId="28" pageNumber="29">other specimens</typeStatus>
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(
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<specimenCode box="[819,966,1650,1669]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="28" pageNumber="29">AMNH 95206</specimenCode>
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,
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<specimenCode box="[985,1046,1650,1669]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="28" pageNumber="29">95207</specimenCode>
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;
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<specimenCode collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">USNM 194785– 194787</specimenCode>
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,
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<specimenCode box="[784,856,1674,1693]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">194790</specimenCode>
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,
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||
<specimenCode box="[867,939,1674,1693]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">194807</specimenCode>
|
||
,
|
||
<specimenCode box="[950,1022,1674,1693]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">194808</specimenCode>
|
||
,
|
||
<specimenCode box="[1033,1187,1674,1693]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">194811–194813</specimenCode>
|
||
), all of which were collected between 2774 and
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="4.267" pageId="28" pageNumber="29" unit="m" value="4267.0">
|
||
<quantity box="[1151,1228,1698,1717]" metricMagnitude="3" metricUnit="m" metricValue="4.267" pageId="28" pageNumber="29" unit="m" value="4267.0">4267 m</quantity>
|
||
above sea level
|
||
</elevation>
|
||
in the
|
||
<location LSID="urn:lsid:plazi:treatment:03FAB267FFAAFFD6FF33FB27D9D5DD20:8E8C55AAFFAAFFD7FC1EF928D8CFD9B8" box="[916,998,1722,1741]" name="Peruvian" pageId="28" pageNumber="29" stateProvince="Cusco">Peruvian</location>
|
||
department of
|
||
<collectingRegion box="[1146,1207,1722,1741]" country="Peru" name="Cusco" pageId="28" pageNumber="29">Cusco</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption pageId="29" pageNumber="30" startId="29.[336,410,194,213]" targetBox="[121,653,286,979]" targetPageId="29">
|
||
<paragraph blockId="29.[159,615,194,269]" pageId="29" pageNumber="30">
|
||
TABLE 5
|
||
<emphasis bold="true" pageId="29" pageNumber="30">
|
||
Measurements (mm) and Weights (g) of
|
||
<taxonomicName authorityName="Anthony" authorityYear="1924" box="[253,522,248,269]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cinnameus">
|
||
<emphasis bold="true" box="[253,522,248,269]" italics="true" pageId="29" pageNumber="30">Thomasomys cinnameus</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="29.[123,652,1044,1742]" lastBlockId="29.[699,1228,193,597]" pageId="29" pageNumber="30">
|
||
<taxonomicName authority="Anthony (1923)" authorityName="Anthony" authorityYear="1923" box="[152,618,1044,1067]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hudsoni">
|
||
<emphasis box="[152,410,1044,1067]" italics="true" pageId="29" pageNumber="30">Thomasomys hudsoni</emphasis>
|
||
<bibRefCitation author="Anthony, H. E." box="[424,618,1044,1067]" pageId="29" pageNumber="30" pagination="1 - 14" refId="ref21317" refString="Anthony, H. E. 1923. Preliminary report on Ecuadorean mammals. No. 3. American Museum Novitates 55: 1 - 14." type="journal article" year="1923">Anthony (1923)</bibRefCitation>
|
||
</taxonomicName>
|
||
is another small Ecuadorean taxon that has been treated without explanation as a synonym or subspecies of
|
||
<taxonomicName authorityName="Thomas" authorityYear="1917" box="[406,534,1132,1155]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[406,534,1132,1155]" italics="true" pageId="29" pageNumber="30">T. gracilis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation author="Cabrera, A." pageId="29" pageNumber="30" pagination="309 - 732" refId="ref21669" refString="Cabrera, A. 1961. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' (Ciencias Zoologicas) 4 (2): 309 - 732." type="journal article" year="1961">Cabrera, 1961</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Musser, G. G. & M. D. Carleton" box="[290,638,1161,1184]" pageId="29" pageNumber="30" pagination="501 - 755" refId="ref23955" refString="Musser, G. G., and M. D. Carleton. 1993. Family Muridae. In D. E. Wilson and D. M. Reeder (editors), Mammal species of the world, 2 nd ed.: 501 - 755. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1993">Musser and Carleton, 1993</bibRefCitation>
|
||
). However, I agree with
|
||
<bibRefCitation author="Anthony, H. E." box="[396,598,1190,1213]" pageId="29" pageNumber="30" pagination="1 - 9" refId="ref21372" refString="Anthony, H. E. 1924 b. Preliminary report on Ecuadorean mammals. No. 6. American Museum Novitates 139: 1 - 9." type="journal article" year="1924">Anthony (1924b)</bibRefCitation>
|
||
that
|
||
<taxonomicName authorityName="Anthony" authorityYear="1923" box="[123,215,1220,1243]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hudsoni">
|
||
<emphasis box="[123,215,1220,1243]" italics="true" pageId="29" pageNumber="30">hudsoni</emphasis>
|
||
</taxonomicName>
|
||
is a distinct species, differing from both
|
||
<taxonomicName authorityName="Thomas" authorityYear="1917" box="[182,271,1249,1272]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[182,271,1249,1272]" italics="true" pageId="29" pageNumber="30">gracilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Anthony" authorityYear="1924" box="[329,453,1249,1272]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cinnameus">
|
||
<emphasis box="[329,453,1249,1272]" italics="true" pageId="29" pageNumber="30">cinnameus</emphasis>
|
||
</taxonomicName>
|
||
in details of coloration and craniodental morphology. Unlike any specimens of the other small species, the
|
||
<typeStatus box="[123,173,1337,1360]" pageId="29" pageNumber="30">type</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="Anthony" authorityYear="1923" box="[221,313,1337,1360]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hudsoni">
|
||
<emphasis box="[221,313,1337,1360]" italics="true" pageId="29" pageNumber="30">hudsoni</emphasis>
|
||
</taxonomicName>
|
||
(AMNH 47690, from Bestión in Provincia
|
||
<collectingRegion box="[336,414,1366,1389]" country="Ecuador" name="Azuay" pageId="29" pageNumber="30">Azuay</collectingRegion>
|
||
) has a rostrum that is peculiarly produced beyond the incisors as a flaring bony tube with a concave (rather than convex) dorsal profile. As in
|
||
<taxonomicName authorityName="Anthony" authorityYear="1924" box="[527,651,1455,1478]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cinnameus">
|
||
<emphasis box="[527,651,1455,1478]" italics="true" pageId="29" pageNumber="30">cinnameus</emphasis>
|
||
</taxonomicName>
|
||
(and unlike
|
||
<taxonomicName authorityName="Thomas" authorityYear="1917" box="[286,375,1484,1507]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[286,375,1484,1507]" italics="true" pageId="29" pageNumber="30">gracilis</emphasis>
|
||
</taxonomicName>
|
||
) genal vibrissae and sphenopalatine vacuities are absent in
|
||
<taxonomicName authorityName="Anthony" authorityYear="1923" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hudsoni">
|
||
<emphasis italics="true" pageId="29" pageNumber="30">hudsoni</emphasis>
|
||
</taxonomicName>
|
||
, but as in
|
||
<taxonomicName authorityName="Thomas" authorityYear="1917" box="[321,410,1543,1566]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[321,410,1543,1566]" italics="true" pageId="29" pageNumber="30">gracilis</emphasis>
|
||
</taxonomicName>
|
||
(and unlike
|
||
<taxonomicName authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cinnameus">
|
||
<emphasis italics="true" pageId="29" pageNumber="30">cinnameus</emphasis>
|
||
</taxonomicName>
|
||
) the incisive foramina are widest anteriorly (near the maxillary/premaxillary suture). The auditory bullae of
|
||
<taxonomicName authorityName="Anthony" authorityYear="1923" box="[507,599,1631,1654]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hudsoni">
|
||
<emphasis box="[507,599,1631,1654]" italics="true" pageId="29" pageNumber="30">hudsoni</emphasis>
|
||
</taxonomicName>
|
||
are larger and more inflated than those of
|
||
<taxonomicName authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cinnameus">
|
||
<emphasis italics="true" pageId="29" pageNumber="30">cinnameus</emphasis>
|
||
</taxonomicName>
|
||
but smaller and less inflated than those of
|
||
<taxonomicName authorityName="Thomas" authorityYear="1917" box="[234,323,1719,1742]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[234,323,1719,1742]" italics="true" pageId="29" pageNumber="30">gracilis</emphasis>
|
||
</taxonomicName>
|
||
. Unfortunately, the molars of the
|
||
<typeStatus box="[785,835,193,216]" pageId="29" pageNumber="30">type</typeStatus>
|
||
(and only known specimen) of
|
||
<taxonomicName authorityName="Anthony" authorityYear="1923" box="[699,791,223,246]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hudsoni">
|
||
<emphasis box="[699,791,223,246]" italics="true" pageId="29" pageNumber="30">hudsoni</emphasis>
|
||
</taxonomicName>
|
||
are too worn to support confident dental comparisons.
