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<document id="7A21FC33B9851E785911C9C25737EBDB" ID-DOI="10.5281/zenodo.5392847" ID-ISSN="1638-9387" ID-Zenodo-Dep="5392847" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630317447715" checkinUser="marcus" docAuthor="Mah, Christopher" docDate="2005" docId="03F287F6FFFAFF9D5D11B6ABFD98F974" docLanguage="en" docName="Zoosystema.2005.27.1.137-161.pdf" docOrigin="Zoosystema 27 (1)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Glyphodiscus pentagonalis Mah 2005, n. sp." docType="treatment" docVersion="3" lastPageNumber="150" masterDocId="FFCBFF8EFFF1FF905E37B559FFACFFFF" masterDocTitle="A phylogeny of Iconaster and Glyphodiscus (Echinodermata, Asteroidea, Valvatida, Goniasteridae) with descriptions of four new species" masterLastPageNumber="161" masterPageNumber="137" pageNumber="148" updateTime="1699224777309" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="99F24DC540C6B1BEA5C002C59A3920B5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9E1F582FEF6829F59E3D78C13D97FE3D">
<mods:title id="963B43764F854DA310813B09B69F040B">A phylogeny of Iconaster and Glyphodiscus (Echinodermata, Asteroidea, Valvatida, Goniasteridae) with descriptions of four new species</mods:title>
</mods:titleInfo>
<mods:name id="8D08AF37809C979773090C21FD88A744" type="personal">
<mods:role id="BEA005549F4F25C022E021A9111BDFA4">
<mods:roleTerm id="0FA71D2F5AB7AF4DBF5C138D3EAF7339">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="621271140980E12F8B83185D09148F93">Mah, Christopher</mods:namePart>
<mods:affiliation id="FE26E71EAE153302B1A80BC580CEB701">Department of Geology, University of Illinois at Urbana-Champaign, 245 NHB, 1301 W. Green, Urbana, Illinois 61801 (USA) mah @ uiuc. edu.</mods:affiliation>
<mods:nameIdentifier id="D45B4038317ED969783FA16AFC5D8F95" type="email">mah@uiuc.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7336390FBCA7365EB0E5739A83785102">text</mods:typeOfResource>
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<mods:titleInfo id="F9F7BCD56B7EBA6E4E6156549F685232">
<mods:title id="9D8C7E884A261DE01940BA2F2E541ED3">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="33F6B0298312CAC28DF776A876EC4BDB">
<mods:date id="C91576C937E02811EA9CE342CD90D330">2005</mods:date>
<mods:detail id="4857F713D7536103E5A76746C647D714" type="volume">
<mods:number id="2F47B8BB4BE7135961B265B98C4B660D">27</mods:number>
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<mods:detail id="AD23E7B6D07993FA95911DF75D202AEF" type="issue">
<mods:number id="BAAA0A4B1CFBC766661B2FB2A86CF760">1</mods:number>
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<mods:extent id="1BB5CC5D0D9FD709395ACE5189F1D656" unit="page">
<mods:start id="2A4E718B1EFF7F781CF474DBCF76BB5B">137</mods:start>
<mods:end id="C15058EF1A174F1EB3EF19008078F2A2">161</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="9AD1BEAE131CEFEE023E70E0E9E02D39">journal article</mods:classification>
<mods:identifier id="FCD690A46E908213D6ADDCA0843263E0" type="DOI">10.5281/zenodo.5392847</mods:identifier>
<mods:identifier id="9D0EBCC8F81A20C04614E9F6A6E7D247" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="3AA5D92236F3E58764F03531DFD6FBD1" type="Zenodo-Dep">5392847</mods:identifier>
</mods:mods>
<treatment id="03F287F6FFFAFF9D5D11B6ABFD98F974" LSID="urn:lsid:plazi:treatment:03F287F6FFFAFF9D5D11B6ABFD98F974" httpUri="http://treatment.plazi.org/id/03F287F6FFFAFF9D5D11B6ABFD98F974" lastPageId="13" lastPageNumber="150" pageId="11" pageNumber="148">
<subSubSection id="C341656BFFFAFF9B5D11B6ABFB2FFBF4" box="[806,1155,1010,1035]" pageId="11" pageNumber="148" type="nomenclature">
<paragraph id="8BE436E0FFFAFF9B5D11B6ABFB2FFBF4" blockId="11.[714,1247,1010,1713]" box="[806,1155,1010,1035]" pageId="11" pageNumber="148">
<heading id="D0AC818CFFFAFF9B5D11B6ABFB2FFBF4" box="[806,1155,1010,1035]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="148" reason="2">
<taxonomicName id="4C5B4D63FFFAFF9B5D11B6ABFBEDFBF4" ID-CoL="3GGSN" authority="Mah, 2005" authorityName="Mah" authorityYear="2005" box="[806,1089,1010,1035]" class="Asteroidea" family="Goniasteridae" genus="Glyphodiscus" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="148" phylum="Echinodermata" rank="species" species="pentagonalis" status="sp. nov.">
<emphasis id="B92FEAF2FFFAFF9B5D11B6ABFBEDFBF4" bold="true" box="[806,1089,1010,1035]" italics="true" pageId="11" pageNumber="148">Glyphodiscus pentagonalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C5789FFFAFF9B5A7FB6ABFB2FFBF4" box="[1096,1155,1010,1035]" pageId="11" pageNumber="148" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C341656BFFFAFF9B5DA5B14BFBBBFBD4" box="[914,1047,1042,1067]" pageId="11" pageNumber="148" type="description">
<paragraph id="8BE436E0FFFAFF9B5DA5B14BFBBBFBD4" blockId="11.[714,1247,1010,1713]" box="[914,1047,1042,1067]" pageId="11" pageNumber="148">
(
<figureCitation id="13602A65FFFAFF9B5DADB14BFBA3FBD4" box="[922,1039,1042,1067]" captionStart="FIG" captionStartId="12.[151,162,1660,1677]" captionTargetBox="[151,1245,368,1606]" captionTargetId="figure-30@12.[145,763,966,1606]" captionTargetPageId="12" captionText="FIG. 3. — A-C, Glyphodiscus magnificus n. sp., New Caledonia, paratype, dry (MNHN EcAs 11682); A, abactinal view; B, superomarginal plate shape; C, oral and adambulacral regions; D, E, Glyphodiscus pentagonalis n. sp., New Caledonia, holotype, dry (MNHN EcAs 11688); D, abactinal view; E, oral and adambulacral regions. Scale bars: A, 5.0 mm; B, D, 3.0 mm; C, 2.0 mm;" figureDoi="http://doi.org/10.5281/zenodo.5392853" httpUri="https://zenodo.org/record/5392853/files/figure.png" pageId="11" pageNumber="148">Fig. 3D, E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C341656BFFFAFF9B5CFDB165FC1DF9BB" pageId="11" pageNumber="148" type="materials_examined">
<paragraph id="8BE436E0FFFAFF9B5CFDB165FB34FB60" blockId="11.[714,1247,1010,1713]" pageId="11" pageNumber="148">
<materialsCitation id="3B333CBDFFFAFF9B5CFDB165FB34FB60" collectingDate="1993-01-27" collectionCode="R, MNHN" elevation="518" latitude="-24.92" location="South Pacific Ocean." longLatPrecision="124" longitude="168.36166" municipality="Eponge" pageId="11" pageNumber="148" specimenCode="SMIB 8" specimenCount="1" typeStatus="holotype">
<typeStatus id="54E08842FFFAFF9B5CFDB165FC9BFBAD" box="[714,823,1084,1107]" pageId="11" pageNumber="148" type="holotype">HOLOTYPE</typeStatus>
. —
<emphasis id="B92FEAF2FFFAFF9B5D5FB165FBE9FBAC" bold="true" box="[872,1093,1084,1107]" pageId="11" pageNumber="148">
<location id="8E84603BFFFAFF9B5D5FB165FBE9FBAC" LSID="urn:lsid:plazi:treatment:03F287F6FFFAFF9D5D11B6ABFD98F974:8E84603BFFFAFF9B5D5FB165FBE9FBAC" box="[872,1093,1084,1107]" latitude="-24.92" longLatPrecision="124" longitude="168.36166" municipality="Eponge" name="South Pacific Ocean." pageId="11" pageNumber="148">South Pacific Ocean.</location>
</emphasis>
<specimenCode id="DBFD9E9BFFFAFF9B5A79B165FB0DFBAC" box="[1102,1185,1084,1107]" collectionCode="SMIB" pageId="11" pageNumber="148">SMIB 8</specimenCode>
, banc
<collectingMunicipality id="6B80AC9AFFFAFF9B5CFDB10FFCB2FB92" box="[714,798,1110,1133]" pageId="11" pageNumber="148">Éponge</collectingMunicipality>
(mont B), stn DW 146,
<geoCoordinate id="EE6F5027FFFAFF9B5A50B10FFB7AFB92" box="[1127,1238,1110,1133]" degrees="24" direction="south" minutes="55. 2" orientation="latitude" pageId="11" pageNumber="148" precision="92" value="-24.92">24°55.2’S</geoCoordinate>
,
<geoCoordinate id="EE6F5027FFFAFF9B5CFDB136FC92FB79" box="[714,830,1135,1158]" degrees="168" direction="east" minutes="21.7" orientation="longitude" pageId="11" pageNumber="148" precision="92" value="168.36166">168°21.7’E</geoCoordinate>
,
<quantity id="4CA39B05FFFAFF9B5D7CB136FC13FB79" box="[843,959,1135,1158]" metricMagnitude="2" metricUnit="m" metricValue="5.18" metricValueMax="5.22" metricValueMin="5.14" pageId="11" pageNumber="148" unit="m" value="518.0" valueMax="522.0" valueMin="514.0">
<elevation id="0076D1D3FFFAFF9B5D7CB136FC13FB79" box="[843,959,1135,1158]" metricMagnitude="2" metricUnit="m" metricValue="5.18" metricValueMax="5.22" metricValueMin="5.14" pageId="11" pageNumber="148" unit="m" value="518.0" valueMax="522.0" valueMin="514.0">514-522 m</elevation>
</quantity>
,
<date id="FFE51020FFFAFF9B5DFCB136FB9DFB79" box="[971,1073,1135,1158]" pageId="11" pageNumber="148" value="1993-01-27">
<collectingDate id="EFA1E9C8FFFAFF9B5DFCB136FB9DFB79" box="[971,1073,1135,1158]" pageId="11" pageNumber="148" value="1993-01-27">27.I.1993</collectingDate>
</date>
,
<specimenCount id="9D5DFD69FFFAFF9B5A0AB136FB02FB79" box="[1085,1198,1135,1158]" pageId="11" pageNumber="148" type="generic">1 specimen</specimenCount>
dry,
<collectionCode id="ED4AAE25FFFAFF9B5CFDB1D1FD76FB60" box="[714,730,1160,1183]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="148">R</collectionCode>
=
<quantity id="4CA39B05FFFAFF9B5CC3B1D1FC97FB60" box="[756,827,1160,1183]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="148" unit="cm" value="1.6">1.6 cm</quantity>
, r =
<quantity id="4CA39B05FFFAFF9B5D51B1D1FC06FB60" box="[870,938,1160,1183]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="148" unit="cm" value="1.3">1.3 cm</quantity>
(
<collectionCode id="ED4AAE25FFFAFF9B5D8FB1D1FBA0FB60" box="[952,1036,1160,1183]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="148">MNHN</collectionCode>
EcAs 11688).
</materialsCitation>
</paragraph>
<paragraph id="8BE436E0FFFAFF9B5CFDB1E8FC1DF9BB" blockId="11.[714,1247,1010,1713]" pageId="11" pageNumber="148">
<typeStatus id="54E08842FFFAFF9B5CFDB1E8FC91FB39" box="[714,829,1201,1224]" pageId="11" pageNumber="148" type="paratype">PARATYPES</typeStatus>
. —
<emphasis id="B92FEAF2FFFAFF9B5D45B1E8FB8BFB37" bold="true" box="[882,1063,1201,1224]" pageId="11" pageNumber="148">
<collectingCountry id="F34C7670FFFAFF9B5D45B1E8FBB3FB37" box="[882,1055,1201,1224]" name="New Caledonia" pageId="11" pageNumber="148">New Caledonia</collectingCountry>
.
</emphasis>
LITHIST, banc Stylaster, stn DW 01,
<geoCoordinate id="EE6F5027FFFAFF9B5D9DB193FBA1FB1E" box="[938,1037,1226,1249]" degrees="23" direction="south" minutes="37.4" orientation="latitude" pageId="11" pageNumber="148" precision="92" value="-23.623333">23°37.4’S</geoCoordinate>
,
<geoCoordinate id="EE6F5027FFFAFF9B5A2EB193FB27FB1E" box="[1049,1163,1226,1249]" degrees="167" direction="east" minutes="42.1" orientation="longitude" pageId="11" pageNumber="148" precision="92" value="167.70166">167°42.1’E</geoCoordinate>
,
<quantity id="4CA39B05FFFAFF9B5AA0B193FB77FB1E" box="[1175,1243,1226,1249]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="11" pageNumber="148" unit="m" value="440.0">440 m</quantity>
,
<date id="FFE51020FFFAFF9B5CFDB1BDFCFCFB04" box="[714,848,1252,1275]" pageId="11" pageNumber="148" value="1999-08-10">10.VIII.1999</date>
,
<specimenCount id="9D5DFD69FFFAFF9B5D6DB1BDFC65FB04" box="[858,969,1252,1275]" pageId="11" pageNumber="148" type="generic">1 specimen</specimenCount>
dry,
<collectionCode id="ED4AAE25FFFAFF9B5DCAB1BDFBA1FB04" box="[1021,1037,1252,1275]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="148">R</collectionCode>
=
<quantity id="4CA39B05FFFAFF9B5A10B1BDFBC2FB04" box="[1063,1134,1252,1275]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="148" unit="cm" value="2.2">2.2 cm</quantity>
, r =
<quantity id="4CA39B05FFFAFF9B5AADB1BDFB73FB04" box="[1178,1247,1252,1275]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="148" unit="cm" value="1.5">1.5 cm</quantity>
(
<collectionCode id="ED4AAE25FFFAFF9B5CE4B1A4FC86FAEB" box="[723,810,1277,1300]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="148">MNHN</collectionCode>
EcAs 11690). —
<collectionCode id="ED4AAE25FFFAFF9B5DC2B1A4FB98FAEB" box="[1013,1076,1277,1300]" pageId="11" pageNumber="148">SMIB</collectionCode>
3, stn DW 05,
<geoCoordinate id="EE6F5027FFFAFF9B5CFDB04FFCB7FAD2" box="[714,795,1302,1325]" degrees="24" direction="south" minutes="55" orientation="latitude" pageId="11" pageNumber="148" precision="925" value="-24.916666">24°55’S</geoCoordinate>
,
<geoCoordinate id="EE6F5027FFFAFF9B5D1FB04FFC25FAD2" box="[808,905,1302,1325]" degrees="168" direction="east" minutes="22" orientation="longitude" pageId="11" pageNumber="148" precision="925" value="168.36667">168°22’E</geoCoordinate>
,
<quantity id="4CA39B05FFFAFF9B5DA2B04FFBA5FAD2" box="[917,1033,1302,1325]" metricMagnitude="2" metricUnit="m" metricValue="5.07" metricValueMax="5.12" metricValueMin="5.02" pageId="11" pageNumber="148" unit="m" value="507.0" valueMax="512.0" valueMin="502.0">502-512 m</quantity>
,
<date id="FFE51020FFFAFF9B5A21B04FFB28FAD2" box="[1046,1156,1302,1325]" pageId="11" pageNumber="148" value="1987-05-21">
21.
<collectionCode id="ED4AAE25FFFAFF9B5A03B04FFBE5FAD2" box="[1076,1097,1302,1325]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="148" type="Museum">V</collectionCode>
.1987
</date>
,
<specimenCount id="9D5DFD69FFFAFF9B5AA6B04FFD54FAB8" pageId="11" pageNumber="148" type="generic">1 specimen</specimenCount>
dry,
<collectionCode id="ED4AAE25FFFAFF9B5D01B069FCEAFAB8" box="[822,838,1328,1351]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="148">R</collectionCode>
=
<quantity id="4CA39B05FFFAFF9B5D51B069FC1DFAB8" box="[870,945,1328,1351]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="148" unit="cm" value="1.6">1.6 cm</quantity>
, r =
<quantity id="4CA39B05FFFAFF9B5DDCB069FB9AFAB8" box="[1003,1078,1328,1351]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="148" unit="cm" value="1.3">1.3 cm</quantity>
(
<collectionCode id="ED4AAE25FFFAFF9B5A7EB069FB0CFAB8" box="[1097,1184,1328,1351]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="148">MNHN</collectionCode>
EcAs 11689). —
<collectionCode id="ED4AAE25FFFAFF9B5D62B010FC06FA9F" box="[853,938,1353,1376]" pageId="11" pageNumber="148">BERYX</collectionCode>
11, stn DW 27,
<geoCoordinate id="EE6F5027FFFAFF9B5A4AB010FB7AFA9F" box="[1149,1238,1353,1376]" degrees="23" direction="south" minutes="37" orientation="latitude" pageId="11" pageNumber="148" precision="925" value="-23.616667">23°37’S</geoCoordinate>
,
<geoCoordinate id="EE6F5027FFFAFF9B5CFDB03BFC81FA86" box="[714,813,1378,1401]" degrees="167" direction="east" minutes="41" orientation="longitude" pageId="11" pageNumber="148" precision="925" value="167.68333">167°41’E</geoCoordinate>
,
<quantity id="4CA39B05FFFAFF9B5D0DB03BFC1DFA86" box="[826,945,1378,1401]" metricMagnitude="2" metricUnit="m" metricValue="4.65" metricValueMax="4.7" metricValueMin="4.6" pageId="11" pageNumber="148" unit="m" value="465.0" valueMax="470.0" valueMin="460.0">460-470 m</quantity>
,
<date id="FFE51020FFFAFF9B5D8AB03BFB81FA86" box="[957,1069,1378,1401]" pageId="11" pageNumber="148" value="1992-10-18">18.X.1992</date>
,
<specimenCount id="9D5DFD69FFFAFF9B5A0DB03BFB02FA86" box="[1082,1198,1378,1401]" pageId="11" pageNumber="148" type="generic">1 specimen</specimenCount>
dry,
<collectionCode id="ED4AAE25FFFAFF9B5CFDB025FD76FA6C" box="[714,730,1404,1427]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="148">R</collectionCode>
=
<quantity id="4CA39B05FFFAFF9B5CCEB025FCEDFA6C" box="[761,833,1404,1427]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="148" unit="cm" value="1.7">1.7 cm</quantity>
, r =
<quantity id="4CA39B05FFFAFF9B5D4FB025FC6CFA6C" box="[888,960,1404,1427]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="148" unit="cm" value="1.4">1.4 cm</quantity>
(
<collectionCode id="ED4AAE25FFFAFF9B5DE5B025FB8BFA6C" box="[978,1063,1404,1427]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="148">MNHN</collectionCode>
EcAs 11687). —
<collectionCode id="ED4AAE25FFFAFF9B5CFDB0CCFCA7FA53" box="[714,779,1429,1452]" pageId="11" pageNumber="148">SMIB</collectionCode>
8, banc Éponge (mont B), stn DW 147,
<geoCoordinate id="EE6F5027FFFAFF9B5CFDB0F7FC81FA3A" box="[714,813,1454,1477]" degrees="24" direction="south" minutes="54.9" orientation="latitude" pageId="11" pageNumber="148" precision="92" value="-24.915">24°54.9’S</geoCoordinate>
,
<geoCoordinate id="EE6F5027FFFAFF9B5D00B0F7FC06FA3A" box="[823,938,1454,1477]" degrees="168" direction="east" minutes="21.8" orientation="longitude" pageId="11" pageNumber="148" precision="92" value="168.36333">168°21.8’E</geoCoordinate>
,
<quantity id="4CA39B05FFFAFF9B5D84B0F7FB8FFA3A" box="[947,1059,1454,1477]" metricMagnitude="2" metricUnit="m" metricValue="5.2" metricValueMax="5.32" metricValueMin="5.08" pageId="11" pageNumber="148" unit="m" value="520.0" valueMax="532.0" valueMin="508.0">508-532 m</quantity>
,
<date id="FFE51020FFFAFF9B5A1AB0F7FB3CFA3A" box="[1069,1168,1454,1477]" pageId="11" pageNumber="148" value="1993-01-27">27.I.1993</date>
,
<specimenCount id="9D5DFD69FFFAFF9B5AACB0F7FCA9FA20" pageId="11" pageNumber="148" type="generic">2 specimens</specimenCount>
dry,
<collectionCode id="ED4AAE25FFFAFF9B5D0AB091FCE1FA20" box="[829,845,1480,1503]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="148">R</collectionCode>
=
<quantity id="4CA39B05FFFAFF9B5D5CB091FC19FA20" box="[875,949,1480,1503]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="148" unit="cm" value="1.4">1.4 cm</quantity>
, r =
<quantity id="4CA39B05FFFAFF9B5DD0B091FB9DFA20" box="[999,1073,1480,1503]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="148" unit="cm" value="1.2">1.2 cm</quantity>
,
<collectionCode id="ED4AAE25FFFAFF9B5A0AB091FBE1FA20" box="[1085,1101,1480,1503]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="148">R</collectionCode>
=
<quantity id="4CA39B05FFFAFF9B5A5CB091FB19FA20" box="[1131,1205,1480,1503]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="148" unit="cm" value="1.2">1.2 cm</quantity>
, r =
<quantity id="4CA39B05FFFAFF9B5CFDB0B8FCBEFA07" box="[714,786,1505,1528]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="148" unit="cm" value="1.1">1.1 cm</quantity>
(
<collectionCode id="ED4AAE25FFFAFF9B5D15B0B8FCD4FA07" box="[802,888,1505,1528]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="148">MNHN</collectionCode>
EcAs 11694). — BATHUS 3, stn DW 819,
<geoCoordinate id="EE6F5027FFFAFF9B5D65B0A3FC07F9EE" box="[850,939,1530,1553]" degrees="23" direction="south" minutes="45" orientation="latitude" pageId="11" pageNumber="148" precision="925" value="-23.75">23 °45’S</geoCoordinate>
,
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,
<quantity id="4CA39B05FFFAFF9B5A78B0A3FB7BF9EE" box="[1103,1239,1530,1553]" metricMagnitude="2" metricUnit="m" metricValue="4.82" metricValueMax="4.86" metricValueMin="4.779999999999999" pageId="11" pageNumber="148" unit="m" value="482.0" valueMax="486.0" valueMin="478.0">478-486 m</quantity>
,
<date id="FFE51020FFFAFF9B5CFDB34DFCECF9D4" box="[714,832,1556,1579]" pageId="11" pageNumber="148" value="1993-11-28">28 XI.1993</date>
,
<specimenCount id="9D5DFD69FFFAFF9B5D7BB34DFC17F9D4" box="[844,955,1556,1579]" pageId="11" pageNumber="148" type="generic">1 specimen</specimenCount>
dry,
<collectionCode id="ED4AAE25FFFAFF9B5DC5B34DFBAEF9D4" box="[1010,1026,1556,1579]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="148">R</collectionCode>
=
<quantity id="4CA39B05FFFAFF9B5A28B34DFBC4F9D4" box="[1055,1128,1556,1579]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="148" unit="cm" value="1.5">1.5 cm</quantity>
, r =
<quantity id="4CA39B05FFFAFF9B5AAEB34DFB73F9D4" box="[1177,1247,1556,1579]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="148" unit="cm" value="1.3">1.3 cm</quantity>
(
<collectionCode id="ED4AAE25FFFAFF9B5CE6B374FC89F9BB" box="[721,805,1581,1604]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="148">MNHN</collectionCode>
EcAs 11695).
</paragraph>
</subSubSection>
<subSubSection id="C341656BFFFAFF9B5CFDB30EFC0FF978" pageId="11" pageNumber="148" type="etymology">
<paragraph id="8BE436E0FFFAFF9B5CFDB30EFC0FF978" blockId="11.[714,1247,1010,1713]" pageId="11" pageNumber="148">ETYMOLOGY. — This species is named for its distinctive pentagonal shape.</paragraph>
</subSubSection>
<subSubSection id="C341656BFFFAFF9D5CFDB3C3FF4CFF7A" lastPageId="13" lastPageNumber="150" pageId="11" pageNumber="148" type="distribution">
<paragraph id="8BE436E0FFFAFF9B5CFDB3C3FB73F94E" blockId="11.[714,1247,1010,1713]" box="[714,1247,1690,1713]" pageId="11" pageNumber="148">
DISTRIBUTION. — Currently known only from
<collectingCountry id="F34C7670FFFAFF9B5A87B3C3FB73F94E" box="[1200,1247,1690,1713]" name="New Caledonia" pageId="11" pageNumber="148">New</collectingCountry>
</paragraph>
<paragraph id="8BE436E0FFFAFF9B5CFDB3EDFC03F934" blockId="11.[714,943,1716,1739]" box="[714,943,1716,1739]" pageId="11" pageNumber="148">
Caledonia.
<quantity id="4CA39B05FFFAFF9B5D0DB3EDFC07F934" box="[826,939,1716,1739]" metricMagnitude="2" metricUnit="m" metricValue="4.96" metricValueMax="5.32" metricValueMin="4.6" pageId="11" pageNumber="148" unit="m" value="496.0" valueMax="532.0" valueMin="460.0">
460-
<specimenCount id="9D5DFD69FFFAFF9B5D5FB3EDFC07F934" box="[872,939,1716,1739]" pageId="11" pageNumber="148" type="generic">532 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="8BE436E0FFFDFF9C5CE8B52DFB7CFF7A" blockId="12.[735,1232,116,133]" box="[735,1232,116,133]" pageId="12" pageNumber="149">
Phylogeny of
<taxonomicName id="4C5B4D63FFFDFF9C5D74B52DFC2CFF7A" authorityName="Sladen" authorityYear="1889" box="[835,896,116,133]" class="Asteroidea" family="Goniasteridae" genus="Iconaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="149" phylum="Echinodermata" rank="genus">
<emphasis id="B92FEAF2FFFDFF9C5D74B52DFC2CFF7A" box="[835,896,116,133]" italics="true" pageId="12" pageNumber="149">Iconaster</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5B4D63FFFDFF9C5D93B52DFC51FF7A" authorityName="Fisher" authorityYear="1917" box="[932,1021,116,133]" class="Asteroidea" family="Goniasteridae" genus="Glyphodiscus" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="149" phylum="Echinodermata" rank="genus">
<emphasis id="B92FEAF2FFFDFF9C5D93B52DFC51FF7A" box="[932,1021,116,133]" italics="true" pageId="12" pageNumber="149">Glyphodiscus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C5B4D63FFFDFF9C5A31B52DFBDBFF7A" box="[1030,1143,116,133]" class="Echinodermata" kingdom="Animalia" pageId="12" pageNumber="159" phylum="Echinodermata" rank="class">Echinodermata</taxonomicName>
,
<taxonomicName id="4C5B4D63FFFDFF9C5A48B52DFB60FF7A" box="[1151,1228,116,133]" class="Asteroidea" kingdom="Animalia" pageId="12" pageNumber="149" phylum="Echinodermata" rank="class">Asteroidea</taxonomicName>
)
</paragraph>
<caption id="DF246668FFFDFF9C5EA0B325FB4CF947" ID-DOI="http://doi.org/10.5281/zenodo.5392853" ID-Zenodo-Dep="5392853" httpUri="https://zenodo.org/record/5392853/files/figure.png" pageId="12" pageNumber="149" startId="12.[151,162,1660,1677]" targetBox="[151,1245,368,1606]" targetPageId="12">
<paragraph id="8BE436E0FFFDFF9C5EA0B325FB4CF947" blockId="12.[151,1248,1660,1720]" pageId="12" pageNumber="149">
FIG. 3. —
<emphasis id="B92FEAF2FFFDFF9C5EDCB325FEA1F972" bold="true" box="[235,269,1660,1677]" pageId="12" pageNumber="149">A -C</emphasis>
,
<taxonomicName id="4C5B4D63FFFDFF9C5F2EB325FE46F972" authority="Mah, 2005" authorityName="Mah" authorityYear="2005" box="[281,490,1660,1677]" class="Asteroidea" family="Goniasteridae" genus="Glyphodiscus" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="149" phylum="Echinodermata" rank="species" species="magnificus" status="sp. nov.">
<emphasis id="B92FEAF2FFFDFF9C5F2EB325FE46F972" box="[281,490,1660,1677]" italics="true" pageId="12" pageNumber="149">Glyphodiscus magnificus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C5789FFFDFF9C5FC6B325FD88F972" box="[497,548,1660,1677]" pageId="12" pageNumber="149" rank="species">n. sp.</taxonomicNameLabel>
, New Caledonia, paratype, dry (MNHN EcAs 11682);
<emphasis id="B92FEAF2FFFDFF9C5DDEB325FC5AF972" bold="true" box="[1001,1014,1660,1677]" pageId="12" pageNumber="149">A</emphasis>
, abactinal view;
<emphasis id="B92FEAF2FFFDFF9C5AB2B325FB3EF972" bold="true" box="[1157,1170,1660,1677]" pageId="12" pageNumber="149">B</emphasis>
, superomarginal plate shape;
<emphasis id="B92FEAF2FFFDFF9C5F60B3C8FEC9F95D" bold="true" box="[343,357,1681,1698]" pageId="12" pageNumber="149">C</emphasis>
, oral and adambulacral regions;
<emphasis id="B92FEAF2FFFDFF9C5CB3B3C8FD3EF95D" bold="true" box="[644,658,1681,1698]" pageId="12" pageNumber="149">D</emphasis>
,
<emphasis id="B92FEAF2FFFDFF9C5CA9B3C8FD06F95D" bold="true" box="[670,682,1681,1698]" pageId="12" pageNumber="149">E</emphasis>
,
<taxonomicName id="4C5B4D63FFFDFF9C5C81B3C8FC30F95D" authority="Mah, 2005" authorityName="Mah" authorityYear="2005" box="[694,924,1681,1698]" class="Asteroidea" family="Goniasteridae" genus="Glyphodiscus" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="149" phylum="Echinodermata" rank="species" species="pentagonalis" status="sp. nov.">
<emphasis id="B92FEAF2FFFDFF9C5C81B3C8FC30F95D" box="[694,924,1681,1698]" italics="true" pageId="12" pageNumber="149">Glyphodiscus pentagonalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C5789FFFDFF9C5D94B3C8FC7BF95D" box="[931,983,1681,1698]" pageId="12" pageNumber="149" rank="species">n. sp.</taxonomicNameLabel>
, New Caledonia, holotype, dry (MNHN EcAs 11688);
<emphasis id="B92FEAF2FFFDFF9C5F60B3FEFEC9F947" bold="true" box="[343,357,1703,1720]" pageId="12" pageNumber="149">D</emphasis>
, abactinal view;
<emphasis id="B92FEAF2FFFDFF9C5FCCB3FEFDABF947" bold="true" box="[507,519,1703,1720]" pageId="12" pageNumber="149">E</emphasis>
, oral and adambulacral regions. Scale bars: A, 5.0 mm; B, D, 3.0 mm; C, 2.0 mm;
</paragraph>
</caption>
<paragraph id="8BE436E0FFFCFF9D5E91B52DFF4CFF7A" blockId="13.[166,224,116,133]" box="[166,224,116,133]" pageId="13" pageNumber="150">Mah C.</paragraph>
</subSubSection>
<subSubSection id="C341656BFFFCFF9D5EA0B5ABFDC9FE69" pageId="13" pageNumber="150" type="diagnosis">
<paragraph id="8BE436E0FFFCFF9D5EA0B5ABFD85FEAA" blockId="13.[151,684,242,406]" pageId="13" pageNumber="150">DIAGNOSIS. — Pentagonal body with few, smooth marginal plates (≈ 4-6) in each interadii. Superomarginal plates at each arm tip triangular in outline, and directly abutting across arm.</paragraph>
<paragraph id="8BE436E0FFFCFF9D5EA0B43FFDC9FE69" blockId="13.[151,684,242,406]" pageId="13" pageNumber="150">APOMORPHY LIST. — 12, marginal plates elongate. 13, enlarged basal superomarginals. 27, arms short.</paragraph>
</subSubSection>
<subSubSection id="C341656BFFFCFF9D5EA0B48BFD98F974" pageId="13" pageNumber="150" type="description">
<paragraph id="8BE436E0FFFCFF9D5EA0B48BFE84FE15" blockId="13.[151,684,466,1707]" box="[151,296,466,491]" pageId="13" pageNumber="150">DESCRIPTION</paragraph>
<paragraph id="8BE436E0FFFCFF9D5EA0B4ABFD94FDD4" blockId="13.[151,684,466,1707]" pageId="13" pageNumber="150">Disk pentagonal. interradial arcs linear. Arms, short and stubby. Body, stout and flat.</paragraph>
<paragraph id="8BE436E0FFFCFF9D5EA0B76BFD3CFBB4" blockId="13.[151,684,466,1707]" pageId="13" pageNumber="150">
Disk area pentagonal. Plates bare, flat. Surface ornamentation absent. Surfaces smooth. Plates more convex on papular regions along radii than on interradial regions of disk. Papular regions composed of six plates surrounding a single seventh plate in the center. Plates flanked at corners by papulae. Abactinals hexagonal to polygonal in shape, especially those around disk center. Plates more elongate, angular at contact between abactinal plates and superomarginal plates. Primary circlet plates at disk center enlarged, plates adjacent to contact with superomarginals smaller. Two enlarged heptagonal plates in each interradius. All plates with complete peripheral border of
<emphasis id="B92FEAF2FFFCFF9D5FD1B68BFE5BFC14" box="[486,503,978,1003]" italics="true" pageId="13" pageNumber="150">c.</emphasis>
20-60 granularrectangular plates. Plates adjacent to superomarginal surface with scattered pedicellariae. Madreporite triangular, flanked by three plates.
</paragraph>
<paragraph id="8BE436E0FFFCFF9D5EA0B10BFE1FFAD4" blockId="13.[151,684,466,1707]" pageId="13" pageNumber="150">
Approximately six marginals in smaller specimens (R = 1.0 cm) vs approximately eight superomarginal plates (R =
<quantity id="4CA39B05FFFCFF9D5F56B1CBFE1DFB54" box="[353,433,1170,1195]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="13" pageNumber="150" unit="cm" value="2.4">2.4 cm</quantity>
) in larger specimens. Typically about two superomarginals per interradius in specimens above R = 1.0 cm. Marginal plates smooth, devoid of surface ornamentation. Plates flat, largely smooth.
</paragraph>
<paragraph id="8BE436E0FFFCFF9D5EA0B06BFD98F974" blockId="13.[151,684,466,1707]" pageId="13" pageNumber="150">Superomarginals at base of arm enlarged and in direct contact. Shape, asymmetrically pentagonal. Enlarged plates approximately 1.5 times those of the interradial superomarginal plates. Superomarginal plates along arm are four-sided but roughly triangular in shape. No enlarged inferomarginals. Inferomarginals are roughly triangular in shape. All marginal plates bordered by a complete ring of rectangular accessories/granules similar to those of the abactinal and actinal surfaces. Inferomarginals smooth, flat.</paragraph>
</subSubSection>
</treatment>
</document>
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