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<document id="CFB33F8EA6A87EA3EE71BDE9E9F46AD4" ID-CLB-Dataset="22490" ID-DOI="10.11646/zootaxa.4767.4.3" ID-GBIF-Dataset="cf8cbc6a-80fe-4a63-874c-11ba7ed044a0" ID-ISSN="1175-5334" ID-Zenodo-Dep="3774902" ID-ZooBank="urn:lsid:zoobank.org:pub:A7123814-F67C-4295-AC00-4FD26AA0B2B2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588142469589" checkinUser="plazi" docAuthor="Golub, Viktor B. &amp; Heiss, Ernst" docDate="2020" docId="03F7575EFFE4FFC8FF71F9345CB8FC07" docLanguage="en" docName="zootaxa.4767.4.3.pdf" docOrigin="Zootaxa 4767 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tingiometra pankowskii Golub &amp; Heiss 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="536" masterDocId="FFCE2F26FFE5FFCDFFE6FFF05E46FFA0" masterDocTitle="Three new species of the lacebug genus Tingiometra from Cretaceous Burmese amber (Hemiptera: Heteroptera: Tingidae: Tingiometrinae)" masterLastPageNumber="542" masterPageNumber="531" pageNumber="532" updateTime="1698837992990" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="68FBF116FC7A5A99A36BD8F97EDAEFA9">Three new species of the lacebug genus Tingiometra from Cretaceous Burmese amber (Hemiptera: Heteroptera: Tingidae: Tingiometrinae)</mods:title>
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<mods:namePart id="1F4CB7C7FB816CB01390D21897980193">Golub, Viktor B.</mods:namePart>
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<mods:namePart id="74C72F075F8EF8CB5AD7BD88ADFC3428">Heiss, Ernst</mods:namePart>
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<treatment id="03F7575EFFE4FFC8FF71F9345CB8FC07" ID-DOI="http://doi.org/10.5281/zenodo.3796701" ID-GBIF-Taxon="163875134" ID-Zenodo-Dep="3796701" LSID="urn:lsid:plazi:treatment:03F7575EFFE4FFC8FF71F9345CB8FC07" httpUri="http://treatment.plazi.org/id/03F7575EFFE4FFC8FF71F9345CB8FC07" lastPageId="5" lastPageNumber="536" pageId="1" pageNumber="532">
<subSubSection id="C344B5C3FFE4FFCCFF71F9345C67F97E" box="[151,545,1731,1758]" pageId="1" pageNumber="532" type="nomenclature">
<paragraph id="8BE1E648FFE4FFCCFF71F9345C67F97E" blockId="1.[151,545,1731,1794]" box="[151,545,1731,1758]" pageId="1" pageNumber="532">
<heading id="D0A95124FFE4FFCCFF71F9345C67F97E" bold="true" box="[151,545,1731,1758]" fontSize="10" level="2" pageId="1" pageNumber="532" reason="2">
<emphasis id="B92A3A5AFFE4FFCCFF71F9345C67F97E" bold="true" box="[151,545,1731,1758]" pageId="1" pageNumber="532">
<taxonomicName id="4C5E9DCBFFE4FFCCFF71F9345FFAF97E" authority="Golub &amp; Heiss, 2020" authorityName="Golub &amp; Heiss" authorityYear="2020" box="[151,444,1731,1758]" class="Insecta" family="Tingidae" genus="Tingiometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="pankowskii" status="sp. nov.">
<emphasis id="B92A3A5AFFE4FFCCFF71F9345FFAF97E" bold="true" box="[151,444,1731,1758]" italics="true" pageId="1" pageNumber="532">Tingiometra pankowskii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2198721FFE4FFCCFE22F9345C67F97E" box="[452,545,1732,1758]" pageId="1" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="8BE1E648FFE4FFCCFF71F9175F53F8A2" blockId="1.[151,545,1731,1794]" box="[151,277,1767,1794]" pageId="1" pageNumber="532">
<figureCitation id="1365FACDFFE4FFCCFF71F9175F56F8A2" box="[151,272,1767,1794]" captionStart-0="FIGURES 14. 1" captionStart-1="FIGURES 58" captionStart-2="FIGURES 912. 9,10" captionStartId-0="3.[151,264,1766,1791]" captionStartId-1="6.[151,264,1962,1987]" captionStartId-2="8.[151,264,1940,1965]" captionTargetBox-0="[278,1305,197,1718]" captionTargetBox-1="[196,1389,186,1932]" captionTargetBox-2="[207,1380,189,1905]" captionTargetId-0="figure@3.[229,1358,178,1743]" captionTargetId-1="figure@6.[188,1400,181,1937]" captionTargetId-2="figure@8.[194,1393,181,1915]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionTargetPageId-2="8" captionText-0="FIGURES 14. 1, Tingiometra burmanica Heiss et al., 2015, holotype, dorsal view 24, Tingiometra pankowskii sp. nov., 2, holotype female (16/19-4), dorsal view, 3, holotype female (16/19-4), head and pronotum, 4, holotype female (16-19-4), pronotum and hemelytra. Scales: 1mm (1,2), 0.5mm (3,4)." captionText-1="FIGURES 58. Tingiometra pankowskii sp. nov., 5, paratype female (16-03), dorsal view, 6, paratype female (15/27), dorsal view, 7, paratype female (?) (15/31), dorsal view, 8, paratype male (?) (17/06), dorsal view. Scales: 1mm." captionText-2="FIGURES 912. 9,10 Tingiometra pankowskii sp. nov., 9, paratype male (?) (7303-1), dorsal view, 10, paratype male (?) (7303-1), ventral view; 11, Tingiometra secunda sp. nov., holotype female (16/19-2), dorsal view; 12, Tingiometra yuripopovi sp. nov., holotype female (18/25), dorsal view. Scales 1mm." figureDoi-0="http://doi.org/10.5281/zenodo.3774904" figureDoi-1="http://doi.org/10.5281/zenodo.3774906" figureDoi-2="http://doi.org/10.5281/zenodo.3774908" httpUri-0="https://zenodo.org/record/3774904/files/figure.png" httpUri-1="https://zenodo.org/record/3774906/files/figure.png" httpUri-2="https://zenodo.org/record/3774908/files/figure.png" pageId="1" pageNumber="532">Figs. 210</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C344B5C3FFE4FFCCFF71F8DA5C0CF858" pageId="1" pageNumber="532" type="materials_examined">
<paragraph id="8BE1E648FFE4FFCCFF71F8DA5A40F8E4" blockId="1.[151,1436,1833,2041]" box="[151,1030,1833,1861]" pageId="1" pageNumber="532">
<emphasis id="B92A3A5AFFE4FFCCFF71F8DA5F0EF8E4" bold="true" box="[151,328,1833,1860]" pageId="1" pageNumber="532">Type Material.</emphasis>
<materialsCitation id="3B36EC15FFE4FFCCFEA9F8DA5A40F8E4" ID-GBIF-Occurrence="2609409777" box="[335,1030,1834,1861]" collectionCode="CEHI" pageId="1" pageNumber="532" specimenCount="1" typeStatus="holotype">
<emphasis id="B92A3A5AFFE4FFCCFEA9F8DA5FFAF8E4" bold="true" box="[335,444,1834,1860]" pageId="1" pageNumber="532">
<typeStatus id="54E558EAFFE4FFCCFEA9F8DA5FFAF8E4" box="[335,444,1834,1860]" pageId="1" pageNumber="532" type="holotype">Holotype</typeStatus>
</emphasis>
female,
<figureCitation id="1365FACDFFE4FFCCFDFBF8DA5CCEF8E4" box="[541,648,1834,1860]" captionStart="FIGURES 14. 1" captionStartId="3.[151,264,1766,1791]" captionTargetBox="[278,1305,197,1718]" captionTargetId="figure@3.[229,1358,178,1743]" captionTargetPageId="3" captionText="FIGURES 14. 1, Tingiometra burmanica Heiss et al., 2015, holotype, dorsal view 24, Tingiometra pankowskii sp. nov., 2, holotype female (16/19-4), dorsal view, 3, holotype female (16/19-4), head and pronotum, 4, holotype female (16-19-4), pronotum and hemelytra. Scales: 1mm (1,2), 0.5mm (3,4)." figureDoi="http://doi.org/10.5281/zenodo.3774904" httpUri="https://zenodo.org/record/3774904/files/figure.png" pageId="1" pageNumber="532">Figs. 24</figureCitation>
, BUB-TING-08 (16/19-4),
<collectionCode id="ED4F7E8DFFE4FFCCFC5BF8DA5A44F8E4" box="[957,1026,1834,1860]" pageId="1" pageNumber="532">CEHI</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE1E648FFE4FFCCFF21F8BE5DA2F82C" blockId="1.[151,1436,1833,2041]" pageId="1" pageNumber="532">
<materialsCitation id="3B36EC15FFE4FFCCFF21F8BE5DA6F82C" ID-GBIF-Occurrence="2609409774" collectionCode="CEHI" pageId="1" pageNumber="532" specimenCount="1" typeStatus="paratype">
<emphasis id="B92A3A5AFFE4FFCCFF21F8BE5F0EF8C8" bold="true" box="[199,328,1870,1896]" pageId="1" pageNumber="532">
<typeStatus id="54E558EAFFE4FFCCFF21F8BE5F05F8C8" box="[199,323,1870,1896]" pageId="1" pageNumber="532" type="paratype">Paratypes</typeStatus>
:
</emphasis>
female,
<figureCitation id="1365FACDFFE4FFCCFE4DF8BE5FA9F8C8" box="[427,495,1869,1896]" captionStart="FIGURES 58" captionStartId="6.[151,264,1962,1987]" captionTargetBox="[196,1389,186,1932]" captionTargetId="figure@6.[188,1400,181,1937]" captionTargetPageId="6" captionText="FIGURES 58. Tingiometra pankowskii sp. nov., 5, paratype female (16-03), dorsal view, 6, paratype female (15/27), dorsal view, 7, paratype female (?) (15/31), dorsal view, 8, paratype male (?) (17/06), dorsal view. Scales: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3774906" httpUri="https://zenodo.org/record/3774906/files/figure.png" pageId="1" pageNumber="532">Fig. 5</figureCitation>
, BUB-TING-08a (16/03); female,
<figureCitation id="1365FACDFFE4FFCCFC9EF8BE5DFAF8C8" box="[888,956,1870,1896]" captionStart="FIGURES 58" captionStartId="6.[151,264,1962,1987]" captionTargetBox="[196,1389,186,1932]" captionTargetId="figure@6.[188,1400,181,1937]" captionTargetPageId="6" captionText="FIGURES 58. Tingiometra pankowskii sp. nov., 5, paratype female (16-03), dorsal view, 6, paratype female (15/27), dorsal view, 7, paratype female (?) (15/31), dorsal view, 8, paratype male (?) (17/06), dorsal view. Scales: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3774906" httpUri="https://zenodo.org/record/3774906/files/figure.png" pageId="1" pageNumber="532">Fig. 6</figureCitation>
, BUB-TING-08b (15/27); female?,
<figureCitation id="1365FACDFFE4FFCCFAB5F8BE5BD1F8C9" box="[1363,1431,1870,1897]" captionStart="FIGURES 58" captionStartId="6.[151,264,1962,1987]" captionTargetBox="[196,1389,186,1932]" captionTargetId="figure@6.[188,1400,181,1937]" captionTargetPageId="6" captionText="FIGURES 58. Tingiometra pankowskii sp. nov., 5, paratype female (16-03), dorsal view, 6, paratype female (15/27), dorsal view, 7, paratype female (?) (15/31), dorsal view, 8, paratype male (?) (17/06), dorsal view. Scales: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3774906" httpUri="https://zenodo.org/record/3774906/files/figure.png" pageId="1" pageNumber="532">Fig. 7</figureCitation>
, BUB-TING-08 c (15/31); male?,
<figureCitation id="1365FACDFFE4FFCCFDE9F8825C17F82C" box="[527,593,1906,1932]" captionStart="FIGURES 58" captionStartId="6.[151,264,1962,1987]" captionTargetBox="[196,1389,186,1932]" captionTargetId="figure@6.[188,1400,181,1937]" captionTargetPageId="6" captionText="FIGURES 58. Tingiometra pankowskii sp. nov., 5, paratype female (16-03), dorsal view, 6, paratype female (15/27), dorsal view, 7, paratype female (?) (15/31), dorsal view, 8, paratype male (?) (17/06), dorsal view. Scales: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3774906" httpUri="https://zenodo.org/record/3774906/files/figure.png" pageId="1" pageNumber="532">Fig. 8</figureCitation>
, BUB-TING-08d (17/06), in
<collectionCode id="ED4F7E8DFFE4FFCCFC7DF8825DA6F82C" box="[923,992,1906,1932]" pageId="1" pageNumber="532">CEHI</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE1E648FFE4FFCCFF21F8665C0CF858" blockId="1.[151,1436,1833,2041]" pageId="1" pageNumber="532">
<emphasis id="B92A3A5AFFE4FFCCFF21F8665F40F810" bold="true" box="[199,262,1942,1968]" pageId="1" pageNumber="532">Note.</emphasis>
<typeStatus id="54E558EAFFE4FFCCFEF6F8675F34F811" box="[272,370,1943,1969]" pageId="1" pageNumber="532" type="paratype">Paratype</typeStatus>
15/27 was formerly regarded and tentatively designated as
<typeStatus id="54E558EAFFE4FFCCFBC5F8675AC5F811" box="[1059,1155,1943,1969]" pageId="1" pageNumber="532" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C5E9DCBFFE4FFCCFB4BF8675FD0F874" authority="(Heiss, Golub &amp; Popov, 2015)" baseAuthorityName="Heiss, Golub &amp; Popov" baseAuthorityYear="2015" class="Insecta" family="Tingidae" genus="Tingiometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="burmanica">
<emphasis id="B92A3A5AFFE4FFCCFB4BF8675B02F810" box="[1197,1348,1942,1968]" italics="true" pageId="1" pageNumber="532">T. burmanica</emphasis>
(
<bibRefCitation id="EFCF9BB9FFE4FFCCFAB0F8665FC8F874" author="Heiss, E. &amp; Golub, V. B. &amp; Popov, Yu. A." pageId="1" pageNumber="532" pagination="1 - 9" refId="ref7677" refString="Heiss, E., Golub, V. B. &amp; Popov, Yu. A. (2015) A new subfamily, genus and species of Tingidae (Hemiptera: Heteroptera) from Burmese amber. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 67, 1 - 9." type="journal article" year="2015">Heiss, Golub &amp; Popov, 2015</bibRefCitation>
)
</taxonomicName>
but now recognized to belong to this new species.
<typeStatus id="54E558EAFFE4FFCCFC36F84B5A7BF875" box="[976,1085,1979,2005]" pageId="1" pageNumber="532" type="paratype">Paratypes</typeStatus>
15/27 and 15/31 are embedded in blocks of transparent artificial resin.
</paragraph>
</subSubSection>
<subSubSection id="C344B5C3FFE7FFCFFF21FF685A7CFDBB" pageId="2" pageNumber="533" type="diagnosis">
<paragraph id="8BE1E648FFE7FFCFFF21FF685A7EFE0F" blockId="2.[151,1437,152,2044]" pageId="2" pageNumber="533">
<emphasis id="B92A3A5AFFE7FFCFFF21FF685F04FF13" bold="true" box="[199,322,152,179]" pageId="2" pageNumber="533">Diagnosis.</emphasis>
This new species differs from
<taxonomicName id="4C5E9DCBFFE7FFCFFD45FF6A5D7DFF13" baseAuthorityName="Heiss, Golub &amp; Popov" baseAuthorityYear="2015" box="[675,827,153,179]" class="Insecta" family="Tingidae" genus="Tingiometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="burmanica">
<emphasis id="B92A3A5AFFE7FFCFFD45FF6A5CF5FF13" box="[675,691,154,179]" italics="true" pageId="2" pageNumber="533">T</emphasis>
.
<emphasis id="B92A3A5AFFE7FFCFFD24FF695D7DFF13" box="[706,827,153,179]" italics="true" pageId="2" pageNumber="533">burmanica</emphasis>
</taxonomicName>
by much wider discoidal area of hemelytra including five large cells and having 1922 rows of small areolae at the widest part (vs 3 large cells and 78 rows of small areolae at the widest part in
<taxonomicName id="4C5E9DCBFFE7FFCFFE33FF125C2FFF5B" baseAuthorityName="Heiss, Golub &amp; Popov" baseAuthorityYear="2015" box="[469,617,225,251]" class="Insecta" family="Tingidae" genus="Tingiometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="burmanica">
<emphasis id="B92A3A5AFFE7FFCFFE33FF125C2FFF5B" box="[469,617,225,251]" italics="true" pageId="2" pageNumber="533">T. burmanica</emphasis>
</taxonomicName>
), narrower subcostal area with about 7 rows of areolae at the widest part (vs 10 rows), narrower and rib-like costal area, wider head (
<quantity id="4CA64BADFFE7FFCFFCC8FEF45D90FEBF" box="[814,982,260,287]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="5.2" metricValueMin="3.0" pageId="2" pageNumber="533" unit="mm" value="0.41" valueMax="0.52" valueMin="0.3">0.300.52 mm</quantity>
; vs
<quantity id="4CA64BADFFE7FFCFFBE6FEF45A21FEBF" box="[1024,1127,260,287]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="533" unit="mm" value="0.25">0.25 mm</quantity>
), pronotum relatively wider (1.301.58 times as wide as long; vs 1.33 times), scutellum relatively wider (1.13 times as wide as long; vs about 1.5 times), median carina lower with only one row of areolae or without areolae (vs two rows of areolae in
<taxonomicName id="4C5E9DCBFFE7FFCFFAB7FEBE5EAFFE2B" baseAuthorityName="Heiss, Golub &amp; Popov" baseAuthorityYear="2015" class="Insecta" family="Tingidae" genus="Tingiometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="burmanica">
<emphasis id="B92A3A5AFFE7FFCFFAB7FEBE5EAFFE2B" italics="true" pageId="2" pageNumber="533">T. burmanica</emphasis>
</taxonomicName>
), The main differences from other new species of
<taxonomicName id="4C5E9DCBFFE7FFCFFCF9FE825A4DFE2B" authority="Golub &amp; Heiss, 2020" authorityName="Golub &amp; Heiss" authorityYear="2020" box="[799,1035,369,395]" class="Insecta" family="Tingidae" genus="Tingiometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="secunda" status="sp. nov.">
<emphasis id="B92A3A5AFFE7FFCFFCF9FE825A4DFE2B" box="[799,1035,369,395]" italics="true" pageId="2" pageNumber="533">Tingiometra secunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2198721FFE7FFCFFBF4FE815A25FE2B" box="[1042,1123,369,395]" pageId="2" pageNumber="533" rank="species">sp. nov</taxonomicNameLabel>
and
<taxonomicName id="4C5E9DCBFFE7FFCFFB7FFE825B54FE2B" authority="Golub &amp; Heiss, 2020" authorityName="Golub &amp; Heiss" authorityYear="2020" box="[1177,1298,370,395]" class="Insecta" family="Tingidae" genus="Tingiometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="yuripopovi" status="sp. nov.">
<emphasis id="B92A3A5AFFE7FFCFFB7FFE825B54FE2B" box="[1177,1298,370,395]" italics="true" pageId="2" pageNumber="533">yuripopovi</emphasis>
</taxonomicName>
<emphasis id="B92A3A5AFFE7FFCFFAFFFE815B35FE2B" bold="true" box="[1305,1395,369,395]" pageId="2" pageNumber="533">
<taxonomicNameLabel id="A2198721FFE7FFCFFAFFFE815B35FE2B" box="[1305,1395,369,395]" pageId="2" pageNumber="533" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are indicated in their descriptions below and in the key to all four
<taxonomicName id="4C5E9DCBFFE7FFCFFCB6FE665D9FFE0F" authorityName="Heiss, Golub &amp; Popov" authorityYear="2015" box="[848,985,406,431]" class="Insecta" family="Tingidae" genus="Tingiometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="B92A3A5AFFE7FFCFFCB6FE665D9FFE0F" box="[848,985,406,431]" italics="true" pageId="2" pageNumber="533">Tingiometra</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="8BE1E648FFE7FFCFFF21FE495A7CFDBB" blockId="2.[151,1437,152,2044]" pageId="2" pageNumber="533">
<emphasis id="B92A3A5AFFE7FFCFFF21FE495F40FE73" bold="true" box="[199,262,441,467]" pageId="2" pageNumber="533">Note.</emphasis>
The position of the
<typeStatus id="54E558EAFFE7FFCFFE05FE495C03FE73" box="[483,581,441,467]" pageId="2" pageNumber="533" type="holotype">holotype</typeStatus>
and
<typeStatus id="54E558EAFFE7FFCFFD9CFE495CA3FE73" box="[634,741,441,467]" pageId="2" pageNumber="533" type="paratype">paratypes</typeStatus>
in the amber pieces are different. In addition, specimens of the type series show morphological differences as an evidence of intraspecific variability. Therefore, the description of the
<typeStatus id="54E558EAFFE7FFCFFF26FDF15F64FDBB" box="[192,290,513,539]" pageId="2" pageNumber="533" type="holotype">holotype</typeStatus>
and the morphological features of the
<typeStatus id="54E558EAFFE7FFCFFD32FDF15D79FDBB" box="[724,831,513,539]" pageId="2" pageNumber="533" type="paratype">paratypes</typeStatus>
are treated separately.
</paragraph>
</subSubSection>
<subSubSection id="C344B5C3FFE7FFC8FF21FDD45F95FCB7" lastPageId="5" lastPageNumber="536" pageId="2" pageNumber="533" type="description">
<paragraph id="8BE1E648FFE7FFCFFF21FDD45C54FD0B" blockId="2.[151,1437,152,2044]" pageId="2" pageNumber="533">
<emphasis id="B92A3A5AFFE7FFCFFF21FDD45FA4FD9F" bold="true" box="[199,482,548,575]" pageId="2" pageNumber="533">
Description of
<typeStatus id="54E558EAFFE7FFCFFE9DFDD55FA4FD9F" box="[379,482,549,575]" pageId="2" pageNumber="533" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="1365FACDFFE7FFCFFE14FDD55C19FD9F" box="[498,607,549,575]" captionStart="FIGURES 14. 1" captionStartId="3.[151,264,1766,1791]" captionTargetBox="[278,1305,197,1718]" captionTargetId="figure@3.[229,1358,178,1743]" captionTargetPageId="3" captionText="FIGURES 14. 1, Tingiometra burmanica Heiss et al., 2015, holotype, dorsal view 24, Tingiometra pankowskii sp. nov., 2, holotype female (16/19-4), dorsal view, 3, holotype female (16/19-4), head and pronotum, 4, holotype female (16-19-4), pronotum and hemelytra. Scales: 1mm (1,2), 0.5mm (3,4)." figureDoi="http://doi.org/10.5281/zenodo.3774904" httpUri="https://zenodo.org/record/3774904/files/figure.png" pageId="2" pageNumber="533">Figs. 24</figureCitation>
). Body elongate, oval, about 2.54 times as long as wide, hemelytra with rows of very small areolae separated by carinate veins. Head and anterior part of pronotum (calli) with very small punctuation, which is much larger at posterior part. At least head dorsally and femora covered by very short, light, adhering and slightly raised hairs.
</paragraph>
<paragraph id="8BE1E648FFE7FFCFFF21FD455C83FC23" blockId="2.[151,1437,152,2044]" pageId="2" pageNumber="533">
<emphasis id="B92A3A5AFFE7FFCFFF21FD455F42FD6F" box="[199,260,693,719]" italics="true" pageId="2" pageNumber="533">Head</emphasis>
2.53 times as long as wide across eyes counting the length of the head from apex of clypeus to posterior margin of eye and 2.77 times counting the length of the head from apex of clypeus to the posterior margin of the frons. Clypeus long, directed anteriorly, horizontal, separated from cheeks by furrows, with longitudinal and moderately wide crest. Apex of head diamond-shaped expanded. Vertex and frons flat. Lateral margins of postocular portion slightly converging posteriorly. Eyes subglobose dorsally, prominently protruding on the head sides and covering the head laterally and partially ventrally.
</paragraph>
<paragraph id="8BE1E648FFE7FFCFFF21FC7D5C42FC4F" blockId="2.[151,1437,152,2044]" pageId="2" pageNumber="533">Antenniferous tubercles very small, adhering. Antennae long and very thin except thickened segment I, segments III and IV shorter and thinner than II; the whole antenna and its segment II 4.9 and 2.27 times as long as width of head across eyes respectively.</paragraph>
<paragraph id="8BE1E648FFE7FFCFFF21FC0A5F2CFAAE" blockId="2.[151,1437,152,2044]" pageId="2" pageNumber="533">
<emphasis id="B92A3A5AFFE7FFCFFF21FC0A5F70FBB3" box="[199,310,1018,1043]" italics="true" pageId="2" pageNumber="533">Pronotum</emphasis>
strongly widening posteriorly, 1.58 times as wide as long at base; its anterior margin not wider than the posterior margin of head. Anterior pronotal margin distinct, posterior one strongly sinuate. Posterior margin not produced and not overlapping scutellum, which is entirely visible in dorsal view. Lateral pronotal margins straight. Pronotal disk slightly convex, with rather large punctures, very low longitudinal median carina without areolae, consisting of a longer posterior part and a shorter anterior part; both parts of carina separated by transverse impression (poorly visible in
<typeStatus id="54E558EAFFE7FFCFFE72FB5B5FB0FB65" box="[404,502,1195,1221]" pageId="2" pageNumber="533" type="holotype">holotype</typeStatus>
due to a deformation of pronotum), posterior part of carina about 34 times as long as anterior one. Paranota forming very thin translucent plates with serrate external margin, without distinct areolae (possibly artefact).
</paragraph>
<paragraph id="8BE1E648FFE7FFCFFF21FAE75A62FAF5" blockId="2.[151,1437,152,2044]" pageId="2" pageNumber="533">
<emphasis id="B92A3A5AFFE7FFCFFF21FAE75F73FA91" box="[199,309,1303,1329]" italics="true" pageId="2" pageNumber="533">Scutellum</emphasis>
dorsally entirely visible, large, triangular and completely exposed; 1.13 times as wide as long; its apex straightly cut, distinctly enlarged; surface of scutellum very finely punctate.
</paragraph>
<paragraph id="8BE1E648FFE7FFCFFF21FAAE5F9AF8A1" blockId="2.[151,1437,152,2044]" pageId="2" pageNumber="533">
<emphasis id="B92A3A5AFFE7FFCFFF21FAAE5F7BFAD8" box="[199,317,1374,1400]" italics="true" pageId="2" pageNumber="533">Hemelytra</emphasis>
submacropterous, not reaching apex of abdomen. Costal area very narrow, with one row of very small areolae. Subcostal, discoidal, and sutural areas distinctly separated by carinate veins. Subcostal area declivous, with about 7 rows of areolae at widest parts, divided by 5 and 6 transverse carinate veins different in both hemelytra. Discoidal area very wide, exteriorly limited by
<emphasis id="B92A3A5AFFE7FFCFFCF8FA3A5D69FA43" box="[798,815,1482,1507]" italics="true" pageId="2" pageNumber="533">R</emphasis>
vein and interiorly limited by
<emphasis id="B92A3A5AFFE7FFCFFB99FA3A5AE6FA44" box="[1151,1184,1482,1508]" italics="true" pageId="2" pageNumber="533">Cu</emphasis>
vein, with 19 irregular rows of areolae at widest part, divided by longitudinal bent carinate vein (evidently
<emphasis id="B92A3A5AFFE7FFCFFBB5FA1E5A2CF9A7" box="[1107,1130,1518,1543]" italics="true" pageId="2" pageNumber="533">M</emphasis>
). Discoidal area is divided into five large cells: external basal and apical cells adjoining to subcostal area and three internal cells (basal, medial and apical) located closer to the longitudinal axis of the body and adjacent to clavus and sutural area of hemelytron; external basal cell the largest. Every large cell of discoidal area contains many small round areolae. Sutural area at the base narrow, with 12 rows of areolae, posteriorly broaden and with about 6 rows of areolae at apex. Reduced apical half of sutural area is part of hemelytral membrane. Clavus distinct, with 7 rows of areolae at widest part, commissure longer than scutellum. The total surface of discoidal and sutural areas and clavus nearly flat. Venation of posterior wings is unclear.
</paragraph>
<paragraph id="8BE1E648FFE7FFCFFF21F8FC5C01F85C" blockId="2.[151,1437,152,2044]" pageId="2" pageNumber="533">
<emphasis id="B92A3A5AFFE7FFCFFF21F8FC5F49F885" box="[199,271,1804,1829]" italics="true" pageId="2" pageNumber="533">Venter</emphasis>
. Bucculae very short (laterally seen), occupying only the apex of head ventrally, closed anteriorly, surrounding the base of labium like a cover. Labium 4-segmented, its apex reaching metasternum. Sternal thoracic plates absent. Metepisternum with long curved groove possibly representing peritreme of the opening of the scent gland apparatus. The posterior margin of metasternum with a process, which narrows to the apex and reaches metacoxae. Abdominal sternites II and III completely fused, sternites IVVIII separated by transverse furrows, sternite VII has two (1+1) narrow lateral projections directed posteriorly. Segment IX is located unusually over the segments VII and VIII and only dorsally visible.
</paragraph>
<caption id="DF21B6C0FFE6FFCEFF71F9165CEAF8E7" ID-DOI="http://doi.org/10.5281/zenodo.3774904" ID-Zenodo-Dep="3774904" httpUri="https://zenodo.org/record/3774904/files/figure.png" pageId="3" pageNumber="534" startId="3.[151,264,1766,1791]" targetBox="[278,1305,197,1718]" targetPageId="3">
<paragraph id="8BE1E648FFE6FFCEFF71F9165CEAF8E7" blockId="3.[151,1437,1766,1863]" pageId="3" pageNumber="534">
<emphasis id="B92A3A5AFFE6FFCEFF71F9165F15F95F" bold="true" box="[151,339,1766,1791]" pageId="3" pageNumber="534">FIGURES 14. 1,</emphasis>
<taxonomicName id="4C5E9DCBFFE6FFCEFEBCF9175D4DF95F" authority="Heiss et al., 2015" authorityName="Heiss" authorityYear="2015" box="[346,779,1766,1791]" class="Insecta" family="Tingidae" genus="Tingiometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="burmanica">
<emphasis id="B92A3A5AFFE6FFCEFEBCF9175C14F95F" bold="true" box="[346,594,1766,1791]" italics="true" pageId="3" pageNumber="534">Tingiometra burmanica</emphasis>
<bibRefCitation id="EFCF9BB9FFE6FFCEFDBFF9165D4DF95F" author="Heiss, E. &amp; Golub, V. B. &amp; Popov, Yu. A." box="[601,779,1766,1791]" pageId="3" pageNumber="534" pagination="1 - 9" refId="ref7677" refString="Heiss, E., Golub, V. B. &amp; Popov, Yu. A. (2015) A new subfamily, genus and species of Tingidae (Hemiptera: Heteroptera) from Burmese amber. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 67, 1 - 9." type="journal article" year="2015">
Heiss
<emphasis id="B92A3A5AFFE6FFCEFD71F9185C96F95F" box="[663,720,1767,1791]" italics="true" pageId="3" pageNumber="534">et al.,</emphasis>
2015
</bibRefCitation>
</taxonomicName>
, holotype, dorsal view
<emphasis id="B92A3A5AFFE6FFCEFC10F9165A5AF95F" bold="true" box="[1014,1052,1766,1791]" pageId="3" pageNumber="534">24</emphasis>
,
<emphasis id="B92A3A5AFFE6FFCEFBCFF9175BDAF95F" bold="true" box="[1065,1436,1766,1791]" pageId="3" pageNumber="534">
<taxonomicName id="4C5E9DCBFFE6FFCEFBCFF9175B63F95F" authority="Golub &amp; Heiss, 2020" authorityName="Golub &amp; Heiss" authorityYear="2020" box="[1065,1317,1766,1791]" class="Insecta" family="Tingidae" genus="Tingiometra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="pankowskii" status="sp. nov.">
<emphasis id="B92A3A5AFFE6FFCEFBCFF9175B63F95F" bold="true" box="[1065,1317,1766,1791]" italics="true" pageId="3" pageNumber="534">Tingiometra pankowskii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2198721FFE6FFCEFACAF9175B38F95F" box="[1324,1406,1767,1791]" pageId="3" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
, 2,
</emphasis>
holotype female (16/19-4), dorsal view,
<emphasis id="B92A3A5AFFE6FFCEFDCAF8FA5C7FF883" bold="true" box="[556,569,1802,1827]" pageId="3" pageNumber="534">3</emphasis>
, holotype female (16/19-4), head and pronotum,
<emphasis id="B92A3A5AFFE6FFCEFBCFF8FA5A7AF883" bold="true" box="[1065,1084,1802,1827]" pageId="3" pageNumber="534">4,</emphasis>
holotype female (16-19-4), pronotum and hemelytra. Scales: 1mm (1,2), 0.5mm (3,4).
</paragraph>
</caption>
<paragraph id="8BE1E648FFE6FFCEFF21F8865A27F830" blockId="3.[151,1437,1909,2044]" box="[199,1121,1909,1936]" pageId="3" pageNumber="534">
<emphasis id="B92A3A5AFFE6FFCEFF21F8865EBAF82F" box="[199,252,1910,1935]" italics="true" pageId="3" pageNumber="534">Legs</emphasis>
long and slender (details of the legs structure of the
<typeStatus id="54E558EAFFE6FFCEFCAFF8855DEDF82F" box="[841,939,1909,1935]" pageId="3" pageNumber="534" type="holotype">holotype</typeStatus>
are not visible).
</paragraph>
<paragraph id="8BE1E648FFE6FFC9FF21F8695AF6FEC8" blockId="3.[151,1437,1909,2044]" lastBlockId="4.[151,1436,152,360]" lastPageId="4" lastPageNumber="535" pageId="3" pageNumber="534">
<emphasis id="B92A3A5AFFE6FFCEFF21F8695C46F813" bold="true" box="[199,512,1944,1971]" pageId="3" pageNumber="534">
Measurements of
<typeStatus id="54E558EAFFE6FFCEFE7FF8695C46F813" box="[409,512,1945,1971]" pageId="3" pageNumber="534" type="holotype">holotype</typeStatus>
</emphasis>
(in mm). Body length 3.00; body width 1.18 (hemelytra slightly apart); head length from apex of clypeus to posterior margin of eye; head length including postocular portion 0.83; length of anteocular portion 0.40; length of eye 0.15; length of postocular portion 0.28; head width across eyes 0.30; width of anteocular part of head 0.125; width of postocular part (approximately, due to an oblique placed head in the piece of amber) 0.20; length of antenna 1,48; length of antennal segments (I, II, III, IV) 0.14, 0.68, 0,23, 0.43; pronotum length 0.43; pronotum width (approximately, due to an oblique placement in the piece of amber and its deformation) 0.68; scutellum length 0.30; scutellum width 0.34; hemelytra length 1.75; width of one wing 0.55; length of commissure between clavus and corium 0.70; length of anterior femora approximately 0.68; hind tibia approximately 1.50 (other segments of legs cannot be measured due to their oblique arrangement in the piece of amber).
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF71FE495CD5FE73" blockId="4.[151,659,441,467]" box="[151,659,441,467]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF71FE495CD5FE73" bold="true" box="[151,659,441,467]" pageId="4" pageNumber="535">
Morphological peculiarities of
<typeStatus id="54E558EAFFE1FFC9FDF1FE495CD5FE73" box="[535,659,441,467]" pageId="4" pageNumber="535" type="paratype">paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF71FDF15DE4FDBB" blockId="4.[151,1437,512,1187]" box="[151,930,512,539]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF71FDF15F0DFDBB" bold="true" box="[151,331,512,539]" pageId="4" pageNumber="535">
<typeStatus id="54E558EAFFE1FFC9FF71FDF15F42FDBB" box="[151,260,513,539]" pageId="4" pageNumber="535" type="paratype">Paratype</typeStatus>
16/03
</emphasis>
(female;
<figureCitation id="1365FACDFFE1FFC9FE50FDF15FBFFDBB" box="[438,505,512,539]" captionStart="FIGURES 58" captionStartId="6.[151,264,1962,1987]" captionTargetBox="[196,1389,186,1932]" captionTargetId="figure@6.[188,1400,181,1937]" captionTargetPageId="6" captionText="FIGURES 58. Tingiometra pankowskii sp. nov., 5, paratype female (16-03), dorsal view, 6, paratype female (15/27), dorsal view, 7, paratype female (?) (15/31), dorsal view, 8, paratype male (?) (17/06), dorsal view. Scales: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3774906" httpUri="https://zenodo.org/record/3774906/files/figure.png" pageId="4" pageNumber="535">Fig. 5</figureCitation>
). Body glabrous, completely hairless.
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF21FDD55E9BFD0B" blockId="4.[151,1437,512,1187]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF21FDD55F42FD9F" box="[199,260,549,575]" italics="true" pageId="4" pageNumber="535">Head</emphasis>
1.38 times as long as wide across eyes counting the length of the head from apex of clypeus to posterior margin of eye, and 2.05 times counting the length of the head from apex of clypeus to the posterior margin of the frons. The whole antenna and its segment II 4.1 and 1.87 times as long as width of head across of eyes respectively.
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF21FD465DD4FCB7" blockId="4.[151,1437,512,1187]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF21FD465F70FD6F" box="[199,310,694,719]" italics="true" pageId="4" pageNumber="535">Pronotum</emphasis>
1.4 times as wide at base as long. Lateral pronotal margins noticeably sinuate. Paranota rather narrow, slightly reflexed, in the form of very thin, semitransparent plates, with two rows of small areolae at anterior half and with one row at posterior half. Scutellum1.61 times as wide as long.
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF21FCD15C83FC23" blockId="4.[151,1437,512,1187]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF21FCD15F7BFC9B" box="[199,317,801,827]" italics="true" pageId="4" pageNumber="535">Hemelytra</emphasis>
submacropterous, covering apex of abdomen and noticeably protruding posteriorly. Subcostal area of hemelytra divided by 4 transverse carinate veins. Discoidal area with 2022 irregular rows of areolae at widest part. Clavus with 6 rows of areolae at widest part.
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF21FC7E5DD8FBFB" blockId="4.[151,1437,512,1187]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF21FC7E5F49FC07" box="[199,271,910,935]" italics="true" pageId="4" pageNumber="535">Venter</emphasis>
. Apex of labium reaching middle coxa. A narrow, slightly curved groove runs along the meso-metapleural junction, possibly representing peritreme of the opening of the scent gland apparatus. Sternite VII has a median and 2 (1+1) lateral projections directed posteriorly; median projection (not clearly visible) triangular, with a wide base, lateral projections have a wide triangular base continuing in a long narrow process pointed at the apex. Median projection represents the subgenital plate of segment VII (
<bibRefCitation id="EFCF9BB9FFE1FFC9FCC2FBED5A5CFB97" author="Drake, C. J. &amp; Davis, N. T." box="[804,1050,1053,1079]" pageId="4" pageNumber="535" pagination="1 - 100" refId="ref7514" refString="Drake, C. J. &amp; Davis, N. T. (1960) The morphology, phylogeny and higher classification of the family Tingidae, including the description of a new genus and species of the subfamily Vianaidinae (Heteroptera). Entomologia Americana, New Series, 39, 1 - 100." type="journal article" year="1960">Drake &amp; Davis, 1960</bibRefCitation>
;
<bibRefCitation id="EFCF9BB9FFE1FFC9FBC0FBED5AA9FB97" author="Heiss, E. &amp; Golub, V. B. &amp; Popov, Yu. A." box="[1062,1263,1052,1079]" pageId="4" pageNumber="535" pagination="1 - 9" refId="ref7677" refString="Heiss, E., Golub, V. B. &amp; Popov, Yu. A. (2015) A new subfamily, genus and species of Tingidae (Hemiptera: Heteroptera) from Burmese amber. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 67, 1 - 9." type="journal article" year="2015">
Heiss
<emphasis id="B92A3A5AFFE1FFC9FB8AFBEE5AD8FB97" box="[1132,1182,1053,1079]" italics="true" pageId="4" pageNumber="535">et al</emphasis>
., 2015
</bibRefCitation>
), but the suture between the plate and the basal part of the segment VII is not visible.
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF21FB965F5CFB03" blockId="4.[151,1437,512,1187]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF21FB965EBAFBDF" box="[199,252,1126,1151]" italics="true" pageId="4" pageNumber="535">Legs</emphasis>
long and slender, femora and tibiae cylindrical; tarsi two-segmented, claws thin and long, pulvilli not discernible.
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF71FB215CFCFB4B" blockId="4.[151,1437,1232,1727]" box="[151,698,1232,1260]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF71FB215F0DFB4B" bold="true" box="[151,331,1232,1259]" pageId="4" pageNumber="535">
<typeStatus id="54E558EAFFE1FFC9FF71FB215F42FB4B" box="[151,260,1233,1259]" pageId="4" pageNumber="535" type="paratype">Paratype</typeStatus>
15/27
</emphasis>
(female.
<figureCitation id="1365FACDFFE1FFC9FE53FB215FBFFB4C" box="[437,505,1233,1260]" captionStart="FIGURES 58" captionStartId="6.[151,264,1962,1987]" captionTargetBox="[196,1389,186,1932]" captionTargetId="figure@6.[188,1400,181,1937]" captionTargetPageId="6" captionText="FIGURES 58. Tingiometra pankowskii sp. nov., 5, paratype female (16-03), dorsal view, 6, paratype female (15/27), dorsal view, 7, paratype female (?) (15/31), dorsal view, 8, paratype male (?) (17/06), dorsal view. Scales: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3774906" httpUri="https://zenodo.org/record/3774906/files/figure.png" pageId="4" pageNumber="535">Fig. 6</figureCitation>
). Body glabrous.
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF21FB055CC6FA3F" blockId="4.[151,1437,1232,1727]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF21FB055F42FAAF" box="[199,260,1269,1295]" italics="true" pageId="4" pageNumber="535">Head</emphasis>
. Crest on clypeus surface narrow. Eyes very large, covering the sides of the head and partly its ventral surface. Ventral surface of head from base of bucculae to almost anterior margin of eyes with low carina. Posterior margin of pronotum almost right, provided with a very narrow strip bearing one row of very small areolae. Long groove extending along the anterior margin of the left metepisternum is apparently an external groove (peritreme) of the opening of the scent gland apparatus.
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF21FA5A5F32F91F" blockId="4.[151,1437,1232,1727]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF21FA5A5F3FFA63" box="[199,377,1450,1475]" italics="true" pageId="4" pageNumber="535">Posterior wings</emphasis>
. Wings widely spread. Venation of posterior wings as follows: vein
<emphasis id="B92A3A5AFFE1FFC9FB96FA595ACCFA63" box="[1136,1162,1449,1475]" italics="true" pageId="4" pageNumber="535">Sc</emphasis>
is not visible apparently due to its location very close to the costal margin or because of a deformation of the anterior margin of the wing. Vein
<emphasis id="B92A3A5AFFE1FFC9FF35FA025EA2F9AB" box="[211,228,1522,1547]" italics="true" pageId="4" pageNumber="535">R</emphasis>
is located very close to costal margin in the basal half of the wing and merges with
<emphasis id="B92A3A5AFFE1FFC9FB4FFA025A86F9AB" box="[1193,1216,1522,1547]" italics="true" pageId="4" pageNumber="535">M</emphasis>
forming
<emphasis id="B92A3A5AFFE1FFC9FACDFA025B15F9AB" box="[1323,1363,1522,1547]" italics="true" pageId="4" pageNumber="535">RM</emphasis>
in the apical part of the wing. Long
<emphasis id="B92A3A5AFFE1FFC9FE05F9E55C42F98F" box="[483,516,1557,1583]" italics="true" pageId="4" pageNumber="535">Cu</emphasis>
located posteriorly, extending to almost wing apex. Short and apical dichotomous branching vein forming two weakly expressed veins are located closer to the posterior margin of the wing. These veins are the secondary ones apparently (
<emphasis id="B92A3A5AFFE1FFC9FD8DF9AD5CCCF9D7" box="[619,650,1629,1655]" italics="true" pageId="4" pageNumber="535">SV</emphasis>
; after
<bibRefCitation id="EFCF9BB9FFE1FFC9FD33F9AD5D8DF9D8" author="Drake, C. J. &amp; Davis, N. T." box="[725,971,1629,1656]" pageId="4" pageNumber="535" pagination="1 - 100" refId="ref7514" refString="Drake, C. J. &amp; Davis, N. T. (1960) The morphology, phylogeny and higher classification of the family Tingidae, including the description of a new genus and species of the subfamily Vianaidinae (Heteroptera). Entomologia Americana, New Series, 39, 1 - 100." type="journal article" year="1960">Drake &amp; Davis, 1960</bibRefCitation>
). Lateral processes of sternite VII extend beyond posterior margin of sternite VIII, so they are visible when viewing this
<typeStatus id="54E558EAFFE1FFC9FBFAF9715A3AF93B" box="[1052,1148,1665,1691]" pageId="4" pageNumber="535" type="paratype">paratype</typeStatus>
dorsally. Apex of labium reaches hind coxae.
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF71F91D5A19F8EF" blockId="4.[151,1436,1772,2016]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF71F91D5F15F8A7" bold="true" box="[151,339,1772,1799]" pageId="4" pageNumber="535">
<typeStatus id="54E558EAFFE1FFC9FF71F91D5F42F8A7" box="[151,260,1773,1799]" pageId="4" pageNumber="535" type="paratype">Paratype</typeStatus>
15/31(
</emphasis>
female?
<figureCitation id="1365FACDFFE1FFC9FE54F91D5FB3F8A7" box="[434,501,1773,1799]" captionStart="FIGURES 58" captionStartId="6.[151,264,1962,1987]" captionTargetBox="[196,1389,186,1932]" captionTargetId="figure@6.[188,1400,181,1937]" captionTargetPageId="6" captionText="FIGURES 58. Tingiometra pankowskii sp. nov., 5, paratype female (16-03), dorsal view, 6, paratype female (15/27), dorsal view, 7, paratype female (?) (15/31), dorsal view, 8, paratype male (?) (17/06), dorsal view. Scales: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3774906" httpUri="https://zenodo.org/record/3774906/files/figure.png" pageId="4" pageNumber="535">Fig. 7</figureCitation>
). Body glabrous, 2.42 times as long as wide. Subcostal area of hemelytra divided by 6 transverse carinate veins. Discoidal area with about 18 rows of areolae at widest part, sutural area with 6 rows of areolae at the apex, clavus with 7 rows of areolae. Apex of labium reaches hind coxae.
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF71F8A95FB0F8D3" blockId="4.[151,1436,1772,2016]" box="[151,502,1880,1907]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF71F8A95F0DF8D3" bold="true" box="[151,331,1880,1907]" pageId="4" pageNumber="535">
<typeStatus id="54E558EAFFE1FFC9FF71F8A95F42F8D3" box="[151,260,1881,1907]" pageId="4" pageNumber="535" type="paratype">Paratype</typeStatus>
17/06
</emphasis>
(male?
<figureCitation id="1365FACDFFE1FFC9FE43F8A95FAEF8D3" box="[421,488,1881,1907]" captionStart="FIGURES 58" captionStartId="6.[151,264,1962,1987]" captionTargetBox="[196,1389,186,1932]" captionTargetId="figure@6.[188,1400,181,1937]" captionTargetPageId="6" captionText="FIGURES 58. Tingiometra pankowskii sp. nov., 5, paratype female (16-03), dorsal view, 6, paratype female (15/27), dorsal view, 7, paratype female (?) (15/31), dorsal view, 8, paratype male (?) (17/06), dorsal view. Scales: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3774906" httpUri="https://zenodo.org/record/3774906/files/figure.png" pageId="4" pageNumber="535">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BE1E648FFE1FFC9FF21F88D5D78F840" blockId="4.[151,1436,1772,2016]" pageId="4" pageNumber="535">
<emphasis id="B92A3A5AFFE1FFC9FF21F88D5F4DF837" box="[199,267,1917,1943]" italics="true" pageId="4" pageNumber="535">Head.</emphasis>
Crest of clypeus rather wide, postocular lobes clearly tapering toward base. Posterior margin of head with two lateral and one medial elongate process pointed posteriorly. Visible ocellum near posterior margin of right eye (it is possible, however, that we are dealing with an artifact).
</paragraph>
<paragraph id="8BE1E648FFE0FFC8FF21FF6A5C86FF13" blockId="5.[151,1437,153,359]" box="[199,704,153,179]" pageId="5" pageNumber="536">
<emphasis id="B92A3A5AFFE0FFC8FF21FF6A5F70FF13" box="[199,310,154,179]" italics="true" pageId="5" pageNumber="536">Pronotum</emphasis>
1.47 times as wide at base as long.
</paragraph>
<paragraph id="8BE1E648FFE0FFC8FF21FF4D5F26FEBF" blockId="5.[151,1437,153,359]" pageId="5" pageNumber="536">
<emphasis id="B92A3A5AFFE0FFC8FF21FF4D5F03FF77" box="[199,325,189,215]" italics="true" pageId="5" pageNumber="536">Hemelytra.</emphasis>
Subcostal area with 6 rows of areolae at widest part, divided by 4 transverse carinate veins. Discoidal area with about 18 rows of areolae at widest part. Sutural area with 6 rows of areolae at widest part. Clavus with 7 rows of areolae.
</paragraph>
<paragraph id="8BE1E648FFE0FFC8FF21FED95C4AFEC7" blockId="5.[151,1437,153,359]" pageId="5" pageNumber="536">
<emphasis id="B92A3A5AFFE0FFC8FF21FED95F57FEE3" bold="true" box="[199,273,297,323]" pageId="5" pageNumber="536">Notes.</emphasis>
Legs of all
<typeStatus id="54E558EAFFE0FFC8FE75FED95FB8FEE3" box="[403,510,297,323]" pageId="5" pageNumber="536" type="paratype">paratypes</typeStatus>
are long and slender, femora and tibiae cylindrical; tarsi two-segmented, claws thin and long, pulvilli not discernible.
</paragraph>
<paragraph id="8BE1E648FFE0FFC8FF71FE655DA1FE0F" blockId="5.[151,1437,405,791]" box="[151,999,405,431]" pageId="5" pageNumber="536">
<emphasis id="B92A3A5AFFE0FFC8FF71FE655F1DFE0F" bold="true" box="[151,347,405,431]" pageId="5" pageNumber="536">
<typeStatus id="54E558EAFFE0FFC8FF71FE655F42FE0F" box="[151,260,405,431]" pageId="5" pageNumber="536" type="paratype">Paratype</typeStatus>
7303-1
</emphasis>
(
<figureCitation id="1365FACDFFE0FFC8FE8CFE655FA4FE0F" box="[362,482,405,431]" captionStart="FIGURES 912. 9,10" captionStartId="8.[151,264,1940,1965]" captionTargetBox="[207,1380,189,1905]" captionTargetId="figure@8.[194,1393,181,1915]" captionTargetPageId="8" captionText="FIGURES 912. 9,10 Tingiometra pankowskii sp. nov., 9, paratype male (?) (7303-1), dorsal view, 10, paratype male (?) (7303-1), ventral view; 11, Tingiometra secunda sp. nov., holotype female (16/19-2), dorsal view; 12, Tingiometra yuripopovi sp. nov., holotype female (18/25), dorsal view. Scales 1mm." figureDoi="http://doi.org/10.5281/zenodo.3774908" httpUri="https://zenodo.org/record/3774908/files/figure.png" pageId="5" pageNumber="536">Figs. 9, 10</figureCitation>
) does not have any different specific features.
</paragraph>
<paragraph id="8BE1E648FFE0FFC8FF21FE495F95FCB7" blockId="5.[151,1437,405,791]" pageId="5" pageNumber="536">
<emphasis id="B92A3A5AFFE0FFC8FF21FE495C55FE73" bold="true" box="[199,531,440,467]" pageId="5" pageNumber="536">
Measurements of
<typeStatus id="54E558EAFFE0FFC8FE7BFE495C55FE73" box="[413,531,441,467]" pageId="5" pageNumber="536" type="paratype">paratypes</typeStatus>
</emphasis>
(in mm). Body length to apex of hemelytra 3.10 (
<typeStatus id="54E558EAFFE0FFC8FBAEFE495AF3FE73" box="[1096,1205,441,467]" pageId="5" pageNumber="536" type="paratype">paratypes</typeStatus>
16/03, 15/31); body width with wings folded in repose 0.90 (
<typeStatus id="54E558EAFFE0FFC8FDBBFE2D5CF9FE57" box="[605,703,477,503]" pageId="5" pageNumber="536" type="paratype">paratype</typeStatus>
17/06), 1.28 (
<typeStatus id="54E558EAFFE0FFC8FCB8FE2D5D86FE57" box="[862,960,477,503]" pageId="5" pageNumber="536" type="paratype">paratype</typeStatus>
15/31); 2.42 times as long as width (
<typeStatus id="54E558EAFFE0FFC8FA87FE2D5E81FDBB" pageId="5" pageNumber="536" type="paratype">paratype</typeStatus>
15/31); head length from apex of clypeus to posterior margin of eye 0.500.54; head length including postocular portion 0.730.80; length of anteocular portion 0.300.43; length of eye 0.100.14; length of postocular portion 0.230.25; head width across eyes 0.330.52; width of anteocular part 0.130.175, width of postocular part 0.250.34; length of antenna 1.56 (
<typeStatus id="54E558EAFFE0FFC8FDFAFD9D5C38FD27" box="[540,638,621,647]" pageId="5" pageNumber="536" type="paratype">paratype</typeStatus>
16/03); length of antennal segments (I, II, III, IV) 0.13, 0.71, 0.28, 0.45 (
<typeStatus id="54E558EAFFE0FFC8FF79FD615F47FD0B" box="[159,257,657,683]" pageId="5" pageNumber="536" type="paratype">paratype</typeStatus>
16/03); pronotum length 0.430.50; pronotum width 0.630.70; scutellum length 0.280.38; scutellum width 0.290.45; hemelytra length 1.782.10; length of commissure between clavus and corium 0.700.80, femora length: fore 0.700.75, middle 0.700.75, hind 0.981.13; tibiae length: fore 1.201.25, middle 1.001.20, hind 1.381.50; tarsi length 0.13.
</paragraph>
</subSubSection>
<subSubSection id="C344B5C3FFE0FFC8FF21FCB55A39FC23" pageId="5" pageNumber="536" type="etymology">
<paragraph id="8BE1E648FFE0FFC8FF21FCB55A39FC23" blockId="5.[151,1436,837,935]" pageId="5" pageNumber="536">
<emphasis id="B92A3A5AFFE0FFC8FF21FCB55F14FCFF" bold="true" box="[199,338,837,863]" pageId="5" pageNumber="536">Etymology:</emphasis>
This species is dedicated to Mark Pankowski (Rockville,
<collectingRegion id="499A28AAFFE0FFC8FC36FCB55A06FCFF" box="[976,1088,837,863]" country="United States of America" name="Maryland" pageId="5" pageNumber="536">Maryland</collectingRegion>
,
<collectingCountry id="F349A6D8FFE0FFC8FBAFFCB55AC3FCFF" box="[1097,1157,837,863]" name="United States of America" pageId="5" pageNumber="536">USA</collectingCountry>
), who acquired the inclusion of
<typeStatus id="54E558EAFFE0FFC8FF0DFC995F0DFC23" box="[235,331,873,899]" pageId="5" pageNumber="536" type="paratype">paratype</typeStatus>
15/27 and generously donated it to the second author for the description.
</paragraph>
</subSubSection>
<subSubSection id="C344B5C3FFE0FFC8FF21FC7C5CB8FC07" box="[199,766,908,935]" pageId="5" pageNumber="536" type="distribution">
<paragraph id="8BE1E648FFE0FFC8FF21FC7C5CB8FC07" blockId="5.[151,1436,837,935]" box="[199,766,908,935]" pageId="5" pageNumber="536">
<emphasis id="B92A3A5AFFE0FFC8FF21FC7C5F27FC07" bold="true" box="[199,353,908,935]" pageId="5" pageNumber="536">Distribution.</emphasis>
Burmese amber, northern
<collectingCountry id="F349A6D8FFE0FFC8FD6DFC7D5CBCFC07" box="[651,762,909,935]" name="Myanmar" pageId="5" pageNumber="536">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>