|
||
</paragraph>
|
||
<paragraph blockId="29.[699,1228,193,597]" pageId="29" pageNumber="30">
|
||
<materialsCitation ID-GBIF-Occurrence="3395013334" collectionCode="FIELD, OBSERVATIONS" county="Rio Papallacta" elevation="3290" location="Papallacta" municipality="Subalpine Rain Forest" pageId="29" pageNumber="30" specimenCount="1">
|
||
<collectionCode box="[728,973,281,304]" pageId="29" pageNumber="30">FIELD OBSERVATIONS</collectionCode>
|
||
: The
|
||
<specimenCount box="[1051,1227,281,304]" pageId="29" pageNumber="30" type="generic">five specimens</specimenCount>
|
||
of
|
||
<taxonomicName authorityName="Anthony" authorityYear="1924" box="[737,1027,311,334]" class="Mammalia" family="Muridae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cinnameus">
|
||
<emphasis box="[737,1027,311,334]" italics="true" pageId="29" pageNumber="30">Thomasomys cinnameus</emphasis>
|
||
</taxonomicName>
|
||
that I collected near
|
||
<location LSID="urn:lsid:plazi:treatment:03FAB267FFAAFFD6FF33FB27D9D5DD20:8E8C55AAFFABFFD6FD7DFEC6D847DE1E" box="[759,878,340,363]" county="Rio Papallacta" municipality="Subalpine Rain Forest" name="Papallacta" pageId="29" pageNumber="30">Papallacta</location>
|
||
in 1980 were trapped at elevations ranging from
|
||
<quantity box="[971,1178,369,392]" metricMagnitude="3" metricUnit="m" metricValue="3.29" metricValueMax="3.38" metricValueMin="3.2" pageId="29" pageNumber="30" unit="m" value="3290.0" valueMax="3380.0" valueMin="3200.0">
|
||
<elevation box="[971,1178,369,392]" metricMagnitude="3" metricUnit="m" metricValue="3.29" metricValueMax="3.38" metricValueMin="3.2" pageId="29" pageNumber="30" unit="m" value="3290.0" valueMax="3380.0" valueMin="3200.0">3200 to 3380 m</elevation>
|
||
</quantity>
|
||
.
|
||
<location LSID="urn:lsid:plazi:treatment:03FAB267FFAAFFD6FF33FB27D9D5DD20:8E8C55AAFFABFFD6FB26FEE3DFE2DEFD" box="[1196,1227,369,392]" county="Rio Papallacta" municipality="Subalpine Rain Forest" name="Of" pageId="29" pageNumber="30">Of</location>
|
||
these, three were taken among mossy boulders in an old lava flow that impounds the
|
||
<location LSID="urn:lsid:plazi:treatment:03FAB267FFAAFFD6FF33FB27D9D5DD20:8E8C55AAFFABFFD6FD31FE5BD868DE95" box="[699,833,457,480]" county="Rio Papallacta" municipality="Subalpine Rain Forest" name="Rio Tambo" pageId="29" pageNumber="30">Río Tambo</location>
|
||
to form
|
||
<location LSID="urn:lsid:plazi:treatment:03FAB267FFAAFFD6FF33FB27D9D5DD20:8E8C55AAFFABFFD6FC3FFE5BDFA6DE95" box="[949,1167,457,480]" county="Rio Papallacta" municipality="Subalpine Rain Forest" name="Laguna Papallacta" pageId="29" pageNumber="30">Laguna Papallacta</location>
|
||
(fig. 1), and two were trapped on the ground in
|
||
<collectingMunicipality box="[699,970,516,539]" pageId="29" pageNumber="30">Subalpine Rain Forest</collectingMunicipality>
|
||
in the valley of the
|
||
<collectingCounty box="[699,868,545,568]" pageId="29" pageNumber="30">Río Papallacta</collectingCounty>
|
||
<quantity box="[876,964,545,568]" metricMagnitude="3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="29" pageNumber="30" unit="km" value="4.0" valueMax="5.0" valueMin="3.0">3–5 km</quantity>
|
||
(by trail) NNW of the town
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |