199 lines
26 KiB
XML
199 lines
26 KiB
XML
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<mods:title id="C609924EC9BBE5518917735FEB06CC5A">A new choristodere (Reptilia: Diapsida) from the Lower Cretaceous of western Liaoning Province, China, and phylogenetic relationships of Monjurosuchidae</mods:title>
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<mods:namePart id="E8B3DEC9CDBE8FA13F97BAF64BCF320C">Gao, Ke-Qin</mods:namePart>
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<mods:namePart id="3456A1E040AB5E7A1AF2CDE28B93FD0A">Fox, Richard C.</mods:namePart>
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<mods:date id="CE92E6BDBA5C0246262BFE5E98CF7B50">2005</mods:date>
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<treatment id="03F787E20F1D820299F1F954BF6BFD49" ID-DOI="http://doi.org/10.5281/zenodo.5735913" ID-GBIF-Taxon="190873757" ID-Zenodo-Dep="5735913" LSID="urn:lsid:plazi:treatment:03F787E20F1D820299F1F954BF6BFD49" httpUri="http://treatment.plazi.org/id/03F787E20F1D820299F1F954BF6BFD49" lastPageNumber="439" pageId="12" pageNumber="439">
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<subSubSection id="C344657F0F1D820299F1F954BB35FE3A" box="[142,358,379,400]" pageId="12" pageNumber="439" type="nomenclature">
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<paragraph id="8BE136F40F1D820299F1F954BB35FE3A" blockId="12.[142,758,379,1904]" box="[142,358,379,400]" pageId="12" pageNumber="439">
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<emphasis id="B92AEAE60F1D820299F1F954BB35FE3A" box="[142,358,379,400]" italics="true" pageId="12" pageNumber="439">
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<collectingRegion id="499AF8160F1D820299F1F954BA9CFE3A" box="[142,207,379,400]" country="American Samoa" name="order" pageId="12" pageNumber="439">Order</collectingRegion>
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<taxonomicName id="4C5E4D770F1D820299A8F954BB35FE3A" authorityName="COPE" authorityYear="1876" box="[215,358,379,400]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="order">Choristodera</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C344657F0F1D820299F1F9B5BF6BFD49" pageId="12" pageNumber="439" type="discussion">
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<paragraph id="8BE136F40F1D820299F1F9B5BAB6FCD6" blockId="12.[142,758,379,1904]" pageId="12" pageNumber="439">
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For more than a century since
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<bibRefCitation id="EFCF4B050F1D82029897F9B5B82DFE1A" author="Cope ED" box="[488,638,410,432]" pageId="12" pageNumber="439" pagination="340 - 359" refId="ref10684" refString="Cope ED. 1876. [1877]. On some extinct reptiles and Batrachia from the Judith River and Fox Hills beds of Montana. Proceedings of the Academy Natural Sciences of Philadelphia 28: 340 - 359." type="journal article" year="1876">Cope’s (1876)</bibRefCitation>
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naming of the group, the order
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<taxonomicName id="4C5E4D770F1D820298EFF996B877FE65" authorityName="COPE" authorityYear="1876" box="[400,548,441,463]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="order">Choristodera</taxonomicName>
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has been known mostly from the
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<taxonomicName id="4C5E4D770F1D8202982AF9F8B870FE47" authorityName="Cope" authorityYear="1876" box="[341,547,471,493]" class="Reptilia" family="Champsosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="family">Champsosauridae</taxonomicName>
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and
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<taxonomicName id="4C5E4D770F1D82029B1CF9F8BA98FDA6" authorityName="Lemone" authorityYear="1884" class="Reptilia" family="Simoedosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="family">Simoedosauridae</taxonomicName>
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. The contents of the
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<taxonomicName id="4C5E4D770F1D820298CBF9D9B818FDA6" authorityName="COPE" authorityYear="1876" box="[436,587,502,524]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="order">Choristodera</taxonomicName>
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, however, have been significantly expanded recently by recognition of the membership of
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<taxonomicName id="4C5E4D770F1D82029816FA1BBBA2FDE3" baseAuthorityName="Evans" baseAuthorityYear="1990" box="[361,497,564,585]" class="Reptilia" family="Cteniogenyidae" genus="Cteniogenys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
