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<document id="4896C3375FD85E4D927782388F7222F5" ID-DOI="10.21248/contrib.entomol.67.2.275-318" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742497" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638268338673" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2017" docId="03F6C753FFE3FFA4FF06DC26FDDBFE04" docLanguage="en" docName="ContribEntomol.67.2.275-318.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Eleusis ecuadoriensis Irmler 2017, spec. nov." docType="treatment" docUuid="774F9A2D-25FE-426C-8A32-C52AB801876B" docUuidSource="ZooBank" docVersion="4" lastPageNumber="282" masterDocId="FFCFBF2BFFE5FFA3FFFFD92CFFDAFF9C" masterDocTitle="The Neotropical species of the genus Eleusis LAPORTE, 1835 with description of the new genus Eleusima (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="318" masterPageNumber="275" pageNumber="281" updateTime="1699437037312" updateUser="plazi">
<mods:mods id="4E0B549C828B81AEFB6D0AD7D8725D8E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BEBB59BD9957529C6CCB5BB2FE15E858">
<mods:title id="46DCCD7950B947A56F34FECD876398C1">The Neotropical species of the genus Eleusis LAPORTE, 1835 with description of the new genus Eleusima (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
</mods:titleInfo>
<mods:name id="1465F21AEFD24319FC8F328DB0C8C4F3" type="personal">
<mods:role id="D342B5E3EFE1B718DE433C8CB2CC407E">
<mods:roleTerm id="655430CAC1DBE256A9BEF92ED48D3551">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="29C815B3968D3886A4EB8DD4D181F079">Irmler, Ulrich</mods:namePart>
</mods:name>
<mods:typeOfResource id="D58322D775C4BC80AE712901D6236176">text</mods:typeOfResource>
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<mods:title id="9255955CA5CEAB2BD7861BF4BF0524AB">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="DADF2D60A5953E5D2DB654B4E04A3B69">
<mods:date id="F8DE9B685D8B05C42B643BDFE6812278">2017</mods:date>
<mods:detail id="F525C9BE104DD4D9462BA54AF3ABD5AC" type="volume">
<mods:number id="EB8A8D57D3BF37DF5F4E618A98BFF3F8">67</mods:number>
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<mods:classification id="E1D329998DAD4F09DD228826DA4AF5A6">journal article</mods:classification>
<mods:identifier id="F49ACFCDFB512DD02B59D43F433758D1" type="DOI">10.21248/contrib.entomol.67.2.275-318</mods:identifier>
<mods:identifier id="DA4CA9A88FFE7A95F53BB2BF78B5052E" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="6E473467C98BAEC654C57193A8587716" type="Zenodo-Dep">5742497</mods:identifier>
<mods:identifier id="5686342D169B60EB468EFB5F99C38720" type="ZooBank">98BC4B71-EE31-4BA0-855A-AAC48C955B38</mods:identifier>
</mods:mods>
<treatment id="03F6C753FFE3FFA4FF06DC26FDDBFE04" LSID="urn:lsid:zoobank.org:act:774F9A2D-25FE-426C-8A32-C52AB801876B" httpUri="http://treatment.plazi.org/id/03F6C753FFE3FFA4FF06DC26FDDBFE04" lastPageId="7" lastPageNumber="282" pageId="6" pageNumber="281">
<subSubSection id="C34525CEFFE3FFA5FF06DC26FCD8FADA" pageId="6" pageNumber="281" type="nomenclature">
<paragraph id="8BE07645FFE3FFA5FF06DC26FD9EFAB9" blockId="6.[249,770,1290,1382]" box="[249,580,1290,1317]" pageId="6" pageNumber="281">
<heading id="D0A8C129FFE3FFA5FF06DC26FD9EFAB9" box="[249,580,1290,1317]" fontSize="12" level="2" pageId="6" pageNumber="281" reason="2">
<taxonomicName id="4C5F0DC6FFE3FFA5FF06DC26FE0EFAB9" ID-CoL="398N5" authority="Irmler, 2017" authorityName="Irmler" authorityYear="2017" box="[249,468,1290,1317]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="ecuadoriensis" status="sp. nov.">
<emphasis id="B92BAA57FFE3FFA5FF06DC26FE0EFAB9" box="[249,468,1290,1317]" italics="true" pageId="6" pageNumber="281">Eleusis ecuadoriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218172CFFE3FFA5FE24DC26FD9EFAB9" box="[475,580,1290,1317]" pageId="6" pageNumber="281" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BE07645FFE3FFA5FF06DC1EFCD8FADA" blockId="6.[249,770,1290,1382]" box="[249,770,1330,1350]" pageId="6" pageNumber="281">
<uri id="FFCE7A47FFE3FFA5FF06DC1EFCD8FADA" box="[249,770,1330,1350]" pageId="6" pageNumber="281">
urn:lsid:zoobank.org:act:
<uuid id="FFF94C90FFE3FFA5FE57DC1EFCD8FADA" box="[424,770,1330,1350]" pageId="6" pageNumber="281">774F9A2D-25FE-426C-8A32-C52AB801876B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFE3FFA5FF06DC7EFEAAFAFA" box="[249,368,1362,1382]" pageId="6" pageNumber="281" type="description">
<paragraph id="8BE07645FFE3FFA5FF06DC7EFEAAFAFA" blockId="6.[249,770,1290,1382]" box="[249,368,1362,1382]" pageId="6" pageNumber="281">
<figureCitation id="13646AC0FFE3FFA5FF06DC7EFEF4FAFA" box="[249,302,1362,1382]" captionStart="Fig" captionStartId="33.[174,207,2052,2074]" captionTargetBox="[204,1383,483,2021]" captionTargetId="figure-103@33.[174,1421,445,2051]" captionTargetPageId="33" captionText="Fig. 13–17: Eleusis rufipennis (13); E. ecuadoriensis (14), Eleusis breviceps (15), E. insignis (16), E. scita (17); aedeagus in lateral (A)" figureDoi="http://doi.org/10.5281/zenodo.5742505" httpUri="https://zenodo.org/record/5742505/files/figure.png" pageId="6" pageNumber="281">Figs 14</figureCitation>
A-C,29F
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFE3FFA5FF2BDCACFD94FA64" pageId="6" pageNumber="281" type="materials_examined">
<paragraph id="8BE07645FFE3FFA5FF2BDCACFD94FA64" blockId="6.[212,817,1407,1528]" pageId="6" pageNumber="281">
<emphasis id="B92BAA57FFE3FFA5FF2BDCACFEABFA0B" bold="true" box="[212,369,1407,1432]" pageId="6" pageNumber="281">Type material</emphasis>
:
<materialsCitation id="3B377C18FFE3FFA5FE7BDCACFD94FA64" collectingDate="2009-11-26" collectionCode="BMNH" collectorName="Yanayacu-Rio Perdido Tr. &amp; L. Sekerka &amp; K. Stajerova" country="Ecuador" elevation="2100" latitude="-0.5833333" location="Cosanga" longLatPrecision="1308" longitude="-77.88333" pageId="6" pageNumber="281" specimenCount="1" stateProvince="Napo" typeStatus="holotype">
male,
<typeStatus id="54E4C8E7FFE3FFA5FE35DCACFDEAFA04" box="[458,560,1408,1432]" pageId="6" pageNumber="281" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34836D5FFE3FFA5FDBFDCACFD44FA04" box="[576,670,1408,1432]" name="Ecuador" pageId="6" pageNumber="281">Ecuador</collectingCountry>
:
<collectingRegion id="499BB8A7FFE3FFA5FD52DCACFCF7FA0B" box="[685,813,1408,1431]" country="Ecuador" name="Napo" pageId="6" pageNumber="281">Napo Prov.</collectingRegion>
,
<location id="8E80209EFFE3FFA5FF2BDC8CFEEAFA24" LSID="urn:lsid:plazi:treatment:03F6C753FFE3FFA4FF06DC26FDDBFE04:8E80209EFFE3FFA5FF2BDC8CFEEAFA24" box="[212,304,1440,1464]" country="Ecuador" latitude="-0.5833333" longLatPrecision="1308" longitude="-77.88333" name="Cosanga" pageId="6" pageNumber="281" stateProvince="Napo">Cosanga</location>
env.,
<collectorName id="26AA1393FFE3FFA5FE99DC8CFDABFA2B" box="[358,625,1440,1464]" pageId="6" pageNumber="281">Yanayacu-Rio Perdido Tr.</collectorName>
(
<geoCoordinate id="EE6B1082FFE3FFA5FD84DC8CFD65FA24" box="[635,703,1440,1464]" degrees="0" direction="south" minutes="35" orientation="latitude" pageId="6" pageNumber="281" precision="925" value="-0.5833333">0°35'S</geoCoordinate>
,
<geoCoordinate id="EE6B1082FFE3FFA5FD39DC8CFCFFFA24" box="[710,805,1440,1464]" degrees="77" direction="west" minutes="53" orientation="longitude" pageId="6" pageNumber="281" precision="925" value="-77.88333">77°53'W</geoCoordinate>
),
<quantity id="4CA7DBA0FFE3FFA5FF2BDCECFEF9FA4B" box="[212,291,1472,1496]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="281" unit="m" value="2100.0">
<elevation id="00729176FFE3FFA5FF2BDCECFEF9FA4B" box="[212,291,1472,1496]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="281" unit="m" value="2100.0">2100 m</elevation>
</quantity>
elev., under bark of dead tree,
<date id="FFE15085FFE3FFA5FD77DCECFD27FA44" box="[648,765,1472,1496]" pageId="6" pageNumber="281" value="2009-11-26">
<collectingDate id="EFA5A96DFFE3FFA5FD77DCECFD27FA44" box="[648,765,1472,1496]" pageId="6" pageNumber="281" value="2009-11-26">26.11.2009</collectingDate>
</date>
, leg.
<collectorName id="26AA1393FFE3FFA5FF2BDCCCFE9BFA64" box="[212,321,1504,1528]" pageId="6" pageNumber="281">L. Sekerka</collectorName>
&amp;
<collectorName id="26AA1393FFE3FFA5FE9FDCCCFE3AFA64" box="[352,480,1504,1528]" pageId="6" pageNumber="281">K. Stajerová</collectorName>
(
<collectionCode id="ED4EEE80FFE3FFA5FE12DCCCFD99FA64" box="[493,579,1504,1528]" pageId="6" pageNumber="281">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFE3FFA5FF2BDF33FE52F7A4" pageId="6" pageNumber="281" type="diagnosis">
<paragraph id="8BE07645FFE3FFA5FF2BDF33FE52F7A4" blockId="6.[212,817,1567,2104]" pageId="6" pageNumber="281">
<emphasis id="B92BAA57FFE3FFA5FF2BDF33FE9BF9AB" bold="true" box="[212,321,1567,1591]" pageId="6" pageNumber="281">Diagnosis</emphasis>
:
<taxonomicName id="4C5F0DC6FFE3FFA5FEB4DF0CFE2EF9A4" authorityName="Irmler" authorityYear="2017" box="[331,500,1568,1592]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="ecuadoriensis">
<emphasis id="B92BAA57FFE3FFA5FEB4DF0CFE2EF9A4" box="[331,500,1568,1592]" italics="true" pageId="6" pageNumber="281">E. ecuadoriensis</emphasis>
</taxonomicName>
is still longer than
<taxonomicName id="4C5F0DC6FFE3FFA5FD4ADF0CFCF0F9AB" authorityName="Irmler" authorityYear="2017" box="[693,810,1568,1591]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B92BAA57FFE3FFA5FD4ADF0CFCF0F9AB" box="[693,810,1568,1591]" italics="true" pageId="6" pageNumber="281">E. castanea</emphasis>
</taxonomicName>
, which are both the largest species of the
<taxonomicName id="4C5F0DC6FFE3FFA5FD85DF6CFD38F9C4" baseAuthorityName="ERICHSON" baseAuthorityYear="1840" box="[634,738,1600,1624]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B92BAA57FFE3FFA5FD85DF6CFD38F9C4" box="[634,738,1600,1624]" italics="true" pageId="6" pageNumber="281">E. humilis</emphasis>
</taxonomicName>
-group. It resembles
<taxonomicName id="4C5F0DC6FFE3FFA5FEA0DF4CFE0EF9EB" authorityName="Irmler" authorityYear="2017" box="[351,468,1632,1655]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B92BAA57FFE3FFA5FEA0DF4CFE0EF9EB" box="[351,468,1632,1655]" italics="true" pageId="6" pageNumber="281">E. castanea</emphasis>
</taxonomicName>
not only in size and shape, but also in colouration. The elytra have the same reddishbrown colouration as in
<taxonomicName id="4C5F0DC6FFE3FFA5FE05DF8CFDB5F92B" authorityName="Irmler" authorityYear="2017" box="[506,623,1696,1719]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B92BAA57FFE3FFA5FE05DF8CFDB5F92B" box="[506,623,1696,1719]" italics="true" pageId="6" pageNumber="281">E. castanea</emphasis>
</taxonomicName>
. Punctation and microsculpture are also extremely similar. The antennae of
<taxonomicName id="4C5F0DC6FFE3FFA5FEDEDFCCFE13F964" authorityName="Irmler" authorityYear="2017" box="[289,457,1760,1784]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="ecuadoriensis">
<emphasis id="B92BAA57FFE3FFA5FEDEDFCCFE13F964" box="[289,457,1760,1784]" italics="true" pageId="6" pageNumber="281">E. ecuadoriensis</emphasis>
</taxonomicName>
are slightly longer than those of
<taxonomicName id="4C5F0DC6FFE3FFA5FF2BDE2CFE93F88B" authorityName="Irmler" authorityYear="2017" box="[212,329,1792,1815]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B92BAA57FFE3FFA5FF2BDE2CFE93F88B" box="[212,329,1792,1815]" italics="true" pageId="6" pageNumber="281">E. castanea</emphasis>
</taxonomicName>
. Whereas they are as long as head and pronotum combined in
<taxonomicName id="4C5F0DC6FFE3FFA5FE5ADE0CFD97F8A4" authorityName="Irmler" authorityYear="2017" box="[421,589,1824,1848]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="ecuadoriensis">
<emphasis id="B92BAA57FFE3FFA5FE5ADE0CFD97F8A4" box="[421,589,1824,1848]" italics="true" pageId="6" pageNumber="281">E. ecuadoriensis</emphasis>
</taxonomicName>
, in
<taxonomicName id="4C5F0DC6FFE3FFA5FD7EDE0CFD2CF8AB" authorityName="Irmler" authorityYear="2017" box="[641,758,1824,1847]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B92BAA57FFE3FFA5FD7EDE0CFD2CF8AB" box="[641,758,1824,1847]" italics="true" pageId="6" pageNumber="281">E. castanea</emphasis>
</taxonomicName>
they are only as long as head and half of pronotum combined. Without dissection of the aedeagus the separation of the two species is difficult. In
<taxonomicName id="4C5F0DC6FFE3FFA5FDFADEACFDA0F80B" authorityName="Irmler" authorityYear="2017" box="[517,634,1920,1943]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B92BAA57FFE3FFA5FDFADEACFDA0F80B" box="[517,634,1920,1943]" italics="true" pageId="6" pageNumber="281">E. castanea</emphasis>
</taxonomicName>
the aedeagus is stout, the apical part of the central lobe is short and the transparent lobe of the paramere is distinctly widened. In
<taxonomicName id="4C5F0DC6FFE3FFA5FF2BDECCFEA6F864" authorityName="Irmler" authorityYear="2017" box="[212,380,2016,2040]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="ecuadoriensis">
<emphasis id="B92BAA57FFE3FFA5FF2BDECCFEA6F864" box="[212,380,2016,2040]" italics="true" pageId="6" pageNumber="281">E. ecuadoriensis</emphasis>
</taxonomicName>
, the aedeagus is slenderer, the apical part is longer and the transparent lobe of the paramere is only slightly widened.
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFE3FFA4FCA9D9D3FD9EFEAB" lastPageId="7" lastPageNumber="282" pageId="6" pageNumber="281" type="description">
<paragraph id="8BE07645FFE3FFA5FCA9D9D3FC7EFEC4" blockId="6.[854,1459,255,2104]" pageId="6" pageNumber="281">
<emphasis id="B92BAA57FFE3FFA5FCA9D9D3FC06FE8B" bold="true" box="[854,988,255,279]" pageId="6" pageNumber="281">Description</emphasis>
: Length:
<quantity id="4CA7DBA0FFE3FFA5FB9ED82CFB6DFE8B" box="[1121,1207,256,280]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="6" pageNumber="281" unit="mm" value="6.3">6.3 mm</quantity>
. Colouration: black; elytra reddish-brown; legs light brown; antennae dark brown.
</paragraph>
<paragraph id="8BE07645FFE3FFA5FCA9D84CFB14FD64" blockId="6.[854,1459,255,2104]" pageId="6" pageNumber="281">
Head:
<quantity id="4CA7DBA0FFE3FFA5FC63D84CFC20FEEB" box="[924,1018,352,376]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="6" pageNumber="281" unit="mm" value="0.79">0.79 mm</quantity>
long,
<quantity id="4CA7DBA0FFE3FFA5FBC3D84CFB41FEEB" box="[1084,1179,352,376]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="6" pageNumber="281" unit="mm" value="0.83">0.83 mm</quantity>
wide; eyes slightly prominent; postocular part wider than preocular part; temples approximately twice as long as eyes; nearly parallel; preocular part to base of antennae parallel; anterior margin of clypeus convergent; supraocular groove deep and wide; punctation weak and irregularly dense; on average, interstices between punctures between two to four times as wide as diameter of punctures; setiferous puncture at each end of supraocular groove; another pair of setiferous punctures on each side of prominent central part of clypeus; elongate microsculpture dense, but not deep; transversely striate on clypeus; longitudinally striate on vertex; surface slightly shiny.
</paragraph>
<paragraph id="8BE07645FFE3FFA5FCA9DA2CFB72FB8B" blockId="6.[854,1459,255,2104]" pageId="6" pageNumber="281">Antennae as long as head and pronotum combined; first antennomere slightly thicker than following antennomeres; more than twice as long as second; third antennomere one third longer than second; following antennomeres slightly increasing in width and decreasing in length; fourth to eighth antennomere longer than wide; ninth and tenth antennomere approximately quadrate; fourth to eleventh antennomere pubescent; all antennomeres with apical setae.</paragraph>
<paragraph id="8BE07645FFE3FFA5FCA9DD0CFB48FA44" blockId="6.[854,1459,255,2104]" pageId="6" pageNumber="281">
Pronotum:
<quantity id="4CA7DBA0FFE3FFA5FC2EDD0CFBEAFBAB" box="[977,1072,1056,1080]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="281" unit="mm" value="0.7">0.70 mm</quantity>
long,
<quantity id="4CA7DBA0FFE3FFA5FB8ADD0CFB09FBAB" box="[1141,1235,1056,1080]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="6" pageNumber="281" unit="mm" value="0.86">0.86 mm</quantity>
wide; widest shortly behind anterior angles; sides in posterior two third smoothly narrowed to posterior angles; in posterior third, interrupted by sharp emargination; posterior angles obtuse; short grooves at anterior angles; punctation as dense and deep as on head; two setiferous punctures in each groove at anterior angles; additional setiferous punctures along lateral margin; lateral interruption marked by setiferous puncture in moderately deep impression; at anterior margin close to anterior angles, with impressions and setiferous punctures on each side of midline; elongate microsculpture as deep and dense as on head; circularly striate at anterior angles; longitudinally striate on disc; surface slightly shiny.
</paragraph>
<paragraph id="8BE07645FFE3FFA5FCA9DCCCFBE0F944" blockId="6.[854,1459,255,2104]" pageId="6" pageNumber="281">
Elytra:
<quantity id="4CA7DBA0FFE3FFA5FC55DCCCFBD2FA6B" box="[938,1032,1504,1528]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" pageId="6" pageNumber="281" unit="mm" value="1.07">1.07 mm</quantity>
long,
<quantity id="4CA7DBA0FFE3FFA5FBA7DCCCFB6CFA6B" box="[1112,1206,1504,1528]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" pageId="6" pageNumber="281" unit="mm" value="1.01">1.01 mm</quantity>
wide; humeral angels nearly rectangular; sides slightly divergent to posterior angles; posterior angles rounded; punctation and microsculpture as dense and deep as on head and pronotum; elongate microsculpture longitudinally striate; one setiferous puncture in central position on each side of suture; several setiferous punctures along lateral margin; surface slightly shiny.
</paragraph>
<paragraph id="8BE07645FFE3FFA5FCA9DFCCFBD7F8A4" blockId="6.[854,1459,255,2104]" pageId="6" pageNumber="281">Abdomen weakly punctate; on each segment, with row of setiferous punctures; isodiametric microsculpture moderately deep.</paragraph>
<paragraph id="8BE07645FFE3FFA4FCA9DE6CFD9EFEAB" blockId="6.[854,1459,255,2104]" lastBlockId="7.[174,779,256,312]" lastPageId="7" lastPageNumber="282" pageId="6" pageNumber="281">Aedeagus slender; in lateral aspect, continuously narrowed to nearly acute apex; in ventral aspect, parallel with triangular apex; in central position of central lobe with large approximately triangular prominence; few sensillae at apex; several sensillae at central prominence; parameres slightly shorter than central lobe; slender; apical transparent lobe nearly as wide as basal scape; long seta at apex of transparent lobe; without short setose fringe; several minute sensillae close to transparent lobe and at base of scape.</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFE2FFA4FF51D84CFDDBFE04" pageId="7" pageNumber="282" type="etymology">
<paragraph id="8BE07645FFE2FFA4FF51D84CFCD1FEE4" blockId="7.[174,779,352,408]" box="[174,779,352,376]" pageId="7" pageNumber="282">
<emphasis id="B92BAA57FFE2FFA4FF51D84CFEFFFEE4" bold="true" box="[174,293,352,376]" pageId="7" pageNumber="282">Etymology</emphasis>
: The species name derived from the country
</paragraph>
<paragraph id="8BE07645FFE2FFA4FF51D8ACFDDBFE04" blockId="7.[174,779,352,408]" box="[174,513,384,408]" pageId="7" pageNumber="282">
<collectingCountry id="F34836D5FFE2FFA4FF51D8ACFED3FE04" box="[174,265,384,408]" name="Ecuador" pageId="7" pageNumber="282">Ecuador</collectingCountry>
, where it was collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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