778 lines
61 KiB
XML
778 lines
61 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4324231" ID-GBIF-Dataset="d5ddaf75-c39b-49b3-bbf5-7e7af2f7ad66" ID-GBIF-Taxon="168144935" ID-Zenodo-Dep="4324231" approvalRequired="5" approvalRequired_for_bibRefs="2" approvalRequired_for_document="1" approvalRequired_for_textStreams="2" checkinTime="1601372077630" checkinUser="plazi" docAuthor="Agudelo‐Zamora, Henry D., Tavera, José, Murillo, Yiskar D. & Ortega‐Lara, Armando" docDate="2020" docId="03F287B4FF90FFD3D575FB857920FEA0" docLanguage="en" docName="jFishBiology.97.4.1-14.pdf" docOrigin="Journal of Fish Biology 97 (4)" docSource="http://dx.doi.org/10.1111/jfb.14527" docStyle="DocumentStyle{}" docTitle="Characidium dule Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara 2020, new species" docType="treatment" docVersion="6" lastPageId="9" lastPageNumber="10" masterDocId="FFCBFFCCFF97FFDAD509FFD07C36FFEF" masterDocTitle="The unknown diversity of the genus Characidium (Characiformes: Crenuchidae) in the Chocó biogeographic region, Colombian Andes: Two new species supported by morphological and molecular data" masterLastPageNumber="14" masterPageNumber="1" pageId="7" pageNumber="8" updateTime="1635496756590" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The unknown diversity of the genus Characidium (Characiformes: Crenuchidae) in the Chocó biogeographic region, Colombian Andes: Two new species supported by morphological and molecular data</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Agudelo‐Zamora, Henry D.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-8571-7235</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tavera, José</mods:namePart>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Murillo, Yiskar D.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ortega‐Lara, Armando</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-2015-7948</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Journal of Fish Biology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-09-27</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>97</mods:number>
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||
</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>14</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.1111/jfb.14527</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1111/jfb.14527</mods:identifier>
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<mods:identifier type="GBIF-Dataset">d5ddaf75-c39b-49b3-bbf5-7e7af2f7ad66</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4058131</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4324231" ID-GBIF-Taxon="168144935" ID-Zenodo-Dep="4324231" LSID="urn:lsid:plazi:treatment:03F287B4FF90FFD3D575FB857920FEA0" httpUri="http://treatment.plazi.org/id/03F287B4FF90FFD3D575FB857920FEA0" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
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<subSubSection pageId="7" pageNumber="8" type="nomenclature">
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<paragraph blockId="7.[124,598,1108,1134]" box="[124,598,1108,1134]" pageId="7" pageNumber="8">
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<emphasis bold="true" box="[124,176,1109,1134]" pageId="7" pageNumber="8">3.12</emphasis>
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|
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<heading bold="true" box="[246,598,1108,1134]" fontSize="10" level="3" pageId="7" pageNumber="8" reason="0">
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<emphasis bold="true" box="[246,598,1108,1134]" pageId="7" pageNumber="8">
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<taxonomicName authority="Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara, 2020" authorityName="Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara" authorityYear="2020" box="[246,441,1108,1134]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="dule" status="new species">
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<emphasis bold="true" box="[246,441,1108,1134]" italics="true" pageId="7" pageNumber="8">Characidium dule</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[455,598,1109,1134]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<caption box="[124,390,1182,1202]" pageId="7" pageNumber="8">
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<paragraph blockId="7.[124,771,1182,1341]" box="[124,390,1182,1202]" pageId="7" pageNumber="8">Figures 6 and 7, and Table 2.</paragraph>
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</caption>
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<paragraph blockId="7.[124,771,1182,1341]" box="[145,599,1217,1237]" pageId="7" pageNumber="8">urn:lsid:zoobank.org:act:FFF769BD-AA23-4FEC-</paragraph>
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<paragraph blockId="7.[124,771,1182,1341]" box="[124,345,1251,1271]" pageId="7" pageNumber="8">AF33-1770A612D41D.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="materials_examined">
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<paragraph blockId="7.[124,771,1182,1341]" pageId="7" pageNumber="8">
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<taxonomicName authorityName="J.T.Reinhardt" authorityYear="1867" box="[164,275,1286,1306]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
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<emphasis box="[164,275,1286,1306]" italics="true" pageId="7" pageNumber="8">Characidium</emphasis>
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</taxonomicName>
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sp. 4
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<emphasis bold="true" box="[337,345,1287,1306]" pageId="7" pageNumber="8">“</emphasis>
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Arquia
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<emphasis bold="true" box="[405,433,1287,1306]" pageId="7" pageNumber="8">” –</emphasis>
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<bibRefCitation author="Agudelo-Zamora, H. & Ortega-Lara, A. & Taphorn, D." box="[441,712,1286,1306]" journalOrPublisher="Zootaxa" pageId="7" pageNumber="8" pagination="249 - 263" part="4768" refId="ref9983" refString="Agudelo-Zamora, H., Ortega-Lara, A., & Taphorn, D. (2020). Characidium chancoense, a new species of south American darter from the Rio Cauca drainage, Colombia (Characiformes: Crenuchidae). Zootaxa, 4768 (2), 249 - 263. https: // doi. org / 10.11646 / zootaxa. 4768.2.6." title="Characidium chancoense, a new species of south American darter from the Rio Cauca drainage, Colombia (Characiformes: Crenuchidae)" type="journal article" year="2020">
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Agudelo-Zamora
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<emphasis box="[603,644,1286,1306]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
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., 2020
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||
</bibRefCitation>
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||
, p. 14 (material examined).
|
||
</paragraph>
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||
<paragraph blockId="7.[124,356,1421,1446]" box="[124,356,1421,1446]" pageId="7" pageNumber="8">
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<emphasis bold="true" box="[124,176,1421,1446]" pageId="7" pageNumber="8">3.13</emphasis>
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|
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<heading bold="true" box="[246,356,1421,1446]" fontSize="10" level="3" pageId="7" pageNumber="8" reason="0">
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<emphasis bold="true" box="[246,356,1421,1446]" pageId="7" pageNumber="8">
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<typeStatus box="[246,356,1421,1446]" pageId="7" pageNumber="8">Holotype</typeStatus>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="7.[124,773,1494,1583]" pageId="7" pageNumber="8">
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<materialsCitation ID-GBIF-Occurrence="2865196302" collectingDate="2017-04-05" collectionCode="IMCN" collectorName="A. Ortega-Lara." country="Colombia" county="Arquia" latitude="8.001972" location="Choco" longLatPrecision="1" longitude="-77.11561" municipality="rio Arquia" pageId="7" pageNumber="8" specimenCode="IMCN 8941" specimenCount="1" typeStatus="holotype">
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<specimenCode box="[124,238,1494,1514]" collectionCode="IMCN" pageId="7" pageNumber="8">IMCN 8941</specimenCode>
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,
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<quantity box="[247,329,1494,1514]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="7" pageNumber="8" unit="mm" value="37.4">37.4 mm</quantity>
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,
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<collectingCountry box="[366,456,1494,1514]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
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,
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,
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<location LSID="urn:lsid:plazi:treatment:03F287B4FF90FFD3D575FB857920FEA0:8E846079FF90FFDDD711FA067E6AFA05" box="[536,604,1494,1514]" country="Colombia" county="Arquia" latitude="8.001972" longLatPrecision="1" longitude="-77.11561" municipality="rio Arquia" name="Unguia" pageId="7" pageNumber="8">Unguia</location>
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,
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<collectingMunicipality box="[613,707,1494,1514]" pageId="7" pageNumber="8">río Arquía</collectingMunicipality>
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, at the village of
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,
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08 00
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<superScript attach="none" box="[372,376,1526,1539]" fontSize="6" pageId="7" pageNumber="8">'</superScript>
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07.1
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<superScript attach="right" box="[418,426,1526,1539]" fontSize="6" pageId="7" pageNumber="8">''</superScript>
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N
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,
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77 06
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<emphasis bold="true" box="[522,526,1526,1539]" italics="true" pageId="7" pageNumber="8">
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<superScript attach="none" box="[522,526,1526,1539]" fontSize="6" pageId="7" pageNumber="8">'</superScript>
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</emphasis>
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56.2
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<emphasis bold="true" box="[569,577,1526,1539]" italics="true" pageId="7" pageNumber="8">
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<superScript attach="right" box="[569,577,1526,1539]" fontSize="6" pageId="7" pageNumber="8">''</superScript>
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W
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,
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<collectorName box="[620,773,1529,1549]" pageId="7" pageNumber="8">A. Ortega-Lara.</collectorName>
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<date box="[124,254,1563,1583]" pageId="7" pageNumber="8" value="2017-04-05">
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<collectingDate box="[124,254,1563,1583]" pageId="7" pageNumber="8" value="2017-04-05">05 April 2017</collectingDate>
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</date>
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</materialsCitation>
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.
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||
</paragraph>
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<paragraph blockId="7.[124,363,1663,1688]" box="[124,363,1663,1688]" pageId="7" pageNumber="8">
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<emphasis bold="true" box="[124,176,1663,1688]" pageId="7" pageNumber="8">3.14</emphasis>
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||
|
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||
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||
<typeStatus box="[246,363,1663,1688]" pageId="7" pageNumber="8">Paratypes</typeStatus>
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</emphasis>
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||
</heading>
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</paragraph>
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||
<paragraph blockId="7.[124,773,1737,1930]" pageId="7" pageNumber="8">
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||
All from
|
||
<materialsCitation ID-GBIF-Occurrence="2865196303" collectionCode="IMCN" country="Colombia" location="Colombia" pageId="7" pageNumber="8" specimenCode="IMCN 8929" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<collectingCountry box="[210,306,1737,1757]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
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.
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<specimenCode box="[315,434,1737,1757]" collectionCode="IMCN" pageId="7" pageNumber="8">IMCN 8929</specimenCode>
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, 7 + 1 C&S
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,
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32.8
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44.7 mm
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<emphasis box="[124,134,1771,1791]" italics="true" pageId="7" pageNumber="8">L</emphasis>
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from the same locality of the holotype
|
||
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;
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<materialsCitation ID-GBIF-Occurrence="2865196301" collectionCode="IMCN" pageId="7" pageNumber="8" specimenCode="IMCN 8942" specimenCount="1" typeStatus="paratype">
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<specimenCode box="[638,769,1771,1791]" collectionCode="IMCN" pageId="7" pageNumber="8">IMCN 8942</specimenCode>
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, 1,
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<subScript attach="left" box="[261,270,1815,1828]" fontSize="5" pageId="7" pageNumber="8">S</subScript>
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||
from the same locality of the holotype
|
||
</materialsCitation>
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;
|
||
<materialsCitation ID-GBIF-Occurrence="2865196304" collectingDate="2019-08-01" collectionCode="CHIFF" collectorName="A. Ortega-Lara & N. Lujan." country="Colombia" county="Tutunendo river" latitude="5.7443614" location="Choco" longLatPrecision="1" longitude="-76.53172" municipality="Quibdo" pageId="7" pageNumber="8" specimenCode="CHIFF 0012" specimenCount="1" typeStatus="paratype">
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, 1,
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,
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<collectingCountry box="[338,434,1841,1861]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
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,
|
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|
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,
|
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<collectingMunicipality box="[520,596,1841,1861]" pageId="7" pageNumber="8">Quibdó</collectingMunicipality>
|
||
,
|
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<collectingCounty box="[606,769,1841,1861]" pageId="7" pageNumber="8">Tutunendo river</collectingCounty>
|
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,
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5 44
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<superScript attach="none" box="[172,176,1873,1886]" fontSize="6" pageId="7" pageNumber="8">'</superScript>
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39.7
|
||
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|
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<superScript attach="right" box="[221,230,1873,1886]" fontSize="6" pageId="7" pageNumber="8">''</superScript>
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N
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</geoCoordinate>
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,
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<geoCoordinate box="[261,406,1873,1895]" degrees="76" direction="west" minutes="31" orientation="longitude" pageId="7" pageNumber="8" precision="1" seconds="54.2" value="-76.53172">
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76 31
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<superScript attach="none" box="[321,325,1873,1886]" fontSize="6" pageId="7" pageNumber="8">'</superScript>
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</emphasis>
|
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54.2
|
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<emphasis bold="true" box="[371,381,1873,1886]" italics="true" pageId="7" pageNumber="8">
|
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<superScript attach="right" box="[371,381,1873,1886]" fontSize="6" pageId="7" pageNumber="8">''</superScript>
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</emphasis>
|
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W
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||
</geoCoordinate>
|
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.
|
||
<collectorName box="[416,565,1875,1895]" pageId="7" pageNumber="8">A. Ortega-Lara</collectorName>
|
||
,
|
||
<collectorName box="[575,664,1875,1895]" pageId="7" pageNumber="8">N. Lujan.</collectorName>
|
||
<date pageId="7" pageNumber="8" value="2019-08-01">
|
||
<collectingDate pageId="7" pageNumber="8" value="2019-08-01">01 August 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="7" pageNumber="8" type="diagnosis">
|
||
<paragraph blockId="7.[821,1056,693,718]" box="[821,1056,693,718]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[821,873,693,718]" pageId="7" pageNumber="8">3.15</emphasis>
|
||
|
|
||
<heading bold="true" box="[942,1056,693,718]" fontSize="10" level="3" pageId="7" pageNumber="8" reason="0">
|
||
<emphasis bold="true" box="[942,1056,693,718]" pageId="7" pageNumber="8">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="7.[821,1470,766,1410]" pageId="7" pageNumber="8">
|
||
<taxonomicName authorityName="Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara" authorityYear="2020" box="[821,974,766,786]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="dule">
|
||
<emphasis box="[821,974,766,786]" italics="true" pageId="7" pageNumber="8">Characidium dule</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from all other
|
||
<taxonomicName authorityName="J.T.Reinhardt" authorityYear="1867" box="[1311,1422,766,786]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1311,1422,766,786]" italics="true" pageId="7" pageNumber="8">Characidium</emphasis>
|
||
</taxonomicName>
|
||
species by having a unique colour pattern in the humeral region that consists of a small marking, similar to a black line, between the supraoccipital and the beginning of the opercle, followed by a light point (golden in live specimens or white in preserved specimens), and a humeral round blotch which is formed by two to three scales (
|
||
<figureCitation box="[825,915,974,994]" captionStart="FIGURE 5" captionStartId="4.[816,907,1916,1935]" captionTargetBox="[818,1461,1393,1876]" captionTargetId="figure-414@4.[818,1461,1393,1876]" captionText="FIGURE 5 Typical habitat of C. dule, Ingará River, San José Del Palmar, Chocó, Colombia" figureDoi="http://doi.org/10.5281/zenodo.4058141" httpUri="https://zenodo.org/record/4058141/files/figure.png" pageId="7" pageNumber="8">Figures 5</figureCitation>
|
||
,
|
||
<figureCitation box="[924,938,974,994]" captionStart="FIGURE 6" captionStartId="6.[119,210,1916,1935]" captionTargetBox="[122,765,1235,1875]" captionTargetId="figure-512@6.[122,765,1235,1875]" captionText="FIGURE 6 C. dule n. sp. IMCN 8941, Holotype 37.4 mm LS. Scale bar = 1 cm" figureDoi="http://doi.org/10.5281/zenodo.4058143" httpUri="https://zenodo.org/record/4058143/files/figure.png" pageId="7" pageNumber="8">6</figureCitation>
|
||
), in addition to a triangular blotch at the end of the caudal peduncle where pigmentation varies in size starting at ½ scale, followed by 1 scale and its base is 2 scales. In mature specimens (recognized by their purplish colour) the caudal triangular point is less visible. It is distinguished from
|
||
<taxonomicName authority="Steindachner 1882" authorityName="Steindachner" authorityYear="1882" box="[1112,1427,1113,1133]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="purpuratum">
|
||
<emphasis box="[1112,1242,1113,1133]" italics="true" pageId="7" pageNumber="8">C. purpuratum</emphasis>
|
||
Steindachner 1882
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Breder 1925" authorityName="Breder" authorityYear="1925" box="[821,1030,1147,1167]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="marshi">
|
||
<emphasis box="[821,905,1147,1167]" italics="true" pageId="7" pageNumber="8">C. marshi</emphasis>
|
||
Breder 1925
|
||
</taxonomicName>
|
||
by having lower number of dorsal (7
|
||
<emphasis bold="true" box="[1377,1389,1148,1167]" pageId="7" pageNumber="8">–</emphasis>
|
||
9
|
||
<emphasis box="[1408,1426,1147,1167]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
|
||
. 11) and anal (6
|
||
<emphasis bold="true" box="[924,936,1183,1202]" pageId="7" pageNumber="8">–</emphasis>
|
||
7
|
||
<emphasis box="[956,975,1182,1202]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
|
||
.
|
||
<quantity box="[986,1022,1182,1202]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="7" pageNumber="8" unit="in" value="8.0">8 in</quantity>
|
||
<taxonomicName authorityName="Steindachner" authorityYear="1882" box="[1029,1161,1182,1202]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="purpuratum">
|
||
<emphasis box="[1029,1161,1182,1202]" italics="true" pageId="7" pageNumber="8">C. purpuratum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity box="[1208,1269,1182,1202]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.778" pageId="7" pageNumber="8" unit="in" value="8.0" valueMax="9.0" valueMin="7.0">
|
||
7
|
||
<emphasis bold="true" box="[1220,1232,1183,1202]" pageId="7" pageNumber="8">–</emphasis>
|
||
9 in
|
||
</quantity>
|
||
<taxonomicName authorityName="Breder" authorityYear="1925" box="[1276,1360,1182,1202]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="marshi">
|
||
<emphasis box="[1276,1360,1182,1202]" italics="true" pageId="7" pageNumber="8">C. marshi</emphasis>
|
||
</taxonomicName>
|
||
) fin rays. It is also distinguished from
|
||
<taxonomicName authorityName="Steindachner" authorityYear="1882" box="[1067,1199,1217,1237]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="purpuratum">
|
||
<emphasis box="[1067,1199,1217,1237]" italics="true" pageId="7" pageNumber="8">C. purpuratum</emphasis>
|
||
</taxonomicName>
|
||
by the presence of scales in the isthmus and the area between the pectorals (
|
||
<emphasis box="[1280,1298,1251,1271]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
|
||
. isthmus and area between the pectoral without scales). It is also distinguished from
|
||
<taxonomicName authorityName="Breder" authorityYear="1925" box="[821,909,1321,1341]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="marshi">
|
||
<emphasis box="[821,909,1321,1341]" italics="true" pageId="7" pageNumber="8">C. marshi</emphasis>
|
||
</taxonomicName>
|
||
by having a smaller number of scales in the lateral line (30
|
||
<emphasis bold="true" box="[851,863,1356,1375]" pageId="7" pageNumber="8">–</emphasis>
|
||
33
|
||
<emphasis box="[896,915,1355,1375]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
|
||
. 31
|
||
<emphasis bold="true" box="[953,965,1356,1375]" pageId="7" pageNumber="8">–</emphasis>
|
||
35) and by the absence of postcleithrum I (
|
||
<emphasis box="[1388,1407,1355,1375]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
|
||
. presence of postcleithrum).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
|
||
<paragraph blockId="7.[821,1079,1490,1515]" box="[821,1079,1490,1515]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[821,873,1490,1515]" pageId="7" pageNumber="8">3.16</emphasis>
|
||
|
|
||
<heading bold="true" box="[942,1079,1490,1515]" fontSize="10" level="3" pageId="7" pageNumber="8" reason="0">
|
||
<emphasis bold="true" box="[942,1079,1490,1515]" pageId="7" pageNumber="8">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="7.[821,1469,1563,1930]" lastBlockId="8.[119,768,1113,1757]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
Morphometric data can be seen in
|
||
<tableCitation box="[1154,1226,1563,1583]" captionStart="TABLE 2" captionStartId="8.[119,197,139,158]" captionTargetBox="[138,1361,186,993]" captionText="TABLE 2 Morphometric data from C. dule n. sp. (n = 9)" httpUri="http://table.plazi.org/id/DF24662AFF9FFFD2D57EFF5B7EA5FF70" pageId="7" pageNumber="8" tableUuid="DF24662AFF9FFFD2D57EFF5B7EA5FF70">Table 2</tableCitation>
|
||
, the largest known specimen not exceeding
|
||
<quantity box="[1015,1098,1598,1618]" metricMagnitude="-2" metricUnit="m" metricValue="4.47" pageId="7" pageNumber="8" unit="mm" value="44.7">44.7 mm</quantity>
|
||
<emphasis box="[1108,1118,1598,1618]" italics="true" pageId="7" pageNumber="8">L</emphasis>
|
||
<subScript attach="left" box="[1118,1127,1607,1620]" fontSize="5" pageId="7" pageNumber="8">S</subScript>
|
||
. Body slightly compressed laterally with slightly convex dorsal profile from snout to supraoccipital fontanelle, ascending slightly convex to origin of dorsal fin, and ascending at another convex angle from beginning of dorsal to adipose fin, descends slightly in a straight line and right angle from adipose to first procuring radius of tail fin. Slightly concave ventral profile, vertex on pelvic fin, ends dentary at anterior part and ventral procurrent rays at posterior. Greatest body depth is at or just before origin of dorsal fin. Slightly convex pointed snout from dorsal margin of orbit. Maxilla short, just barely exceeding anterior margin of orbital, exceeding posterior nostril. Circular orbit larger than snout length, similar in size to maxilla; iris round. Dorsal base fin with chromatophores forming a band.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="DF24662AFF9FFFD2D57EFF5B7EA5FF70" box="[119,659,139,159]" httpUri="http://table.plazi.org/id/DF24662AFF9FFFD2D57EFF5B7EA5FF70" pageId="8" pageNumber="9" startId="8.[119,197,139,158]" targetBox="[138,1361,186,993]" targetIsTable="true" targetPageId="8">
|
||
<paragraph blockId="8.[119,218,139,158]" box="[119,659,139,159]" lastBlockId="8.[248,659,139,159]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[119,218,139,158]" pageId="8" pageNumber="9">TABLE 2</emphasis>
|
||
Morphometric data from
|
||
<taxonomicName authority="Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara, 2020" authorityName="Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara" authorityYear="2020" box="[482,542,139,159]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="dule" status="n. sp.">
|
||
<emphasis box="[482,542,139,159]" italics="true" pageId="8" pageNumber="9">C. dule</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[547,597,139,159]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<emphasis box="[606,617,139,159]" italics="true" pageId="8" pageNumber="9">n</emphasis>
|
||
= 9)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="8" pageNumber="9">
|
||
<table box="[138,1361,186,993]" gridcols="7" gridrows="22" pageId="8" pageNumber="9">
|
||
<tr box="[138,1361,186,205]" gridrow="0" pageId="8" pageNumber="9" rowspan-0="1">
|
||
<th box="[453,535,186,205]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">
|
||
<heading bold="true" box="[453,535,187,205]" fontSize="7" level="6" pageId="8" pageNumber="9" reason="2">Holotype</heading>
|
||
</th>
|
||
<th box="[631,716,186,205]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">
|
||
<heading bold="true" box="[631,716,187,205]" fontSize="7" level="6" pageId="8" pageNumber="9" reason="2">Minimum</heading>
|
||
</th>
|
||
<th box="[811,900,186,205]" gridcol="3" gridrow="0" pageId="8" pageNumber="9">
|
||
<heading bold="true" box="[811,900,187,205]" fontSize="7" level="6" pageId="8" pageNumber="9" reason="2">Maximum</heading>
|
||
</th>
|
||
<th box="[995,1084,186,205]" gridcol="4" gridrow="0" pageId="8" pageNumber="9">
|
||
<heading bold="true" box="[995,1050,187,205]" fontSize="7" level="6" pageId="8" pageNumber="9" reason="2">Range</heading>
|
||
</th>
|
||
<th box="[1180,1231,186,205]" gridcol="5" gridrow="0" pageId="8" pageNumber="9">
|
||
<heading bold="true" box="[1180,1231,187,205]" fontSize="7" level="6" pageId="8" pageNumber="9" reason="2">
|
||
<emphasis bold="true" box="[1180,1231,187,205]" pageId="8" pageNumber="9">Mean</emphasis>
|
||
</heading>
|
||
</th>
|
||
<th box="[1325,1361,186,205]" gridcol="6" gridrow="0" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[1325,1361,186,205]" pageId="8" pageNumber="9">S.D.</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[138,1361,228,246]" gridrow="1" pageId="8" pageNumber="9" rowspan-6="1">
|
||
<th box="[138,358,228,246]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">TL</th>
|
||
<td box="[453,535,228,246]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">45.07</td>
|
||
<td box="[631,716,228,246]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">41.9</td>
|
||
<td box="[811,900,228,246]" gridcol="3" gridrow="1" pageId="8" pageNumber="9">54.3</td>
|
||
<td box="[995,1084,228,246]" gridcol="4" gridrow="1" pageId="8" pageNumber="9">
|
||
41.9
|
||
<emphasis bold="true" box="[1034,1045,228,246]" pageId="8" pageNumber="9">–</emphasis>
|
||
54.3
|
||
</td>
|
||
<td box="[1180,1231,228,246]" gridcol="5" gridrow="1" pageId="8" pageNumber="9">48.2</td>
|
||
</tr>
|
||
<tr box="[138,1361,265,284]" gridrow="2" pageId="8" pageNumber="9" rowspan-6="1">
|
||
<th box="[138,358,265,284]" gridcol="0" gridrow="2" pageId="8" pageNumber="9">SL</th>
|
||
<td box="[453,535,265,284]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">37.42</td>
|
||
<td box="[631,716,265,284]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">32.8</td>
|
||
<td box="[811,900,265,284]" gridcol="3" gridrow="2" pageId="8" pageNumber="9">44.7</td>
|
||
<td box="[995,1084,265,284]" gridcol="4" gridrow="2" pageId="8" pageNumber="9">
|
||
32,8
|
||
<emphasis bold="true" box="[1034,1045,266,284]" pageId="8" pageNumber="9">–</emphasis>
|
||
44.7
|
||
</td>
|
||
<td box="[1180,1231,265,284]" gridcol="5" gridrow="2" pageId="8" pageNumber="9">38.8</td>
|
||
</tr>
|
||
<tr box="[138,1361,302,321]" gridrow="3" pageId="8" pageNumber="9" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
||
<th box="[138,358,302,321]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">% SL</th>
|
||
</tr>
|
||
<tr box="[138,1361,340,358]" gridrow="4" pageId="8" pageNumber="9">
|
||
<th box="[138,358,340,358]" gridcol="0" gridrow="4" pageId="8" pageNumber="9">Head length</th>
|
||
<td box="[453,535,340,358]" gridcol="1" gridrow="4" pageId="8" pageNumber="9">21.9</td>
|
||
<td box="[631,716,340,358]" gridcol="2" gridrow="4" pageId="8" pageNumber="9">21.7</td>
|
||
<td box="[811,900,340,358]" gridcol="3" gridrow="4" pageId="8" pageNumber="9">27.2</td>
|
||
<td box="[995,1084,340,358]" gridcol="4" gridrow="4" pageId="8" pageNumber="9">
|
||
21.7
|
||
<emphasis bold="true" box="[1034,1045,340,358]" pageId="8" pageNumber="9">–</emphasis>
|
||
27.2
|
||
</td>
|
||
<td box="[1180,1231,340,358]" gridcol="5" gridrow="4" pageId="8" pageNumber="9">23.7</td>
|
||
<td box="[1325,1361,340,358]" gridcol="6" gridrow="4" pageId="8" pageNumber="9">1.6</td>
|
||
</tr>
|
||
<tr box="[138,1361,378,396]" gridrow="5" pageId="8" pageNumber="9">
|
||
<th box="[138,358,378,396]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">Pectoral distance</th>
|
||
<td box="[453,535,378,396]" gridcol="1" gridrow="5" pageId="8" pageNumber="9">23.1</td>
|
||
<td box="[631,716,378,396]" gridcol="2" gridrow="5" pageId="8" pageNumber="9">20.9</td>
|
||
<td box="[811,900,378,396]" gridcol="3" gridrow="5" pageId="8" pageNumber="9">25.9</td>
|
||
<td box="[995,1084,378,396]" gridcol="4" gridrow="5" pageId="8" pageNumber="9">
|
||
20.9
|
||
<emphasis bold="true" box="[1034,1045,378,396]" pageId="8" pageNumber="9">–</emphasis>
|
||
25.9
|
||
</td>
|
||
<td box="[1180,1231,378,396]" gridcol="5" gridrow="5" pageId="8" pageNumber="9">23.4</td>
|
||
<td box="[1325,1361,378,396]" gridcol="6" gridrow="5" pageId="8" pageNumber="9">1.7</td>
|
||
</tr>
|
||
<tr box="[138,1361,415,433]" gridrow="6" pageId="8" pageNumber="9">
|
||
<th box="[138,358,415,433]" gridcol="0" gridrow="6" pageId="8" pageNumber="9">Predorsal distance</th>
|
||
<td box="[453,535,415,433]" gridcol="1" gridrow="6" pageId="8" pageNumber="9">44.3</td>
|
||
<td box="[631,716,415,433]" gridcol="2" gridrow="6" pageId="8" pageNumber="9">44.8</td>
|
||
<td box="[811,900,415,433]" gridcol="3" gridrow="6" pageId="8" pageNumber="9">49.5</td>
|
||
<td box="[995,1084,415,433]" gridcol="4" gridrow="6" pageId="8" pageNumber="9">
|
||
44.8
|
||
<emphasis bold="true" box="[1034,1045,415,433]" pageId="8" pageNumber="9">–</emphasis>
|
||
49.5
|
||
</td>
|
||
<td box="[1180,1231,415,433]" gridcol="5" gridrow="6" pageId="8" pageNumber="9">46.8</td>
|
||
<td box="[1325,1361,415,433]" gridcol="6" gridrow="6" pageId="8" pageNumber="9">1.4</td>
|
||
</tr>
|
||
<tr box="[138,1361,452,470]" gridrow="7" pageId="8" pageNumber="9">
|
||
<th box="[138,358,452,470]" gridcol="0" gridrow="7" pageId="8" pageNumber="9">Prepelvic distance</th>
|
||
<td box="[453,535,452,470]" gridcol="1" gridrow="7" pageId="8" pageNumber="9">49.5</td>
|
||
<td box="[631,716,452,470]" gridcol="2" gridrow="7" pageId="8" pageNumber="9">52.1</td>
|
||
<td box="[811,900,452,470]" gridcol="3" gridrow="7" pageId="8" pageNumber="9">56.3</td>
|
||
<td box="[995,1084,452,470]" gridcol="4" gridrow="7" pageId="8" pageNumber="9">
|
||
52.1
|
||
<emphasis bold="true" box="[1034,1045,452,470]" pageId="8" pageNumber="9">–</emphasis>
|
||
56.3
|
||
</td>
|
||
<td box="[1180,1231,452,470]" gridcol="5" gridrow="7" pageId="8" pageNumber="9">53.7</td>
|
||
<td box="[1325,1361,452,470]" gridcol="6" gridrow="7" pageId="8" pageNumber="9">1.6</td>
|
||
</tr>
|
||
<tr box="[138,1361,490,508]" gridrow="8" pageId="8" pageNumber="9">
|
||
<th box="[138,358,490,508]" gridcol="0" gridrow="8" pageId="8" pageNumber="9">Preanal distance</th>
|
||
<td box="[453,535,490,508]" gridcol="1" gridrow="8" pageId="8" pageNumber="9">71.1</td>
|
||
<td box="[631,716,490,508]" gridcol="2" gridrow="8" pageId="8" pageNumber="9">74.9</td>
|
||
<td box="[811,900,490,508]" gridcol="3" gridrow="8" pageId="8" pageNumber="9">80.3</td>
|
||
<td box="[995,1084,490,508]" gridcol="4" gridrow="8" pageId="8" pageNumber="9">
|
||
74.9
|
||
<emphasis bold="true" box="[1034,1045,490,508]" pageId="8" pageNumber="9">–</emphasis>
|
||
80.3
|
||
</td>
|
||
<td box="[1180,1231,490,508]" gridcol="5" gridrow="8" pageId="8" pageNumber="9">76.7</td>
|
||
<td box="[1325,1361,490,508]" gridcol="6" gridrow="8" pageId="8" pageNumber="9">1.8</td>
|
||
</tr>
|
||
<tr box="[138,1361,527,545]" gridrow="9" pageId="8" pageNumber="9">
|
||
<th box="[138,358,527,545]" gridcol="0" gridrow="9" pageId="8" pageNumber="9">Anal-apex distance</th>
|
||
<td box="[453,535,527,545]" gridcol="1" gridrow="9" pageId="8" pageNumber="9">77.9</td>
|
||
<td box="[631,716,527,545]" gridcol="2" gridrow="9" pageId="8" pageNumber="9">82.1</td>
|
||
<td box="[811,900,527,545]" gridcol="3" gridrow="9" pageId="8" pageNumber="9">88.6</td>
|
||
<td box="[995,1084,527,545]" gridcol="4" gridrow="9" pageId="8" pageNumber="9">
|
||
82.1
|
||
<emphasis bold="true" box="[1034,1045,527,545]" pageId="8" pageNumber="9">–</emphasis>
|
||
88.6
|
||
</td>
|
||
<td box="[1180,1231,527,545]" gridcol="5" gridrow="9" pageId="8" pageNumber="9">85</td>
|
||
<td box="[1325,1361,527,545]" gridcol="6" gridrow="9" pageId="8" pageNumber="9">1.8</td>
|
||
</tr>
|
||
<tr box="[138,1361,564,582]" gridrow="10" pageId="8" pageNumber="9">
|
||
<th box="[138,358,564,582]" gridcol="0" gridrow="10" pageId="8" pageNumber="9">Body width</th>
|
||
<td box="[453,535,564,582]" gridcol="1" gridrow="10" pageId="8" pageNumber="9">12.4</td>
|
||
<td box="[631,716,564,582]" gridcol="2" gridrow="10" pageId="8" pageNumber="9">11.2</td>
|
||
<td box="[811,900,564,582]" gridcol="3" gridrow="10" pageId="8" pageNumber="9">12.9</td>
|
||
<td box="[995,1084,564,582]" gridcol="4" gridrow="10" pageId="8" pageNumber="9">
|
||
11.2
|
||
<emphasis bold="true" box="[1034,1045,564,582]" pageId="8" pageNumber="9">–</emphasis>
|
||
12.9
|
||
</td>
|
||
<td box="[1180,1231,564,582]" gridcol="5" gridrow="10" pageId="8" pageNumber="9">11.9</td>
|
||
<td box="[1325,1361,564,582]" gridcol="6" gridrow="10" pageId="8" pageNumber="9">0.7</td>
|
||
</tr>
|
||
<tr box="[138,1361,601,620]" gridrow="11" pageId="8" pageNumber="9">
|
||
<th box="[138,358,601,620]" gridcol="0" gridrow="11" pageId="8" pageNumber="9">Body depth at DO</th>
|
||
<td box="[453,535,601,620]" gridcol="1" gridrow="11" pageId="8" pageNumber="9">22.4</td>
|
||
<td box="[631,716,601,620]" gridcol="2" gridrow="11" pageId="8" pageNumber="9">20.7</td>
|
||
<td box="[811,900,601,620]" gridcol="3" gridrow="11" pageId="8" pageNumber="9">24.6</td>
|
||
<td box="[995,1084,601,620]" gridcol="4" gridrow="11" pageId="8" pageNumber="9">
|
||
20.7
|
||
<emphasis bold="true" box="[1034,1045,602,620]" pageId="8" pageNumber="9">–</emphasis>
|
||
24.6
|
||
</td>
|
||
<td box="[1180,1231,601,620]" gridcol="5" gridrow="11" pageId="8" pageNumber="9">22.3</td>
|
||
<td box="[1325,1361,601,620]" gridcol="6" gridrow="11" pageId="8" pageNumber="9">1.5</td>
|
||
</tr>
|
||
<tr box="[138,1361,638,657]" gridrow="12" pageId="8" pageNumber="9">
|
||
<th box="[138,358,638,657]" gridcol="0" gridrow="12" pageId="8" pageNumber="9">Body depth at AO</th>
|
||
<td box="[453,535,638,657]" gridcol="1" gridrow="12" pageId="8" pageNumber="9">17.3</td>
|
||
<td box="[631,716,638,657]" gridcol="2" gridrow="12" pageId="8" pageNumber="9">15.7</td>
|
||
<td box="[811,900,638,657]" gridcol="3" gridrow="12" pageId="8" pageNumber="9">18.2</td>
|
||
<td box="[995,1084,638,657]" gridcol="4" gridrow="12" pageId="8" pageNumber="9">
|
||
15.7
|
||
<emphasis bold="true" box="[1034,1045,639,657]" pageId="8" pageNumber="9">–</emphasis>
|
||
18.2
|
||
</td>
|
||
<td box="[1180,1231,638,657]" gridcol="5" gridrow="12" pageId="8" pageNumber="9">16.9</td>
|
||
<td box="[1325,1361,638,657]" gridcol="6" gridrow="12" pageId="8" pageNumber="9">1</td>
|
||
</tr>
|
||
<tr box="[138,1361,676,695]" gridrow="13" pageId="8" pageNumber="9">
|
||
<th box="[138,358,676,695]" gridcol="0" gridrow="13" pageId="8" pageNumber="9">Caudal peduncle depth</th>
|
||
<td box="[453,535,676,695]" gridcol="1" gridrow="13" pageId="8" pageNumber="9">11.1</td>
|
||
<td box="[631,716,676,695]" gridcol="2" gridrow="13" pageId="8" pageNumber="9">10.6</td>
|
||
<td box="[811,900,676,695]" gridcol="3" gridrow="13" pageId="8" pageNumber="9">12</td>
|
||
<td box="[995,1084,676,695]" gridcol="4" gridrow="13" pageId="8" pageNumber="9">
|
||
10.6
|
||
<emphasis bold="true" box="[1050,1061,676,694]" pageId="8" pageNumber="9">–</emphasis>
|
||
12
|
||
</td>
|
||
<td box="[1180,1231,676,695]" gridcol="5" gridrow="13" pageId="8" pageNumber="9">11.4</td>
|
||
<td box="[1325,1361,676,695]" gridcol="6" gridrow="13" pageId="8" pageNumber="9">0.5</td>
|
||
</tr>
|
||
<tr box="[138,1361,714,732]" gridrow="14" pageId="8" pageNumber="9" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
||
<th box="[138,358,714,732]" gridcol="0" gridrow="14" pageId="8" pageNumber="9">% HL</th>
|
||
</tr>
|
||
<tr box="[138,1361,750,769]" gridrow="15" pageId="8" pageNumber="9">
|
||
<th box="[138,358,750,769]" gridcol="0" gridrow="15" pageId="8" pageNumber="9">Snout length</th>
|
||
<td box="[453,535,750,769]" gridcol="1" gridrow="15" pageId="8" pageNumber="9">22.8</td>
|
||
<td box="[631,716,750,769]" gridcol="2" gridrow="15" pageId="8" pageNumber="9">23</td>
|
||
<td box="[811,900,750,769]" gridcol="3" gridrow="15" pageId="8" pageNumber="9">25.7</td>
|
||
<td box="[995,1084,750,769]" gridcol="4" gridrow="15" pageId="8" pageNumber="9">
|
||
23
|
||
<emphasis bold="true" box="[1034,1045,751,769]" pageId="8" pageNumber="9">–</emphasis>
|
||
25.7
|
||
</td>
|
||
<td box="[1180,1231,750,769]" gridcol="5" gridrow="15" pageId="8" pageNumber="9">24.2</td>
|
||
<td box="[1325,1361,750,769]" gridcol="6" gridrow="15" pageId="8" pageNumber="9">1</td>
|
||
</tr>
|
||
<tr box="[138,1361,788,807]" gridrow="16" pageId="8" pageNumber="9">
|
||
<th box="[138,358,788,807]" gridcol="0" gridrow="16" pageId="8" pageNumber="9">Snout-maxillary tip</th>
|
||
<td box="[453,535,788,807]" gridcol="1" gridrow="16" pageId="8" pageNumber="9">25.6</td>
|
||
<td box="[631,716,788,807]" gridcol="2" gridrow="16" pageId="8" pageNumber="9">23.2</td>
|
||
<td box="[811,900,788,807]" gridcol="3" gridrow="16" pageId="8" pageNumber="9">27.1</td>
|
||
<td box="[995,1084,788,807]" gridcol="4" gridrow="16" pageId="8" pageNumber="9">
|
||
23.2
|
||
<emphasis bold="true" box="[1034,1045,788,806]" pageId="8" pageNumber="9">–</emphasis>
|
||
27.1
|
||
</td>
|
||
<td box="[1180,1231,788,807]" gridcol="5" gridrow="16" pageId="8" pageNumber="9">24.9</td>
|
||
<td box="[1325,1361,788,807]" gridcol="6" gridrow="16" pageId="8" pageNumber="9">1.2</td>
|
||
</tr>
|
||
<tr box="[138,1361,826,844]" gridrow="17" pageId="8" pageNumber="9">
|
||
<th box="[138,358,826,844]" gridcol="0" gridrow="17" pageId="8" pageNumber="9">Anterior naris-eye</th>
|
||
<td box="[453,535,826,844]" gridcol="1" gridrow="17" pageId="8" pageNumber="9">9.7</td>
|
||
<td box="[631,716,826,844]" gridcol="2" gridrow="17" pageId="8" pageNumber="9">7.1</td>
|
||
<td box="[811,900,826,844]" gridcol="3" gridrow="17" pageId="8" pageNumber="9">9.5</td>
|
||
<td box="[995,1084,826,844]" gridcol="4" gridrow="17" pageId="8" pageNumber="9">
|
||
7.1
|
||
<emphasis bold="true" box="[1045,1056,826,844]" pageId="8" pageNumber="9">–</emphasis>
|
||
9.5
|
||
</td>
|
||
<td box="[1180,1231,826,844]" gridcol="5" gridrow="17" pageId="8" pageNumber="9">8.5</td>
|
||
<td box="[1325,1361,826,844]" gridcol="6" gridrow="17" pageId="8" pageNumber="9">0.8</td>
|
||
</tr>
|
||
<tr box="[138,1361,863,881]" gridrow="18" pageId="8" pageNumber="9">
|
||
<th box="[138,358,863,881]" gridcol="0" gridrow="18" pageId="8" pageNumber="9">Posterior naris - eye</th>
|
||
<td box="[453,535,863,881]" gridcol="1" gridrow="18" pageId="8" pageNumber="9">5.1</td>
|
||
<td box="[631,716,863,881]" gridcol="2" gridrow="18" pageId="8" pageNumber="9">1.8</td>
|
||
<td box="[811,900,863,881]" gridcol="3" gridrow="18" pageId="8" pageNumber="9">5.8</td>
|
||
<td box="[995,1084,863,881]" gridcol="4" gridrow="18" pageId="8" pageNumber="9">
|
||
1.8
|
||
<emphasis bold="true" box="[1045,1056,863,881]" pageId="8" pageNumber="9">–</emphasis>
|
||
5.8
|
||
</td>
|
||
<td box="[1180,1231,863,881]" gridcol="5" gridrow="18" pageId="8" pageNumber="9">4</td>
|
||
<td box="[1325,1361,863,881]" gridcol="6" gridrow="18" pageId="8" pageNumber="9">1.3</td>
|
||
</tr>
|
||
<tr box="[138,1361,900,919]" gridrow="19" pageId="8" pageNumber="9">
|
||
<th box="[138,358,900,919]" gridcol="0" gridrow="19" pageId="8" pageNumber="9">Cheek</th>
|
||
<td box="[453,535,900,919]" gridcol="1" gridrow="19" pageId="8" pageNumber="9">21.8</td>
|
||
<td box="[631,716,900,919]" gridcol="2" gridrow="19" pageId="8" pageNumber="9">19.7</td>
|
||
<td box="[811,900,900,919]" gridcol="3" gridrow="19" pageId="8" pageNumber="9">22,6</td>
|
||
<td box="[995,1084,900,919]" gridcol="4" gridrow="19" pageId="8" pageNumber="9">
|
||
19.7
|
||
<emphasis bold="true" box="[1034,1045,900,918]" pageId="8" pageNumber="9">–</emphasis>
|
||
22.6
|
||
</td>
|
||
<td box="[1180,1231,900,919]" gridcol="5" gridrow="19" pageId="8" pageNumber="9">20.9</td>
|
||
<td box="[1325,1361,900,919]" gridcol="6" gridrow="19" pageId="8" pageNumber="9">1</td>
|
||
</tr>
|
||
<tr box="[138,1361,937,956]" gridrow="20" pageId="8" pageNumber="9">
|
||
<th box="[138,358,937,956]" gridcol="0" gridrow="20" pageId="8" pageNumber="9">Orbital diameter</th>
|
||
<td box="[453,535,937,956]" gridcol="1" gridrow="20" pageId="8" pageNumber="9">26.3</td>
|
||
<td box="[631,716,937,956]" gridcol="2" gridrow="20" pageId="8" pageNumber="9">22.5</td>
|
||
<td box="[811,900,937,956]" gridcol="3" gridrow="20" pageId="8" pageNumber="9">29.2</td>
|
||
<td box="[995,1084,937,956]" gridcol="4" gridrow="20" pageId="8" pageNumber="9">
|
||
22.5
|
||
<emphasis bold="true" box="[1034,1045,938,956]" pageId="8" pageNumber="9">–</emphasis>
|
||
29.2
|
||
</td>
|
||
<td box="[1180,1231,937,956]" gridcol="5" gridrow="20" pageId="8" pageNumber="9">25.7</td>
|
||
<td box="[1325,1361,937,956]" gridcol="6" gridrow="20" pageId="8" pageNumber="9">2</td>
|
||
</tr>
|
||
<tr box="[138,1361,975,993]" gridrow="21" pageId="8" pageNumber="9">
|
||
<th box="[138,358,975,993]" gridcol="0" gridrow="21" pageId="8" pageNumber="9">Interorbital distance</th>
|
||
<td box="[453,535,975,993]" gridcol="1" gridrow="21" pageId="8" pageNumber="9">11.3</td>
|
||
<td box="[631,716,975,993]" gridcol="2" gridrow="21" pageId="8" pageNumber="9">8,3</td>
|
||
<td box="[811,900,975,993]" gridcol="3" gridrow="21" pageId="8" pageNumber="9">15.5</td>
|
||
<td box="[995,1084,975,993]" gridcol="4" gridrow="21" pageId="8" pageNumber="9">
|
||
8.3
|
||
<emphasis bold="true" box="[1034,1045,975,993]" pageId="8" pageNumber="9">–</emphasis>
|
||
15,5
|
||
</td>
|
||
<td box="[1180,1231,975,993]" gridcol="5" gridrow="21" pageId="8" pageNumber="9">12.8</td>
|
||
<td box="[1325,1361,975,993]" gridcol="6" gridrow="21" pageId="8" pageNumber="9">2.7</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="8.[119,1217,1018,1038]" box="[119,1217,1018,1038]" pageId="8" pageNumber="9">
|
||
<tableNote box="[119,1217,1018,1038]" pageId="8" pageNumber="9" targetBox="[138,1361,186,993]" targetPageId="8">Abbreviations: DO, dorsal origin; AO, anal origin; HL, head length; S.D; standard deviation; LS, standard length; TL, total length.</tableNote>
|
||
</paragraph>
|
||
<paragraph blockId="8.[119,768,1113,1757]" pageId="8" pageNumber="9">Dorsal-fin rays ii7(1), 9*(8); anal-fin rays i6(7), 7*(2); pectoral-fin rays iii7 (1), 8*(6), 9(2); pelvic-fin rays i6(1), 7*(8); principal caudal-fin rays in dorsal lobe 9*(9), in ventral lobe 8*(9).</paragraph>
|
||
<paragraph blockId="8.[119,768,1113,1757]" pageId="8" pageNumber="9">Scales cycloid. Lateral line complete; pored lateral-line scales 30(2), 31*(2), 32(3), 33(2) Scales above lateral line 4*(9), below 2*(9). Predorsal scales 8*(4), 9(5). Circumpeduncular scales 11*(2), 12*(7). Isthmus and area between the pectoral with scales.</paragraph>
|
||
<paragraph blockId="8.[119,768,1113,1757]" pageId="8" pageNumber="9">
|
||
Mesethmoid bone separated from frontal, parietal fontanel reduced width size of the post-parietal bars, postcleitrum (PC) I absent but PC II and III present. Medial process of first rib with ventral vertex paddle-shaped, wider in vertex which decreases ventrally to tip of rib, dorsal upper process has an L shape without concave space between process and base to vertebral join (
|
||
<figureCitation box="[448,540,1633,1653]" captionStart="FIGURE 8" captionStartId="7.[1107,1198,141,160]" captionTargetBox="[125,1057,139,619]" captionTargetId="figure-619@7.[124,1062,138,622]" captionText="FIGURE 8 Detail in the shape of the first rib (a)–(c) and swimbladder (b)–(d) in (a) and (b). C. tatama n. sp. (b, ventral view) IMCN 8924 (1C&S) and (c) and (d) C. dule n. sp. (d, dorsal view) IMCN 8929 (1C&S). The right lobule in (d) is damaged. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.4058149" httpUri="https://zenodo.org/record/4058149/files/figure.png" pageId="8" pageNumber="9">Figure 8c</figureCitation>
|
||
.); swimbladder with two chambers, anterior bilobulated and laterally wider than that of
|
||
<taxonomicName authorityName="Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara" authorityYear="2020" box="[119,214,1702,1722]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="tatama">
|
||
<emphasis box="[119,214,1702,1722]" italics="true" pageId="8" pageNumber="9">C. tatama</emphasis>
|
||
</taxonomicName>
|
||
, posterior chamber shorter than anterior chamber and rounded (
|
||
<figureCitation box="[207,298,1737,1757]" captionStart="FIGURE 8" captionStartId="7.[1107,1198,141,160]" captionTargetBox="[125,1057,139,619]" captionTargetId="figure-619@7.[124,1062,138,622]" captionText="FIGURE 8 Detail in the shape of the first rib (a)–(c) and swimbladder (b)–(d) in (a) and (b). C. tatama n. sp. (b, ventral view) IMCN 8924 (1C&S) and (c) and (d) C. dule n. sp. (d, dorsal view) IMCN 8929 (1C&S). The right lobule in (d) is damaged. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.4058149" httpUri="https://zenodo.org/record/4058149/files/figure.png" pageId="8" pageNumber="9">Figure 8d</figureCitation>
|
||
.).
|
||
</paragraph>
|
||
<paragraph blockId="8.[119,527,1837,1862]" box="[119,527,1837,1862]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[119,171,1837,1862]" pageId="8" pageNumber="9">3.17</emphasis>
|
||
|
|
||
<heading bold="true" box="[241,527,1837,1862]" fontSize="10" level="3" pageId="8" pageNumber="9" reason="0">
|
||
<emphasis bold="true" box="[241,527,1837,1862]" pageId="8" pageNumber="9">Colour in live specimens</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="8.[119,767,1910,1965]" lastBlockId="8.[816,1465,1113,1306]" pageId="8" pageNumber="9">
|
||
Mature specimens purplish. This colour is observed predominantly on dorsal, anal, adipose, caudal and pelvic fins, extending also over ventral area (
|
||
<figureCitation box="[938,1017,1113,1133]" captionStart="FIGURE 9" captionStartId="8.[816,907,1916,1935]" captionTargetBox="[820,1462,1682,1873]" captionTargetId="figure-574@8.[818,1463,1681,1875]" captionTargetPageId="8" captionText="FIGURE 9 C. dule n. sp. IMCN 8929, Paratype. Live specimen. Scale bar = 1 cm" figureDoi="http://doi.org/10.5281/zenodo.4058151" httpUri="https://zenodo.org/record/4058151/files/figure.png" pageId="8" pageNumber="9">Figure 9</figureCitation>
|
||
). Lateral line with golden shades predominantly between pelvic and anal fins. Head, with golden line behind eye, followed by a black line and golden point at end of opercle, as well as a black humeral blotch. First portion of dorsal fin (first two rays) golden in colour. Head with a band that is smaller than pupil, extending from front edge of eye to beginning of maxillary bone.
|
||
</paragraph>
|
||
<paragraph blockId="8.[816,1136,1386,1411]" box="[816,1136,1386,1411]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[816,868,1386,1411]" pageId="8" pageNumber="9">3.18</emphasis>
|
||
|
|
||
<heading bold="true" box="[938,1136,1386,1411]" fontSize="10" level="3" pageId="8" pageNumber="9" reason="0">
|
||
<emphasis bold="true" box="[938,1136,1386,1411]" pageId="8" pageNumber="9">Colour in alcohol</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="8.[816,1465,1459,1618]" lastBlockId="9.[124,772,142,266]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
Mature specimens purple with 7
|
||
<emphasis bold="true" box="[1114,1126,1460,1479]" pageId="8" pageNumber="9">–</emphasis>
|
||
8 bars, with highly pigmented dorsal, anal and caudal fins (
|
||
<figureCitation box="[1010,1097,1494,1514]" captionStart="FIGURE 6" captionStartId="6.[119,210,1916,1935]" captionTargetBox="[122,765,1235,1875]" captionTargetId="figure-512@6.[122,765,1235,1875]" captionText="FIGURE 6 C. dule n. sp. IMCN 8941, Holotype 37.4 mm LS. Scale bar = 1 cm" figureDoi="http://doi.org/10.5281/zenodo.4058143" httpUri="https://zenodo.org/record/4058143/files/figure.png" pageId="8" pageNumber="9">Figures 6</figureCitation>
|
||
and
|
||
<figureCitation box="[1144,1158,1494,1514]" captionStart="FIGURE 8" captionStartId="7.[1107,1198,141,160]" captionTargetBox="[125,1057,139,619]" captionTargetId="figure-619@7.[124,1062,138,622]" captionText="FIGURE 8 Detail in the shape of the first rib (a)–(c) and swimbladder (b)–(d) in (a) and (b). C. tatama n. sp. (b, ventral view) IMCN 8924 (1C&S) and (c) and (d) C. dule n. sp. (d, dorsal view) IMCN 8929 (1C&S). The right lobule in (d) is damaged. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.4058149" httpUri="https://zenodo.org/record/4058149/files/figure.png" pageId="8" pageNumber="9">8</figureCitation>
|
||
). In all the specimens, transverse band present and smaller in size (1/2 scale) than humeral and caudal spots. Unmarked belly. Humeral blotch reaching two to three scales and concave extension towards pectoral fin, being faint in immature specimens and dark in mature specimens and coming into contact with last rays of the pectoral fin (
|
||
<figureCitation box="[436,515,177,197]" captionStart="FIGURE 7" captionStartId="6.[816,907,1916,1935]" captionTargetBox="[874,1405,1414,1876]" captionTargetId="figure-537@6.[874,1405,1414,1876]" captionText="FIGURE 7 Detail of the composition of spots in C. dule n. sp. IMCN 8929, Paratype 38.24 mm LS, arrows show the detail. Scale bar = 1 cm" figureDoi="http://doi.org/10.5281/zenodo.4058147" httpUri="https://zenodo.org/record/4058147/files/figure.png" pageId="9" pageNumber="10">Figure 7</figureCitation>
|
||
). Dorsal anterior portion of maxilla pigmented forming a line reaching the eye, ventral posterior portion of maxilla without pigment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4058151" ID-Zenodo-Dep="4058151" httpUri="https://zenodo.org/record/4058151/files/figure.png" pageId="8" pageNumber="9" startId="8.[816,907,1916,1935]" targetBox="[820,1462,1682,1873]" targetPageId="8">
|
||
<paragraph blockId="8.[816,1427,1915,1965]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[816,928,1916,1935]" pageId="8" pageNumber="9">FIGURE 9</emphasis>
|
||
<taxonomicName authority="Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara, 2020" authorityName="Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara" authorityYear="2020" box="[958,1018,1915,1935]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="dule" status="n. sp.">
|
||
<emphasis box="[958,1018,1915,1935]" italics="true" pageId="8" pageNumber="9">C. dule</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1023,1073,1915,1935]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
IMCN 8929, Paratype. Live specimen. Scale bar = 1 cm
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="9" pageNumber="10" type="biology_ecology">
|
||
<paragraph blockId="9.[124,176,346,371]" box="[124,176,346,371]" pageId="9" pageNumber="10">
|
||
<emphasis bold="true" box="[124,176,346,371]" pageId="9" pageNumber="10">3.19</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="9.[207,214,346,371]" box="[207,214,346,371]" pageId="9" pageNumber="10">|</paragraph>
|
||
<paragraph blockId="9.[246,339,346,371]" box="[246,339,346,371]" pageId="9" pageNumber="10">
|
||
<heading bold="true" box="[246,339,346,371]" fontSize="10" level="3" pageId="9" pageNumber="10" reason="0">
|
||
<emphasis bold="true" box="[246,339,346,371]" pageId="9" pageNumber="10">Ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="9.[124,772,419,578]" pageId="9" pageNumber="10">
|
||
The Arquía River (
|
||
<typeStatus box="[291,333,419,439]" pageId="9" pageNumber="10">type</typeStatus>
|
||
locality) is a tributary of steep banks with large slopes, with good riparian coverage on both flanks and a predominance of a rocky substrate with large edges, which allow the formation of backwaters and small pools where sand and organic materials are deposited. A tributary of transparent waters (Figure 10).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="10" type="distribution">
|
||
<paragraph blockId="9.[124,176,658,683]" box="[124,176,658,683]" pageId="9" pageNumber="10">
|
||
<emphasis bold="true" box="[124,176,658,683]" pageId="9" pageNumber="10">3.20</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="9.[207,214,658,683]" box="[207,214,658,683]" pageId="9" pageNumber="10">|</paragraph>
|
||
<paragraph blockId="9.[246,388,658,683]" box="[246,388,658,683]" pageId="9" pageNumber="10">
|
||
<heading bold="true" box="[246,388,658,683]" fontSize="10" level="3" pageId="9" pageNumber="10" reason="0">
|
||
<emphasis bold="true" box="[246,388,658,683]" pageId="9" pageNumber="10">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="9.[124,772,731,821]" pageId="9" pageNumber="10">
|
||
<taxonomicName authorityName="Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara" authorityYear="2020" box="[124,279,731,751]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="dule">
|
||
<emphasis box="[124,279,731,751]" italics="true" pageId="9" pageNumber="10">Characidium dule</emphasis>
|
||
</taxonomicName>
|
||
is endemic to the
|
||
<collectingRegion box="[454,514,731,751]" country="Panama" name="Embera-Wounaan" pageId="9" pageNumber="10">Chocó</collectingRegion>
|
||
biogeographical region and inhabits rivers or streams tributaries to the Atrato River, from its upper reaches to near the mouth into the Urabá Gulf (Figure 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="10" type="etymology">
|
||
<paragraph blockId="9.[124,176,901,926]" box="[124,176,901,926]" pageId="9" pageNumber="10">
|
||
<emphasis bold="true" box="[124,176,901,926]" pageId="9" pageNumber="10">3.21</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="9.[207,214,901,926]" box="[207,214,901,926]" pageId="9" pageNumber="10">|</paragraph>
|
||
<paragraph blockId="9.[246,372,901,926]" box="[246,372,901,926]" pageId="9" pageNumber="10">
|
||
<heading bold="true" box="[246,372,901,926]" fontSize="10" level="3" pageId="9" pageNumber="10" reason="0">
|
||
<emphasis bold="true" box="[246,372,901,926]" pageId="9" pageNumber="10">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="9.[124,773,974,1098]" pageId="9" pageNumber="10">
|
||
A noun in apposition dedicated to the Dule (Kuna) ethnic group of Maggilagundiwala or Resguardo de Arquía in the Colombian
|
||
<collectingRegion box="[707,769,1009,1029]" country="Panama" name="Embera-Wounaan" pageId="9" pageNumber="10">Chocó</collectingRegion>
|
||
, whose language is under a high risk of extinction and where most of the
|
||
<typeStatus box="[160,201,1078,1098]" pageId="9" pageNumber="10">type</typeStatus>
|
||
specimens were collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="10" type="conservation">
|
||
<paragraph blockId="9.[124,176,1178,1203]" box="[124,545,1178,1203]" lastBlockId="9.[246,545,1178,1203]" pageId="9" pageNumber="10">
|
||
<emphasis bold="true" box="[124,176,1178,1203]" pageId="9" pageNumber="10">3.22</emphasis>
|
||
|
|
||
<heading bold="true" box="[246,545,1178,1203]" fontSize="10" level="3" pageId="9" pageNumber="10" reason="0">
|
||
<emphasis bold="true" box="[246,545,1178,1203]" pageId="9" pageNumber="10">Conservation assessment</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="9.[124,772,1251,1410]" lastBlockId="9.[821,1469,142,335]" pageId="9" pageNumber="10">
|
||
<taxonomicName authorityName="Agudelo‐Zamora & Tavera & Murillo & Ortega‐Lara" authorityYear="2020" box="[124,280,1251,1271]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="dule">
|
||
<emphasis box="[124,280,1251,1271]" italics="true" pageId="9" pageNumber="10">Characidium dule</emphasis>
|
||
</taxonomicName>
|
||
is so far known from two localities in tributaries of the Atrato River. The EOO estimated for the species is 5.701.
|
||
<quantity box="[710,758,1286,1306]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="km" value="2.0">2 km</quantity>
|
||
<superScript attach="left" box="[758,767,1282,1295]" fontSize="5" pageId="9" pageNumber="10">2</superScript>
|
||
, and the AOO estimated is
|
||
<quantity box="[373,432,1321,1341]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="10" unit="km" value="21.0">21 km</quantity>
|
||
<superScript attach="left" box="[431,440,1316,1329]" fontSize="5" pageId="9" pageNumber="10">2</superScript>
|
||
(
|
||
<emphasis box="[453,502,1321,1341]" italics="true" pageId="9" pageNumber="10">sensu</emphasis>
|
||
<bibRefCitation author="Bachman, S. & Moat, J. & Hill, A. W. & de la Torre, J. & Scott, B." box="[508,706,1321,1341]" journalOrPublisher="ZooKeys" pageId="9" pageNumber="10" pagination="117 - 126" part="150" refId="ref10245" refString="Bachman, S., Moat, J., Hill, A. W., de la Torre, J., & Scott, B. (2011). Supporting red list threat assessments with GeoCAT: geospatial conservation assessment tool. ZooKeys, 150, 117 - 126." title="Supporting red list threat assessments with GeoCAT: geospatial conservation assessment tool" type="journal article" year="2011">
|
||
Bachman
|
||
<emphasis box="[600,640,1321,1341]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
|
||
., 2011
|
||
</bibRefCitation>
|
||
). Additional collecting efforts should be conducted in the region to better understand the geographic distribution of the species and the existent threats. From the points assessed here for the distribution of
|
||
<taxonomicName box="[1402,1464,142,162]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="dule">
|
||
<emphasis box="[1402,1464,142,162]" italics="true" pageId="9" pageNumber="10">C. dule</emphasis>
|
||
</taxonomicName>
|
||
, the species is tentatively categorized as Vulnerable (VU) D2 under the geospatial aspect because the population is restricted to less than
|
||
<quantity box="[821,879,246,266]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="km" value="20.0">20 km</quantity>
|
||
<superScript attach="left" box="[879,888,242,255]" fontSize="5" pageId="9" pageNumber="10">2</superScript>
|
||
and the number of localities is less than five (
|
||
<collectionCode box="[1318,1370,246,266]" pageId="9" pageNumber="10">IUCN</collectionCode>
|
||
Standards and Petitions Subcommittee, 2019), with accelerated deforestation inside Colombian national parks (
|
||
<bibRefCitation author="Clerici, N. & Armenteras, D. & Kareiva, P. & Botero, R. & Ramirez-Delgado, J. P. & Forero-Medina, G." box="[1124,1292,315,335]" journalOrPublisher="Scientific Reports" pageId="9" pageNumber="10" pagination="1 - 10" part="10" refId="ref10585" refString="Clerici, N., Armenteras, D., Kareiva, P., Botero, R., Ramirez-Delgado, J. P., Forero-Medina, G., et al. (2020). Deforestation in Colombian protected areas increased duringpost-conflict periods. Scientific Reports, 10 (1), 1 - 10." title="Deforestation in Colombian protected areas increased duringpost-conflict periods" type="journal article" year="2020">
|
||
Clerici
|
||
<emphasis box="[1188,1226,315,335]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
|
||
., 2020
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |