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<document id="DE92C48898AD11EE805B96965F61BA65" ID-DOI="10.11646/zootaxa.3990.3.1" ID-GBIF-Dataset="cf747765-b9bf-42ed-8479-35355e712db0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244301" ID-ZooBank="378C0359-E0E4-4CCC-821D-649144E37A63" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461268581225" checkinUser="plazi" docAuthor="Toledo, Mario &amp; Michat, Mariano C." docDate="2015" docId="03EE87FEFFE4FFEC91A6F889FEE2A4FB" docLanguage="en" docName="zt03990p354.pdf" docOrigin="Zootaxa 3990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccomimus youngi Toledo &amp; Michat, 2015, sp. n." docType="treatment" docVersion="9" lastPageNumber="325" masterDocId="FFD7FF86FFF3FFF49131FFBDFFF8A03E" masterDocTitle="Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)" masterLastPageNumber="354" masterPageNumber="301" pageNumber="324" updateTime="1698611165842" updateUser="plazi">
<mods:mods id="19A8E41D48D75870CF349C88F9088D31" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="73ACC12CF47E35DA30474A46FCBB7FB0">
<mods:title id="4F08BCE00EAC3E3E3748B7B08F27E158">Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)</mods:title>
</mods:titleInfo>
<mods:name id="D327D7FBE92645B74F8A1638397E98DC" type="personal">
<mods:role id="535AF72508EBAB7EB57EE74989365D52">
<mods:roleTerm id="6631AEDC21DA95E0A7C1E8CA3D37507B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF2FB3A57680318D161BBC96533181A2">Toledo, Mario</mods:namePart>
</mods:name>
<mods:name id="6FFC3D6E3E4022E99C5D65E740DC8100" type="personal">
<mods:role id="F17EE7250155BC5FC60C55E6864893A5">
<mods:roleTerm id="3894D60B8F6738D86F37E6F38F954843">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A5473E4852AE2DDA95F7D4688DAD255E">Michat, Mariano C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CEA5ACAC46D6A4EA175F72C686D6E9CC">text</mods:typeOfResource>
<mods:relatedItem id="88B9D7C6C829D03DA1FD2AA5613E607C" type="host">
<mods:titleInfo id="05331CA9C4AE1AD69F14FED812CF18E4">
<mods:title id="5AC3360496436887F337B49372F96DAE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="92442E6EF1DEB6963DFB8A8C642D2363">
<mods:date id="541B090FE57CF1E0AFDC4A5A2C4B913A">2015</mods:date>
<mods:detail id="73881E00D36FC3BBA485E5A0DC32B64C" type="volume">
<mods:number id="E0BAB203B7094B0BEF26BEB19831D5D0">3990</mods:number>
</mods:detail>
<mods:detail id="FDE9E37AC3E033342D8632C26C1E3521" type="issue">
<mods:number id="EECAA0C79E1D5EDA85E6F9E6172D5EAB">3</mods:number>
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<mods:extent id="3D17FE5CFF9A68ADEB077749E4F1DD09" unit="page">
<mods:start id="180BED3F9AC390A460C922893A0DB9EF">301</mods:start>
<mods:end id="880304CA71744864C346C90CF47C2CAC">354</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2AA9CCD4A35075C0415984B82089CC51">journal article</mods:classification>
<mods:identifier id="6F54B7A9EC4CAE8612F0EB6ACCC749DF" type="DOI">10.11646/zootaxa.3990.3.1</mods:identifier>
<mods:identifier id="D689C566A9642507F2A92913F8B42EFB" type="GBIF-Dataset">cf747765-b9bf-42ed-8479-35355e712db0</mods:identifier>
<mods:identifier id="6AC89CA63EB7B91489846C65B2CD5E0A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8406BEA12785A5574ABA4C527C4018BB" type="Zenodo-Dep">244301</mods:identifier>
<mods:identifier id="EA905F17CDC48D4634FA92F2F18A2259" type="ZooBank">378C0359-E0E4-4CCC-821D-649144E37A63</mods:identifier>
</mods:mods>
<treatment id="03EE87FEFFE4FFEC91A6F889FEE2A4FB" ID-DOI="http://doi.org/10.5281/zenodo.5692062" ID-GBIF-Taxon="119638444" ID-Zenodo-Dep="5692062" LSID="urn:lsid:plazi:treatment:03EE87FEFFE4FFEC91A6F889FEE2A4FB" httpUri="http://treatment.plazi.org/id/03EE87FEFFE4FFEC91A6F889FEE2A4FB" lastPageId="24" lastPageNumber="325" pageId="23" pageNumber="324">
<subSubSection id="C35D6563FFE4FFE391A6F889FEC9A751" pageId="23" pageNumber="324" type="nomenclature">
<paragraph id="8BF836E8FFE4FFE391A6F889FE2DA770" blockId="23.[151,469,1844,1903]" box="[151,469,1844,1870]" pageId="23" pageNumber="324">
<heading id="D0B08184FFE4FFE391A6F889FE2DA770" bold="true" box="[151,469,1844,1870]" fontSize="11" level="1" pageId="23" pageNumber="324" reason="1">
<emphasis id="B933EAFAFFE4FFE391A6F889FE2DA770" bold="true" box="[151,469,1844,1870]" pageId="23" pageNumber="324">
<taxonomicName id="4C474D6BFFE4FFE391A6F889FE74A770" ID-CoL="8LZJC" box="[151,396,1844,1870]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="youngi" status="sp. nov.">
<emphasis id="B933EAFAFFE4FFE391A6F889FE74A770" bold="true" box="[151,396,1844,1870]" italics="true" pageId="23" pageNumber="324">Laccomimus youngi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2005781FFE4FFE390A2F889FE2DA770" box="[403,469,1844,1870]" pageId="23" pageNumber="324" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF836E8FFE4FFE391A6F8EBFEC9A751" blockId="23.[151,469,1844,1903]" box="[151,305,1878,1903]" pageId="23" pageNumber="324">
(
<figureCitation id="137C2A6DFFE4FFE391AEF8EBFF0BA751" box="[159,243,1878,1903]" captionStart="FIGURES 88 – 90" captionStartId="41.[151,264,1919,1941]" captionTargetBox="[225,1366,198,1894]" captionTargetId="figure@41.[204,1370,193,1898]" captionTargetPageId="41" captionText="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." httpUri="https://zenodo.org/record/244314/files/figure.png" pageId="23" pageNumber="324">Fig. 90</figureCitation>
; 127)
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE4FFEC91A6F823FABAA15B" lastPageId="24" lastPageNumber="325" pageId="23" pageNumber="324" type="materials_examined">
<paragraph id="8BF836E8FFE4FFE391A6F823FC68A789" blockId="23.[151,912,1950,1975]" box="[151,912,1950,1975]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391A6F823FECDA789" bold="true" box="[151,309,1950,1975]" pageId="23" pageNumber="324">
<typeStatus id="54FC884AFFE4FFE391A6F823FF2AA789" box="[151,210,1950,1975]" pageId="23" pageNumber="324">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3507678FFE4FFE3900DF823FE7BA789" box="[316,387,1950,1975]" name="Brazil" pageId="23" pageNumber="324">Brazil</collectingCountry>
, Mato Grosso, Jacaré, Parque Nacional Xingu.
</paragraph>
<paragraph id="8BF836E8FFEBFFEC91F6FF2AFDA5A123" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="325">
<emphasis id="B933EAFAFFEBFFEC91F6FF2AFE8DA08E" bold="true" box="[199,373,151,176]" pageId="24" pageNumber="325">
<typeStatus id="54FC884AFFEBFFEC91F6FF2AFEFAA08E" box="[199,258,151,176]" pageId="24" pageNumber="325">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54FC884AFFEBFFEC904DFF2AFE1DA08E" box="[380,485,151,176]" pageId="24" pageNumber="325" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED56AE2DFFEBFFEC9303FF25FD72A08E" LSID="urn:lsid:biocol.org:col:34871" box="[562,650,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="325">USNM</collectionCode>
): dissected by us; median lobe of aedeagus and parameres included in
<collectionCode id="ED56AE2DFFEBFFEC91A6FF00FF10A0EA" box="[151,232,189,212]" pageId="24" pageNumber="325">DMHF</collectionCode>
on plastic label pinned under the
<typeStatus id="54FC884AFFEBFFEC9357FF01FD30A0EB" box="[614,712,188,213]" pageId="24" pageNumber="325" type="holotype">holotype</typeStatus>
“
<collectingCountry id="F3507678FFEBFFEC93EBFF01FCDBA0EB" box="[730,803,188,213]" name="Brazil" pageId="24" pageNumber="325">Brazil</collectingCountry>
, MG [= Mato Grosso], Jacare, PNX [= Parque Nacional Xingu],
<date id="FFF91028FFEBFFEC91CBFF62FEA2A0C6" box="[250,346,223,248]" pageId="24" pageNumber="325" value="1965-11">XI.1965</date>
, Alvarenga” [rectangular, white, handwritten], “
<typeStatus id="54FC884AFFEBFFEC92A2FF5DFBD2A0C6" box="[915,1066,224,248]" pageId="24" pageNumber="325" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C474D6BFFEBFFEC9506FF5DFAE6A0C9" box="[1079,1310,224,247]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="youngi">
<emphasis id="B933EAFAFFEBFFEC9506FF5DFAE6A0C9" box="[1079,1310,224,247]" italics="true" pageId="24" pageNumber="325">Laccomimus youngi</emphasis>
</taxonomicName>
Toledo &amp; Michat” [red, rectangular, printed label].
</paragraph>
<paragraph id="8BF836E8FFEBFFEC91F6FE95FABAA15B" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="325">
<typeStatus id="54FC884AFFEBFFEC91F6FE95FECCA17E" box="[199,308,296,320]" pageId="24" pageNumber="325" type="paratype">Paratypes</typeStatus>
(7 exs, all bearing red, printed label “
<typeStatus id="54FC884AFFEBFFEC93D0FE95FC97A17E" box="[737,879,296,320]" pageId="24" pageNumber="325" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C474D6BFFEBFFEC9246FE95FBA1A101" box="[887,1113,296,319]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="youngi">
<emphasis id="B933EAFAFFEBFFEC9246FE95FBA1A101" box="[887,1113,296,319]" italics="true" pageId="24" pageNumber="325">Laccomimus youngi</emphasis>
</taxonomicName>
Toledo &amp; Michat”):
<emphasis id="B933EAFAFFEBFFEC9478FE9AFA63A17E" bold="true" box="[1353,1435,295,320]" pageId="24" pageNumber="325">
<collectingCountry id="F3507678FFEBFFEC9478FE9AFA6EA17E" box="[1353,1430,295,320]" name="Brazil" pageId="24" pageNumber="325">Brazil</collectingCountry>
:
</emphasis>
same data as
<typeStatus id="54FC884AFFEBFFEC901AFEF1FE75A15B" box="[299,397,332,357]" pageId="24" pageNumber="325" type="holotype">holotype</typeStatus>
(7 exs
<collectionCode id="ED56AE2DFFEBFFEC90EEFEF1FDCCA15A" LSID="urn:lsid:biocol.org:col:34871" box="[479,564,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="325">USNM</collectionCode>
: most of them unlabelled: printed label added by the present authors).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFEBFFEC91F6FED2FEF5A1EE" pageId="24" pageNumber="325" type="diagnosis">
<paragraph id="8BF836E8FFEBFFEC91F6FED2FEF5A1EE" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="325">
<emphasis id="B933EAFAFFEBFFEC91F6FED2FEB9A1B6" bold="true" box="[199,321,367,392]" pageId="24" pageNumber="325">Diagnosis.</emphasis>
Very close to
<taxonomicName id="4C474D6BFFEBFFEC90C8FECCFD8DA1B6" box="[505,629,368,392]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFEBFFEC90C8FECCFD8DA1B6" box="[505,629,368,392]" italics="true" pageId="24" pageNumber="325">L. bolivari</emphasis>
</taxonomicName>
but a bit shorter and with less clearly defined markings on elytra. Aedeagus similar to that of
<taxonomicName id="4C474D6BFFEBFFEC90E9FE28FDA9A192" box="[472,593,404,428]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFEBFFEC90E9FE28FDA9A192" box="[472,593,404,428]" italics="true" pageId="24" pageNumber="325">L. bolivari</emphasis>
</taxonomicName>
but longer, with apex ending in distinctive expansion, resembling an axe (
<figureCitation id="137C2A6DFFEBFFEC91AEFE0AFF0BA1EE" box="[159,243,439,464]" captionStart="FIGURES 88 – 90" captionStartId="41.[151,264,1919,1941]" captionTargetBox="[225,1366,198,1894]" captionTargetId="figure@41.[204,1370,193,1898]" captionTargetPageId="41" captionText="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." httpUri="https://zenodo.org/record/244314/files/figure.png" pageId="24" pageNumber="325">Fig. 90</figureCitation>
a).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFEBFFEC91F6FE61FE7AA40B" pageId="24" pageNumber="325" type="description">
<paragraph id="8BF836E8FFEBFFEC91F6FE61FA85A226" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="325">
<emphasis id="B933EAFAFFEBFFEC91F6FE61FEA1A1CB" bold="true" box="[199,345,476,501]" pageId="24" pageNumber="325">Description.</emphasis>
TL: 2.0–
<quantity id="4CBF9B0DFFEBFFEC90F2FE61FDE4A1CA" box="[451,540,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="24" pageNumber="325" unit="mm" value="2.3">2.3 mm</quantity>
; MW: 1.0–
<quantity id="4CBF9B0DFFEBFFEC93AAFE61FD09A1CA" box="[667,753,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="325" unit="mm" value="1.2">1.2 mm</quantity>
(
<typeStatus id="54FC884AFFEBFFEC9231FE61FC90A1CB" box="[768,872,476,501]" pageId="24" pageNumber="325" type="holotype">holotype</typeStatus>
: TL:
<quantity id="4CBF9B0DFFEBFFEC9294FE61FC06A1CA" box="[933,1022,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="24" pageNumber="325" unit="mm" value="2.3">2.3 mm</quantity>
; MW:
<quantity id="4CBF9B0DFFEBFFEC957DFE61FB4DA1CA" box="[1100,1205,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="24" pageNumber="325" unit="mm" value="1.15">1.15 mm</quantity>
); A-TL: 2.0 mm; A- MW:
<quantity id="4CBF9B0DFFEBFFEC91E9FDBDFED6A229" box="[216,302,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="24" pageNumber="325" unit="mm" value="1.1">1.1 mm</quantity>
(NUM: 7 exs); body drop-shaped, short, regularly tapering to apex, pale markings poorly defined.
</paragraph>
<paragraph id="8BF836E8FFEBFFEC91F6FD99FBCBA296" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="325">
<emphasis id="B933EAFAFFEBFFEC91F6FD99FEEFA202" box="[199,279,548,572]" italics="true" pageId="24" pageNumber="325">Colour</emphasis>
(based on old specimens: colouration might be altered). Head and elytra reddish-brown; pronotum paler, with more or less defined dark elongate marking along hind margin. Elytra with poorly defined pale subbasal band–normally not fragmented–and subapical lateral area. Whole underside testaceous or pale-brown. Antennae and mouthparts testaceous. Legs testaceous except for slightly darkened metatarsi.
</paragraph>
<paragraph id="8BF836E8FFEBFFEC91F6FD09FCCCA306" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="325">
<emphasis id="B933EAFAFFEBFFEC91F6FD09FE1FA2F2" box="[199,487,692,716]" italics="true" pageId="24" pageNumber="325">Structures and sculpture</emphasis>
. Head and pronotum as in preceding species: iridescence lacking or faint on pronotum. Elytra with hardly visible subapical sinuation, gently narrowed at tip; smooth with quite impressed reticulation and shallow grooves on basal half; iridescence faint. Iridescence on underside missing or almost so. Prosternal and metacoxal processes as in preceding species.
</paragraph>
<paragraph id="8BF836E8FFEBFFEC91F6FCF9FB68A3D2" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="325">
<emphasis id="B933EAFAFFEBFFEC91F6FCF9FEF9A362" box="[199,257,836,860]" italics="true" pageId="24" pageNumber="325">Male</emphasis>
. Last abdominal ventrite tectiform, with antero-lateral sides visibly sinuate. Outer claw of forelegs scimitar-shaped. Median lobe of aedeagus, in left lateral view (
<figureCitation id="137C2A6DFFEBFFEC926FFCDAFC4AA3BE" box="[862,946,871,896]" captionStart="FIGURES 88 – 90" captionStartId="41.[151,264,1919,1941]" captionTargetBox="[225,1366,198,1894]" captionTargetId="figure@41.[204,1370,193,1898]" captionTargetPageId="41" captionText="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." httpUri="https://zenodo.org/record/244314/files/figure.png" pageId="24" pageNumber="325">Fig. 90</figureCitation>
a), straight as in
<taxonomicName id="4C474D6BFFEBFFEC955AFCD4FB1AA3BE" box="[1131,1250,872,896]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFEBFFEC955AFCD4FB1AA3BE" box="[1131,1250,872,896]" italics="true" pageId="24" pageNumber="325">L. bolivari</emphasis>
</taxonomicName>
but longer, with more elongate, narrow, almost stem-like apex, ending in distinctive expansion, widely rounded dorsally, short and acute ventrally, resembling an axe. In dorsal view as in
<taxonomicName id="4C474D6BFFEBFFEC9235FC0CFC82A3F6" box="[772,890,944,968]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFEBFFEC9235FC0CFC82A3F6" box="[772,890,944,968]" italics="true" pageId="24" pageNumber="325">L. bolivari</emphasis>
</taxonomicName>
but more slender. Left paramere straight as in
<taxonomicName id="4C474D6BFFEBFFEC94B4FC0CFF08A3D2" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFEBFFEC94B4FC0CFF08A3D2" italics="true" pageId="24" pageNumber="325">L. bolivari</emphasis>
</taxonomicName>
but blunt, less acute apically, very short in other dissected specimens (
<figureCitation id="137C2A6DFFEBFFEC9539FC69FBA2A3D3" box="[1032,1114,980,1005]" captionStart="FIGURES 88 – 90" captionStartId="41.[151,264,1919,1941]" captionTargetBox="[225,1366,198,1894]" captionTargetId="figure@41.[204,1370,193,1898]" captionTargetPageId="41" captionText="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." httpUri="https://zenodo.org/record/244314/files/figure.png" pageId="24" pageNumber="325">Fig. 90</figureCitation>
d, e).
</paragraph>
<paragraph id="8BF836E8FFEBFFEC91F6FC45FE7AA40B" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="325">
<emphasis id="B933EAFAFFEBFFEC91F6FC45FEE3A42E" box="[199,283,1016,1040]" italics="true" pageId="24" pageNumber="325">Female</emphasis>
. Last abdominal ventrite triangular, very narrowly rounded at apex, almost pointed. Genital pieces as in preceding species.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFEBFFEC91F6FB82FBA2A466" box="[199,1114,1087,1112]" pageId="24" pageNumber="325" type="distribution">
<paragraph id="8BF836E8FFEBFFEC91F6FB82FBA2A466" blockId="24.[151,1437,151,1221]" box="[199,1114,1087,1112]" pageId="24" pageNumber="325">
<emphasis id="B933EAFAFFEBFFEC91F6FB82FE98A466" bold="true" box="[199,352,1087,1112]" pageId="24" pageNumber="325">Distribution.</emphasis>
<collectingCountry id="F3507678FFEBFFEC9056FB82FE56A466" box="[359,430,1087,1112]" name="Brazil" pageId="24" pageNumber="325">Brazil</collectingCountry>
, Mato Grosso (
<figureCitation id="137C2A6DFFEBFFEC936BFB82FD44A466" box="[602,700,1087,1112]" captionStart="FIGURES 126 – 129" captionStartId="49.[151,264,1953,1975]" captionTargetBox="[157,1432,197,1931]" captionTargetId="figure@49.[151,1436,193,1932]" captionTargetPageId="49" captionText="FIGURES 126 – 129. Distribution maps of Laccomimus species: 126) L. bordoni in South America; 127) L. bolivari (full circles), L. youngi (single open square in Brazil); 128) L. pumilio (open circles), L. alvarengi in Central America (single triangle in Panamá), L. spangleri (stars), L. bordoni in Central America (asterisks); 129) L. improvidus." httpUri="https://zenodo.org/record/244322/files/figure.png" pageId="24" pageNumber="325">Fig. 127</figureCitation>
). Known only from the
<typeStatus id="54FC884AFFEBFFEC92F8FBFDFC01A466" box="[969,1017,1088,1112]" pageId="24" pageNumber="325">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFEBFFEC91F6FBD9FA97A442" box="[199,1391,1124,1149]" pageId="24" pageNumber="325" type="etymology">
<paragraph id="8BF836E8FFEBFFEC91F6FBD9FA97A442" blockId="24.[151,1437,151,1221]" box="[199,1391,1124,1149]" pageId="24" pageNumber="325">
<emphasis id="B933EAFAFFEBFFEC91F6FBD9FEB6A443" bold="true" box="[199,334,1124,1149]" pageId="24" pageNumber="325">Etymology.</emphasis>
Species dedicated to Prof. Frank N. Young. The specific epithet is a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFEBFFEC91F6FB3AFEE2A4FB" pageId="24" pageNumber="325" type="discussion">
<paragraph id="8BF836E8FFEBFFEC91F6FB3AFEE2A4FB" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="325">
<emphasis id="B933EAFAFFEBFFEC91F6FB3AFEC3A49E" bold="true" box="[199,315,1159,1184]" pageId="24" pageNumber="325">Remarks.</emphasis>
<taxonomicName id="4C474D6BFFEBFFEC9075FB34FE57A4A1" box="[324,431,1160,1184]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="youngi">
<emphasis id="B933EAFAFFEBFFEC9075FB34FE57A4A1" box="[324,431,1160,1184]" italics="true" pageId="24" pageNumber="325">L. youngi</emphasis>
</taxonomicName>
is very close to
<taxonomicName id="4C474D6BFFEBFFEC9341FB34FD1FA49E" box="[624,743,1160,1184]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFEBFFEC9341FB34FD1FA49E" box="[624,743,1160,1184]" italics="true" pageId="24" pageNumber="325">L. bolivari</emphasis>
</taxonomicName>
, from which it can be distinguished only by minor aedeagal differences.
</paragraph>
</subSubSection>
</treatment>
</document>
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