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<emphasis id="B92AEAE60F1D82029816FA1BBBA2FDE3" box="[361,497,564,585]" italics="true" pageId="12" pageNumber="439">Cteniogenys</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C5E4D770F1D82029B56FA1CB88BFDE2" box="[553,728,563,584]" class="Reptilia" family="Simoedosauridae" genus="Monjurosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
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<emphasis id="B92AEAE60F1D82029B56FA1CB88BFDE2" box="[553,728,563,584]" italics="true" pageId="12" pageNumber="439">Monjurosuchus</emphasis>
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</taxonomicName>
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in this group (
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<bibRefCitation id="EFCF4B050F1D82029868FA7DBBFBFDCD" author="Evans SE" box="[279,424,594,615]" pageId="12" pageNumber="439" pagination="577 - 589" refId="ref10832" refString="Evans SE. 1989. New material of Cteniogenys (Reptilia: Diapsida) and a reassessment of the systematic position of the genus. Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte 1989: 577 - 589." type="journal article" year="1989">Evans, 1989</bibRefCitation>
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,
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<bibRefCitation id="EFCF4B050F1D820298C9FA7DBBBCFDCD" author="Evans SE" box="[438,495,594,615]" pageId="12" pageNumber="439" pagination="205 - 237" refId="ref10871" refString="Evans SE. 1990. The skull of Cteniogenys, a choristodere (Reptilia: Archosauromorpha) from the Middle Jurassic of Oxforshire. Zoological Journal of the Linnean Society 99: 205 - 237." type="journal article" year="1990">1990</bibRefCitation>
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;
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<bibRefCitation id="EFCF4B050F1D82029882FA7DB8E5FDC2" author="Gao K & Evans S & Ji Q & Norell M & Ji S" box="[509,694,594,616]" pageId="12" pageNumber="439" pagination="417 - 421" refId="ref11034" refString="Gao K, Evans S, Ji Q, Norell M, Ji S. 2000. Exceptional fossil material of a semi-aquatic reptile from China: the resolution of an enigma. Journal of Vertebrate Paleontology 20: 417 - 421." type="journal article" year="2000">
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Gao
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<emphasis id="B92AEAE60F1D82029B4BFA7CB837FDCD" box="[564,612,594,616]" italics="true" pageId="12" pageNumber="439">et al</emphasis>
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., 2000
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</bibRefCitation>
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), and by the discovery of new morphotypes including the long-necked
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<taxonomicName id="4C5E4D770F1D8202985CFAA0BB82FD0E" box="[291,465,655,676]" class="Reptilia" family="Hyphalosauridae" genus="Hyphalosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
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<emphasis id="B92AEAE60F1D8202985CFAA0BB82FD0E" box="[291,465,655,676]" italics="true" pageId="12" pageNumber="439">Hyphalosaurus</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="F34976640F1D82029B5DFAA0B834FD0F" box="[546,615,655,677]" name="China" pageId="12" pageNumber="439">China</collectingCountry>
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(
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<bibRefCitation id="EFCF4B050F1D82029B03FAA0BA95FD6E" author="Gao KQ & Tang Z & Wang X" pageId="12" pageNumber="439" pagination="1 - 8" refId="ref11123" refString="Gao KQ, Tang Z, Wang X. 1999. A long-necked diapsid reptile from the Upper Jurassic / Lower Cretaceous of Liaoning Province, northeastern China. Vertebrata Palasiatica 37: 1 - 8." type="journal article" year="1999">
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Gao
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<emphasis id="B92AEAE60F1D82029BC8FABFB8BBFD0E" box="[695,744,655,677]" italics="true" pageId="12" pageNumber="439">et al</emphasis>
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||
., 1999
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</bibRefCitation>
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) and
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<taxonomicName id="4C5E4D770F1D8202986FFA81BB2AFD69" box="[272,377,686,707]" class="Reptilia" genus="Shokawa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
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<emphasis id="B92AEAE60F1D8202986FFA81BB2AFD69" box="[272,377,686,707]" italics="true" pageId="12" pageNumber="439">Shokawa</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="F34976640F1D820298B9FA81B85FFD69" box="[454,524,686,707]" name="Japan" pageId="12" pageNumber="439">Japan</collectingCountry>
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(
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<bibRefCitation id="EFCF4B050F1D82029B5FFA81BA9BFD49" author="Evans SE & Manabe M" pageId="12" pageNumber="439" pagination="101 - 119" refId="ref10977" refString="Evans SE, Manabe M. 1999. A choristoderan reptile from the Lower Cretaceous of Japan. Special Papers in Palaeontology 60: 101 - 119." type="journal article" year="1999">Evans & Manabe, 1999</bibRefCitation>
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). By current understanding, the order
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<taxonomicName id="4C5E4D770F1D82029BDDFAE2BA84FCAB" authorityName="COPE" authorityYear="1876" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="order">Choristodera</taxonomicName>
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includes five families:
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<taxonomicName id="4C5E4D770F1D820298A3FAC4B8D2FCAB" authorityName="Seiffert" authorityYear="1975" box="[476,641,747,769]" class="Reptilia" family="Cteniogenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="family">Cteniogenidae</taxonomicName>
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,
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<taxonomicName id="4C5E4D770F1D82029BF2FAC4BAA7FC8A" class="Reptilia" family="Monjurosuchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="family">Monjurosuchidae</taxonomicName>
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,
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<taxonomicName id="4C5E4D770F1D82029870FB25BB86FC8A" authorityName="Gao & Fox" authorityYear="2005" box="[271,469,778,800]" class="Reptilia" family="Hyphalosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="family">Hyphalosauridae</taxonomicName>
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,
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<taxonomicName id="4C5E4D770F1D8202988FFB25B8E7FC8A" authorityName="Lemone" authorityYear="1884" box="[496,692,778,800]" class="Reptilia" family="Simoedosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="family">Simoedosauridae</taxonomicName>
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||
and
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<taxonomicName id="4C5E4D770F1D820299F1FB06BB0DFC95" authorityName="Cope" authorityYear="1876" box="[142,350,809,831]" class="Reptilia" family="Champsosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="family">Champsosauridae</taxonomicName>
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. The relationships of these families are depicted in cladograms resulting from this study (
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<figureCitation id="13652A710F1D820299E9FB49BA8BFCD6" box="[150,216,870,892]" captionStart="Figure 6" captionStartId="11.[162,240,1825,1844]" captionTargetBox="[170,768,810,1793]" captionTargetId="graphics-586@11.[201,760,976,1793]" captionTargetPageId="11" captionText="Figure 6. Comparison of the two equally parsimonious trees (TL = 123 steps; CI = 0.85; RI = 0.87) derived from analysis of data presented in Appendices 1 and 2." figureDoi="http://doi.org/10.5281/zenodo.4687375" httpUri="https://zenodo.org/record/4687375/files/figure.png" pageId="12" pageNumber="439">Fig. 6</figureCitation>
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||
).
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||
</paragraph>
|
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<paragraph id="8BE136F40F1D820299D8FBAAB843F95F" blockId="12.[142,758,379,1904]" pageId="12" pageNumber="439">
|
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The taxonomically problematic taxon
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||
<taxonomicName id="4C5E4D770F1D82029B09FBAABA91FC12" authorityName="Hecht" authorityYear="1992" class="Reptilia" genus="Lazarussuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAE60F1D82029B09FBAABA91FC12" italics="true" pageId="12" pageNumber="439">Lazarussuchus</emphasis>
|
||
</taxonomicName>
|
||
is excluded from the order
|
||
<taxonomicName id="4C5E4D770F1D82029B75FB8CB8CDFC13" authorityName="COPE" authorityYear="1876" box="[522,670,931,953]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="order">Choristodera</taxonomicName>
|
||
in this paper (see
|
||
<bibRefCitation id="EFCF4B050F1D82029872FBEDBB83FC7D" author="Gao KQ & Fox RC" box="[269,464,962,983]" pageId="12" pageNumber="439" pagination="303 - 353" refId="ref11077" refString="Gao KQ, Fox RC. 1998. New choristoderes (Reptilia: Diapsida) from the Upper Cretaceous and Palaeocene, Alberta and Saskatchewan, Canada, and phylogenetic relationships of Choristodera. Zoological Journal of the Linnean Society 124: 303 - 353." type="journal article" year="1998">Gao & Fox, 1998</bibRefCitation>
|
||
, for discussion). Independent phylogenetic analyses consistently place
|
||
<taxonomicName id="4C5E4D770F1D82029BBBFBCEBB5CFBBE" authorityName="Hecht" authorityYear="1992" class="Reptilia" genus="Lazarussuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAE60F1D82029BBBFBCEBB5CFBBE" italics="true" pageId="12" pageNumber="439">Lazarussuchus</emphasis>
|
||
</taxonomicName>
|
||
outside the choristoderan clade (
|
||
<bibRefCitation id="EFCF4B050F1D82029BD8FC2FBB56FB99" author="Gao KQ & Fox RC" pageId="12" pageNumber="439" pagination="303 - 353" refId="ref11077" refString="Gao KQ, Fox RC. 1998. New choristoderes (Reptilia: Diapsida) from the Upper Cretaceous and Palaeocene, Alberta and Saskatchewan, Canada, and phylogenetic relationships of Choristodera. Zoological Journal of the Linnean Society 124: 303 - 353." type="journal article" year="1998">Gao & Fox, 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCF4B050F1D82029866FC30B86DFB99" author="Evans SE & Manabe M" box="[281,574,1054,1076]" pageId="12" pageNumber="439" pagination="101 - 119" refId="ref10977" refString="Evans SE, Manabe M. 1999. A choristoderan reptile from the Lower Cretaceous of Japan. Special Papers in Palaeontology 60: 101 - 119." type="journal article" year="1999">Evans & Manabe, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCF4B050F1D82029B2DFC31BA94FBF9" author="Ksepka DT & Gao K-Q & Norell MA" pageId="12" pageNumber="439" pagination="1 - 22" refId="ref11393" refString="Ksepka DT, Gao K-Q, Norell MA. 2005. A new choristodere from the Cretaceous of Mongolia. American Museum Novitates 3468: 1 - 22." type="journal article" year="2005">
|
||
Ksepka
|
||
<emphasis id="B92AEAE60F1D82029BC8FC30B8BBFB99" box="[695,744,1054,1076]" italics="true" pageId="12" pageNumber="439">et al</emphasis>
|
||
., 2005
|
||
</bibRefCitation>
|
||
; this paper; but see also
|
||
<bibRefCitation id="EFCF4B050F1D82029B78FC12BA9BFBDB" author="Evans SE & Klembara J" pageId="12" pageNumber="439" pagination="171 - 184" refId="ref10943" refString="Evans SE, Klembara J. 2005. A choristoderan reptile (Reptilia: Diapsida) from the lower Miocene of northwest Bohemia. Journal of Vertebrate Palaeontology 25: 171 - 184." type="journal article" year="2005">Evans & Klembara, 2005</bibRefCitation>
|
||
, for different interpretations).
|
||
<taxonomicName id="4C5E4D770F1D82029B34FC74B8A5FBDA" authorityName="Hecht" authorityYear="1992" box="[587,758,1115,1136]" class="Reptilia" genus="Lazarussuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAE60F1D82029B34FC74B8A5FBDA" box="[587,758,1115,1136]" italics="true" pageId="12" pageNumber="439">Lazarussuchus</emphasis>
|
||
</taxonomicName>
|
||
shares with choristoderes several character states including: parietal foramen absent (4-1; also absent in many other aquatic reptiles); caudal zygapophyses lie nearly or completely vertical (58-1); and fibula narrow at proximal end and flared at distal end (70-1). None of these are individually diagnosable for the
|
||
<taxonomicName id="4C5E4D770F1D82029BDEFD3CBA8AFAE2" authorityName="COPE" authorityYear="1876" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="order">Choristodera</taxonomicName>
|
||
, and possible homologies of these characters in
|
||
<taxonomicName id="4C5E4D770F1D820299F1FD7EBB6AFACC" authorityName="Hecht" authorityYear="1992" box="[142,313,1361,1382]" class="Reptilia" genus="Lazarussuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAE60F1D820299F1FD7EBB6AFACC" box="[142,313,1361,1382]" italics="true" pageId="12" pageNumber="439">Lazarussuchus</emphasis>
|
||
</taxonomicName>
|
||
with choristoderes need to be further investigated. According to
|
||
<bibRefCitation id="EFCF4B050F1D820298BDFD40B834FA2C" author="Hecht MK" box="[450,615,1391,1414]" pageId="12" pageNumber="439" pagination="115 - 131" refId="ref11209" refString="Hecht MK. 1992. A new choristodere (Reptilia, Diapsida) from the Oligocene of France: an example of the Lazarus effect. Geobios 25: 115 - 131." type="journal article" year="1992">Hecht’s (1992)</bibRefCitation>
|
||
description,
|
||
<taxonomicName id="4C5E4D770F1D820299F1FDA1BB6AFA09" authorityName="Hecht" authorityYear="1992" box="[142,313,1422,1443]" class="Reptilia" genus="Lazarussuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAE60F1D820299F1FDA1BB6AFA09" box="[142,313,1422,1443]" italics="true" pageId="12" pageNumber="439">Lazarussuchus</emphasis>
|
||
</taxonomicName>
|
||
displays several features that are not otherwise known in choristoderes: paired narial openings located far back from the tip of the snout; prefrontals entering narial opening; vertebral centrum amphicoelous and notochordal; and presence of four sacral vertebrae with fused ribs. In addition, uniquely diagnostic character states, such as presence/absence of a neomorph bone in the braincase and presence/ absence of a pterygoquadrate foramen, are unknown for
|
||
<taxonomicName id="4C5E4D770F1D820299C5FE8DBB36F91D" authorityName="Hecht" authorityYear="1992" box="[186,357,1698,1719]" class="Reptilia" genus="Lazarussuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAE60F1D820299C5FE8DBB36F91D" box="[186,357,1698,1719]" italics="true" pageId="12" pageNumber="439">Lazarussuchus</emphasis>
|
||
</taxonomicName>
|
||
. Until these anatomical ambiguities can be clarified,
|
||
<taxonomicName id="4C5E4D770F1D820298FFFEEEB878F97C" authorityName="Hecht" authorityYear="1992" box="[384,555,1729,1750]" class="Reptilia" genus="Lazarussuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAE60F1D820298FFFEEEB878F97C" box="[384,555,1729,1750]" italics="true" pageId="12" pageNumber="439">Lazarussuchus</emphasis>
|
||
</taxonomicName>
|
||
is best treated as a taxon outside the
|
||
<taxonomicName id="4C5E4D770F1D8202980BFEF0B85FF95F" authorityName="COPE" authorityYear="1876" box="[372,524,1759,1781]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="order">Choristodera</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE136F40F1D820299D8FED1BF6BFD49" blockId="12.[142,758,379,1904]" lastBlockId="12.[806,1422,195,739]" pageId="12" pageNumber="439">
|
||
With the exclusion of
|
||
<taxonomicName id="4C5E4D770F1D820298E4FED1B815F8B9" authorityName="Hecht" authorityYear="1992" box="[411,582,1790,1811]" class="Reptilia" genus="Lazarussuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAE60F1D820298E4FED1B815F8B9" box="[411,582,1790,1811]" italics="true" pageId="12" pageNumber="439">Lazarussuchus</emphasis>
|
||
</taxonomicName>
|
||
, the order
|
||
<taxonomicName id="4C5E4D770F1D82029BBFFED1BAA6F899" authorityName="COPE" authorityYear="1876" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="order">Choristodera</taxonomicName>
|
||
can be unambiguously diagnosed by the following derived character states: prefrontals having median sutural contact for their entire length (2-2); parietal foramen absent (4-1; possibly homoplastic in
|
||
<taxonomicName id="4C5E4D770F1D82029A59F8CDB982FF5D" authorityName="Hecht" authorityYear="1992" box="[806,977,226,247]" class="Reptilia" genus="Lazarussuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAE60F1D82029A59F8CDB982FF5D" box="[806,977,226,247]" italics="true" pageId="12" pageNumber="439">Lazarussuchus</emphasis>
|
||
</taxonomicName>
|
||
and in many other aquatic reptiles); choana retracted close to midpoint of marginal tooth row (20-1); pterygoid flange consisting of pterygoid and ectopterygoid with a horizontal overlap (23-1); basipterygoid/pterygoid joint a sutural contact (24-1); parasphenoid and pterygoid having a clear sutural contact (25-1); postorbital process of jugal much shorter than anteroventral process (28-1); interpterygoid vacuity enclosed anteriorly by pterygoids and posteriorly by parasphenoid (33-1; further modification in
|
||
<taxonomicName id="4C5E4D770F1D82029C64F9D9B9D8FD81" authorityName="Cope" authorityYear="1876" class="Reptilia" family="Champsosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Choristodera" pageId="12" pageNumber="439" phylum="Chordata" rank="family">Champsosauridae</taxonomicName>
|
||
); neomorph in braincase present as part of external wall of braincase and medial wall of temporal fossa (40-1; unknown for
|
||
<taxonomicName id="4C5E4D770F1D82029D2BFA7DBEACFDCD" authorityName="Hecht" authorityYear="1992" box="[1108,1279,594,615]" class="Reptilia" genus="Lazarussuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eosuchia" pageId="12" pageNumber="439" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAE60F1D82029D2BFA7DBEACFDCD" box="[1108,1279,594,615]" italics="true" pageId="12" pageNumber="439">Lazarussuchus</emphasis>
|
||
</taxonomicName>
|
||
); free odontoid process unfused to axis (50-1); vertebral centra amphiplatyan with notochordal canal closed (52-1); sacral vertebrae three in number (56-1); sacral and caudal ribs remain free from vertebrae (57-1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |