577 lines
68 KiB
XML
577 lines
68 KiB
XML
<document id="B89FCBEC53E313E40DBEED26ADDE8C0D" ID-DOI="10.11646/zootaxa.3990.3.1" ID-GBIF-Dataset="cf747765-b9bf-42ed-8479-35355e712db0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244301" ID-ZooBank="378C0359-E0E4-4CCC-821D-649144E37A63" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461268581225" checkinUser="plazi" docAuthor="Toledo, Mario & Michat, Mariano C." docDate="2015" docId="03EE87FEFFE3FFE691A6F8B2FDCAA469" docLanguage="en" docName="zt03990p354.pdf" docOrigin="Zootaxa 3990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccomimus alvarengi Toledo & Michat, 2015, sp. n." docType="treatment" docVersion="9" lastPageNumber="319" masterDocId="FFD7FF86FFF3FFF49131FFBDFFF8A03E" masterDocTitle="Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)" masterLastPageNumber="354" masterPageNumber="301" pageNumber="317" updateTime="1698611165842" updateUser="plazi">
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<mods:mods id="46929D88BFB5F1E46FC853448F16E800" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="82313350D56856A491CB69CE8487C7F8">
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<mods:title id="91A5558385536621916B77773E9BD72B">Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="66E692046B72541F3270525BCA92BC8C" type="personal">
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<mods:role id="94133FA94A303284A756D967263724D3">
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<mods:roleTerm id="98A5CD7A6F06020A7836E4B626430FAB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9E55BB7B6AE5C0E0C3A1029C7EFE2087">Toledo, Mario</mods:namePart>
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</mods:name>
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<mods:name id="CFFCECDE0DAB6ED63C308D6F58017401" type="personal">
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<mods:role id="CFB65DCE810D830714D66444552802C0">
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<mods:roleTerm id="5513A922B4CE1B3C8539D7286811EB52">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A72434A90B124174AEA725940ECD146E">Michat, Mariano C.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="799F4CE95939004171634420DD8E798D">text</mods:typeOfResource>
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<mods:titleInfo id="A8BFA553C79E6AC4A0A3E2E0FBC701A1">
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<mods:title id="0C43E8BC3239365B64FC8390EA65E788">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="71D304B36B311592D5A6AF1442E8451A">
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<mods:date id="EBD2DDFC4C7B87301D0CB8CB81F910FF">2015</mods:date>
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<mods:detail id="33449826375A2AF9C1E40F7DB0F63F7D" type="volume">
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<mods:number id="23FE43402380066BFA2C9EFB598A4EFC">3990</mods:number>
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</mods:detail>
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<mods:detail id="4A44910CA17F16E531DD02148025566D" type="issue">
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<mods:number id="7E2D46D77BE8D02DDE4A2A0F4E59F192">3</mods:number>
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<mods:start id="77E2BA86530F97822894EC85D3426042">301</mods:start>
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<mods:end id="10B79A24A26D9237E9351D5562DD7D39">354</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="7CBEE722A1E241230E1B3546B73BB369">journal article</mods:classification>
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<mods:identifier id="0B7302379AF6FF9B05FB5CEA2737F2DA" type="DOI">10.11646/zootaxa.3990.3.1</mods:identifier>
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<mods:identifier id="C4742DB1A91467501B3907B6E8449231" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="131AA3E5192FB839716452D314B33C4F" type="Zenodo-Dep">244301</mods:identifier>
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<mods:identifier id="AA448FAAEF6884D839A49A4884765244" type="ZooBank">378C0359-E0E4-4CCC-821D-649144E37A63</mods:identifier>
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</mods:mods>
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<treatment id="03EE87FEFFE3FFE691A6F8B2FDCAA469" ID-DOI="http://doi.org/10.5281/zenodo.5692052" ID-GBIF-Taxon="119638445" ID-Zenodo-Dep="5692052" LSID="urn:lsid:plazi:treatment:03EE87FEFFE3FFE691A6F8B2FDCAA469" httpUri="http://treatment.plazi.org/id/03EE87FEFFE3FFE691A6F8B2FDCAA469" lastPageId="18" lastPageNumber="319" pageId="16" pageNumber="317">
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<subSubSection id="C35D6563FFE3FFE491A6F8B2FC0FA775" pageId="16" pageNumber="317" type="nomenclature">
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<paragraph id="8BF836E8FFE3FFE491A6F8B2FE0CA714" blockId="16.[151,1015,1807,1867]" box="[151,500,1807,1834]" pageId="16" pageNumber="317">
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||
<heading id="D0B08184FFE3FFE491A6F8B2FE0CA714" bold="true" box="[151,500,1807,1834]" fontSize="11" level="1" pageId="16" pageNumber="317" reason="1">
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||
<emphasis id="B933EAFAFFE3FFE491A6F8B2FE0CA714" bold="true" box="[151,500,1807,1834]" pageId="16" pageNumber="317">
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||
<taxonomicName id="4C474D6BFFE3FFE491A6F8B2FE53A717" ID-CoL="8LXJS" box="[151,427,1807,1833]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="alvarengi" status="sp. nov.">
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<emphasis id="B933EAFAFFE3FFE491A6F8B2FE53A717" bold="true" box="[151,427,1807,1833]" italics="true" pageId="16" pageNumber="317">Laccomimus alvarengi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2005781FFE3FFE49083F8ADFE0CA714" box="[434,500,1808,1834]" pageId="16" pageNumber="317" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF836E8FFE3FFE491A6F88FFC0FA775" blockId="16.[151,1015,1807,1867]" box="[151,1015,1842,1867]" pageId="16" pageNumber="317">
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(
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<figureCitation id="137C2A6DFFE3FFE491AEF88FFF1FA775" box="[159,231,1842,1867]" captionStart="FIGURES 1 – 7. 1" captionStartId="29.[151,264,1905,1927]" captionTargetBox="[169,1412,219,1874]" captionTargetId="figure@29.[169,1413,193,1876]" captionTargetPageId="29" captionText="FIGURES 1 – 7. 1) Habitus of Laccomimus bolivari, dorsal view (Photo A. E. Z. Short); 2) habitus of Laccodytes apalodes, dorsal view (Photo A. E. Z. Short); 3 – 7) SEMs of Laccomimus species (females): 3) habitus of L. alvarengi, dorsal view; 4) habitus of L. bordoni, dorsal view; 5) pronotum of L. alvarengi, dorsal view; 6) pronotum of L. bordoni, dorsal view; 7) pronotum of L. bordoni, dorsolateral view." httpUri="https://zenodo.org/record/244302/files/figure.png" pageId="16" pageNumber="317">Figs 3</figureCitation>
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; 5; 11; 14–15; 28; 39; 57; 64; 69; 77–81; 94–102; 111; 118; 125; 128)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6563FFE3FFE591A6F8C7FAE2A546" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317" type="materials_examined">
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<paragraph id="8BF836E8FFE3FFE491A6F8C7FC89A7AD" blockId="16.[151,1436,1914,2011]" box="[151,881,1914,1939]" pageId="16" pageNumber="317">
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<emphasis id="B933EAFAFFE3FFE491A6F8C7FECDA7AD" bold="true" box="[151,309,1914,1939]" pageId="16" pageNumber="317">
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<typeStatus id="54FC884AFFE3FFE491A6F8C7FF2AA7AD" box="[151,210,1914,1939]" pageId="16" pageNumber="317">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F3507678FFE3FFE4900DF8C7FE7BA7AD" box="[316,387,1914,1939]" name="Brazil" pageId="16" pageNumber="317">Brazil</collectingCountry>
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, Mato Grosso, Jacaré, Xingu National Park.
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</paragraph>
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<paragraph id="8BF836E8FFE3FFE591F6F823FECCA0C6" blockId="16.[151,1436,1914,2011]" lastBlockId="17.[151,1437,151,2013]" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317">
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<emphasis id="B933EAFAFFE3FFE491F6F823FE80A789" bold="true" box="[199,376,1950,1975]" pageId="16" pageNumber="317">
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<typeStatus id="54FC884AFFE3FFE491F6F823FEFAA789" box="[199,258,1950,1975]" pageId="16" pageNumber="317">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54FC884AFFE3FFE490B3F823FE13A789" box="[386,491,1950,1975]" pageId="16" pageNumber="317" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="ED56AE2DFFE3FFE4930FF822FD6EA789" LSID="urn:lsid:biocol.org:col:34871" box="[574,662,1951,1975]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="317">USNM</collectionCode>
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): dissected for genitalia probably by Young; genitalia and last two abdominal ventrites glued on separate label under the
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<typeStatus id="54FC884AFFE3FFE4923BF87FFC94A7E5" box="[778,876,1986,2011]" pageId="16" pageNumber="317" type="holotype">holotype</typeStatus>
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“
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<collectingCountry id="F3507678FFE3FFE492B0F87FFC32A7E5" box="[897,970,1986,2011]" name="Brazil" pageId="16" pageNumber="317">Brazil</collectingCountry>
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, Matto [sic] Grosso, Jacaré, P.N. Xingu,
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<date id="FFF91028FFE2FFE591A6FF2AFF0EA08E" box="[151,246,151,176]" pageId="17" pageNumber="318" value="1961-11">XI.1961</date>
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, Alvarenga & Werner leg” [printed], “FIC” [small, pink, printed label]”, “
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<taxonomicName id="4C474D6BFFE2FFE5950FFF25FA6AA08E" authority="Young" authorityName="Young" box="[1086,1426,152,176]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="alvarengi">
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<emphasis id="B933EAFAFFE2FFE5950FFF25FAC6A08E" box="[1086,1342,152,176]" italics="true" pageId="17" pageNumber="318">Laccomimus alvarengi</emphasis>
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Young
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</taxonomicName>
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” [white label, handwritten by Young], “
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<typeStatus id="54FC884AFFE2FFE59363FF01FD11A0EA" box="[594,745,188,212]" pageId="17" pageNumber="318" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4C474D6BFFE2FFE593C3FF00FC0CA0EA" box="[754,1012,188,212]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="alvarengi">
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<emphasis id="B933EAFAFFE2FFE593C3FF00FC0CA0EA" box="[754,1012,188,212]" italics="true" pageId="17" pageNumber="318">Laccomimus alvarengi</emphasis>
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</taxonomicName>
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Toledo & Michat” [red, rectangular, printed label].
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</paragraph>
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<paragraph id="8BF836E8FFE2FFE591F6FEB8FAE2A546" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
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<typeStatus id="54FC884AFFE2FFE591F6FEB8FECCA123" box="[199,308,261,285]" pageId="17" pageNumber="318" type="paratype">Paratypes</typeStatus>
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(200 exs all bearing red, printed label “
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<typeStatus id="54FC884AFFE2FFE5923EFEB9FC65A122" box="[783,925,260,284]" pageId="17" pageNumber="318" type="paratype">PARATYPE</typeStatus>
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<taxonomicName id="4C474D6BFFE2FFE59299FEB8FB52A122" box="[936,1194,260,284]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="alvarengi">
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<emphasis id="B933EAFAFFE2FFE59299FEB8FB52A122" box="[936,1194,260,284]" italics="true" pageId="17" pageNumber="318">Laccomimus alvarengi</emphasis>
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</taxonomicName>
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Toledo & Michat”):
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<emphasis id="B933EAFAFFE2FFE591A6FE9AFF11A17E" bold="true" box="[151,233,295,320]" pageId="17" pageNumber="318">
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<collectingCountry id="F3507678FFE2FFE591A6FE9AFF1DA17E" box="[151,229,295,320]" name="Brazil" pageId="17" pageNumber="318">Brazil</collectingCountry>
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:
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</emphasis>
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same data as
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<typeStatus id="54FC884AFFE2FFE590B9FE9AFE12A17E" box="[392,490,295,320]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
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(10 exs
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<collectionCode id="ED56AE2DFFE2FFE5937DFE95FD5AA17E" LSID="urn:lsid:biocol.org:col:34871" box="[588,674,296,320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
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); same data as
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<typeStatus id="54FC884AFFE2FFE5927DFE9AFC56A17E" box="[844,942,295,320]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
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except for “
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<date id="FFF91028FFE2FFE5950AFE9AFB62A17E" box="[1083,1178,295,320]" pageId="17" pageNumber="318" value="1965-11">XI.1965</date>
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” (65 exs
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<collectionCode id="ED56AE2DFFE2FFE59436FE95FAA0A17E" LSID="urn:lsid:biocol.org:col:34871" box="[1287,1368,296,320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
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–most of them without label); “Mato Grosso, Cuiabá,
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<date id="FFF91028FFE2FFE59397FEF1FCDEA15A" box="[678,806,332,356]" pageId="17" pageNumber="318" value="1972-03-30">30.III.1972</date>
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, W.H. Withcomb” (8 exs
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<collectionCode id="ED56AE2DFFE2FFE59575FEF1FB62A15A" LSID="urn:lsid:biocol.org:col:34871" box="[1092,1178,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
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); idem but “
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<date id="FFF91028FFE2FFE59411FEF1FA69A15A" box="[1312,1425,332,356]" pageId="17" pageNumber="318" value="1972-04-09">9.IV.1972</date>
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” (6 exs
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<collectionCode id="ED56AE2DFFE2FFE591C3FECDFEB0A1B6" LSID="urn:lsid:biocol.org:col:34871" box="[242,328,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
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); idem but “
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<date id="FFF91028FFE2FFE590D9FECDFD9EA1B6" box="[488,614,368,392]" pageId="17" pageNumber="318" value="1972-04-11">11.IV.1972</date>
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” (3 exs
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<collectionCode id="ED56AE2DFFE2FFE593EBFECDFCC8A1B6" LSID="urn:lsid:biocol.org:col:34871" box="[730,816,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
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); idem but “
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<date id="FFF91028FFE2FFE592E1FECDFBB7A1B6" box="[976,1103,368,392]" pageId="17" pageNumber="318" value="1972-04-21">21.IV.1972</date>
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” (1 ex.
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<collectionCode id="ED56AE2DFFE2FFE595F1FECDFAEEA1B6" LSID="urn:lsid:biocol.org:col:34871" box="[1216,1302,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
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||
); idem but “
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<date id="FFF91028FFE2FFE59190FE29FED9A192" box="[161,289,404,428]" pageId="17" pageNumber="318" value="1972-04-22">22.IV.1972</date>
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” (7 exs
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<collectionCode id="ED56AE2DFFE2FFE590B9FE29FE20A193" LSID="urn:lsid:biocol.org:col:34871" box="[392,472,404,429]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
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–four of which without label); “Mato Grosso, Rio Bento Gomes (Pantanal),
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<date id="FFF91028FFE2FFE59467FE29FF2AA1EE" pageId="17" pageNumber="318" value="1993" valueMax="1994">1993– 1994</date>
|
||
, E. Stuhr leg.”, “Flussabschnitt bei [= section of river at] Periquito,
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<geoCoordinate id="EE73502FFFE2FFE592CBFE0AFBA7A1EE" box="[1018,1119,439,464]" direction="south" orientation="latitude" pageId="17" pageNumber="318" precision="925" value="-15.9">15°54' S</geoCoordinate>
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,
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<geoCoordinate id="EE73502FFFE2FFE5955EFE0AFB27A1EE" box="[1135,1247,439,464]" direction="west" orientation="longitude" pageId="17" pageNumber="318" precision="925" value="-56.55">56°33' W</geoCoordinate>
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” (1 ex.
|
||
<collectionCode id="ED56AE2DFFE2FFE59472FE05FA68A1EE" box="[1347,1424,440,464]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="318">NMW</collectionCode>
|
||
); “Rondôna,
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||
<quantity id="4CBF9B0DFFE2FFE5902BFE61FE9CA1CB" box="[282,356,476,501]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="17" pageNumber="318" unit="km" value="62.0">62 km</quantity>
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SW Ariquemes, nr. fz. Rancho Grande,
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||
<date id="FFF91028FFE2FFE59270FE61FC27A1CA" box="[833,991,476,500]" pageId="17" pageNumber="318" value="1994-09-08" valueMax="1994-09-20" valueMin="1994-09-08">8–20.IX.1994</date>
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||
, JE Eger & C.W. O’Brien leg” (3 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE591A6FDBDFF2DA226" box="[151,213,512,536]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="17" pageNumber="318">ZSM</collectionCode>
|
||
); “Minas Gerais, Prov. Cordisburgo, Faz. Pontinha,
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||
<date id="FFF91028FFE2FFE59211FDBDFC70A226" box="[800,904,512,536]" pageId="17" pageNumber="318" value="1997-12">XII.1997</date>
|
||
, leg. F.Z. Vaz-de-Mello, coll. L. Hendrich” (16 exs
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||
<collectionCode id="ED56AE2DFFE2FFE591FBFD99FEF0A202" box="[202,264,548,572]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="17" pageNumber="318">ZSM</collectionCode>
|
||
); “Par. Curitiba,
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<date id="FFF91028FFE2FFE590E9FD99FDA1A203" box="[472,601,548,573]" pageId="17" pageNumber="318" value="1969-06-28">28.VI.1969</date>
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, leg. P.& P. Spangler” (1 ex.
|
||
<collectionCode id="ED56AE2DFFE2FFE592FCFD99FBDBA202" LSID="urn:lsid:biocol.org:col:34871" box="[973,1059,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
|
||
).
|
||
<emphasis id="B933EAFAFFE2FFE5950DFD99FB46A203" bold="true" box="[1084,1214,548,573]" pageId="17" pageNumber="318">
|
||
<collectingCountry id="F3507678FFE2FFE5950DFD99FB42A203" box="[1084,1210,548,573]" name="Argentina" pageId="17" pageNumber="318">Argentina</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
“Prov. Corrientes, Mbrucuyá N.P.,
|
||
<date id="FFF91028FFE2FFE5907EFDFAFE1FA25E" box="[335,487,583,608]" pageId="17" pageNumber="318" value="2004-01-15" valueMax="2004-01-17" valueMin="2004-01-15">15–17.I.2004</date>
|
||
, light trap, leg. M.C. Michat” (24 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE592A9FDF5FC0DA25E" box="[920,1013,584,608]" httpUri="http://grbio.org/cool/654h-avhc" name="Museo de Historia Natural de la Ciudad de Mexico" pageId="17" pageNumber="318">MCMC</collectionCode>
|
||
).
|
||
<emphasis id="B933EAFAFFE2FFE59539FDFAFB9DA25E" bold="true" box="[1032,1125,583,608]" pageId="17" pageNumber="318">
|
||
<collectingCountry id="F3507678FFE2FFE59539FDFAFB98A25E" box="[1032,1120,583,608]" name="Bolivia" pageId="17" pageNumber="318">Bolivia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
“Beni,
|
||
<quantity id="4CBF9B0DFFE2FFE5958CFDF5FAFDA25E" box="[1213,1285,583,608]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="318" unit="km" value="40.0">40 km</quantity>
|
||
E San Borja, Estacion Biologica Beni, Estancia El Porvenir,
|
||
<date id="FFF91028FFE2FFE59385FDD1FCBBA2BB" box="[692,835,620,645]" pageId="17" pageNumber="318" value="1987-09-06" valueMax="1987-09-08" valueMin="1987-09-06">6–8.IX.1987</date>
|
||
, M. E. Steiner”, “At black light, open grass savanna and marsh” (7 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE59057FD2DFE44A296" LSID="urn:lsid:biocol.org:col:34871" box="[358,444,656,680]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
|
||
); “Guayaramerin, Beni, river bank, Soil Zoological Exp., Balogh, Mahunka, Zicsi”, “No. 428, beaten from vegetation,
|
||
<date id="FFF91028FFE2FFE590D3FD09FDA7A2F3" box="[482,607,692,717]" pageId="17" pageNumber="318" value="1966-12-02">2.XII.1966</date>
|
||
” (2 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE5938CFD09FCEEA2F2" box="[701,790,692,716]" pageId="17" pageNumber="318">HNMB</collectionCode>
|
||
); “El Beni, Beni Station, NE of San Borja,
|
||
<date id="FFF91028FFE2FFE59431FD09FA75A2F2" box="[1280,1421,692,716]" pageId="17" pageNumber="318" value="1988-07-31">
|
||
31.
|
||
<collectionCode id="ED56AE2DFFE2FFE59411FD09FAB4A2F2" box="[1312,1356,692,716]" pageId="17" pageNumber="318">VII</collectionCode>
|
||
.1988
|
||
</date>
|
||
”, “Robert W. Brooks ex., at lights, BIOLAT-SI/MAB” (2 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE59202FD65FC8EA2CE" box="[819,886,728,752]" pageId="17" pageNumber="318">VAIP</collectionCode>
|
||
); “El Beni, El Porvenir Station, NE of San Borja,
|
||
<date id="FFF91028FFE2FFE591A6FD41FEC3A32B" box="[151,315,764,789]" pageId="17" pageNumber="318" value="1988-08-06" valueMax="1988-08-09" valueMin="1988-08-06">6–9.VIII.1988</date>
|
||
”, “Robert W. Brooks ex., at lights, BIOLAT-SI/MAB” (1 ex.
|
||
<collectionCode id="ED56AE2DFFE2FFE592CBFD41FBC5A32A" box="[1018,1085,764,788]" pageId="17" pageNumber="318">VAIP</collectionCode>
|
||
); “Santa Crus, Sara,
|
||
<quantity id="4CBF9B0DFFE2FFE5941BFD41FA9DA32B" box="[1322,1381,764,789]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="318" unit="km" value="2.0">2 km</quantity>
|
||
NW Santa Rosa,
|
||
<date id="FFF91028FFE2FFE59010FCA2FE60A306" box="[289,408,799,824]" pageId="17" pageNumber="318" value="1969-02-21">21.II.1969</date>
|
||
,
|
||
<collectionCode id="ED56AE2DFFE2FFE59095FC9DFE2EA309" LSID="urn:lsid:biocol.org:col:12647" box="[420,470,800,823]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12647" name="Belfast Natural History and Philosophical Society" pageId="17" pageNumber="318">BLT</collectionCode>
|
||
A. Martinez & R.E. Woodruff” (5 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE592B6FC9DFC25A306" LSID="urn:lsid:biocol.org:col:34871" box="[903,989,800,824]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
|
||
); idem “
|
||
<quantity id="4CBF9B0DFFE2FFE5950AFC9DFB8CA306" box="[1083,1140,799,824]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="318" unit="km" value="3.0">3 km</quantity>
|
||
S Santa Rosa,
|
||
<date id="FFF91028FFE2FFE5942AFCA2FA69A306" box="[1307,1425,799,824]" pageId="17" pageNumber="318" value="1969-02-22">22.II.1969</date>
|
||
” (7 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE591DAFCF9FEB9A362" LSID="urn:lsid:biocol.org:col:34871" box="[235,321,836,860]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
|
||
).
|
||
<emphasis id="B933EAFAFFE2FFE59069FCF9FE30A363" bold="true" box="[344,456,836,861]" pageId="17" pageNumber="318">
|
||
<collectingCountry id="F3507678FFE2FFE59069FCF9FE3BA363" box="[344,451,836,861]" name="Ecuador" pageId="17" pageNumber="318">Ecuador</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
“Esmeraldas,
|
||
<quantity id="4CBF9B0DFFE2FFE59345FCF9FD49A363" box="[628,689,836,861]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="318" unit="km" value="9.0">9 km</quantity>
|
||
S San Lorenzo,
|
||
<date id="FFF91028FFE2FFE5924FFCF9FC0BA363" box="[894,1011,836,861]" pageId="17" pageNumber="318" value="1979-05-25">25.V.1979</date>
|
||
, ultraviolet light, Jos. J. Anderson”, “
|
||
<collectingCountry id="F3507678FFE2FFE59190FCDAFEF9A3BE" box="[161,257,871,896]" name="Ecuador" pageId="17" pageNumber="318">Ecuador</collectingCountry>
|
||
Peace Corps, Smithsonian Institution Aquatic Insects Survey” (3 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE59507FCD5FB7FA3BE" LSID="urn:lsid:biocol.org:col:34871" box="[1078,1159,872,896]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
|
||
–one ex. without label); “Los Rios, Babahoyo,
|
||
<date id="FFF91028FFE2FFE590ACFC31FDE7A39B" box="[413,543,908,933]" pageId="17" pageNumber="318" value="1975-06-21">21.VI.1975</date>
|
||
, Cohen, Langley & Monnig”, “
|
||
<collectingCountry id="F3507678FFE2FFE592BAFC31FC13A39B" box="[907,1003,908,933]" name="Ecuador" pageId="17" pageNumber="318">Ecuador</collectingCountry>
|
||
Peace Corps, Smithsonian Institution Aquatic Insects Survey” (2 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE59336FC0DFDA5A3F6" LSID="urn:lsid:biocol.org:col:34871" box="[519,605,944,968]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
|
||
).
|
||
<emphasis id="B933EAFAFFE2FFE59342FC12FD26A3F6" bold="true" box="[627,734,943,968]" pageId="17" pageNumber="318">
|
||
<collectingCountry id="F3507678FFE2FFE59342FC12FD21A3F6" box="[627,729,943,968]" name="Panama" pageId="17" pageNumber="318">Panamá</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
“C. Z. [= Canal Zone], Albrook Forest site,
|
||
<date id="FFF91028FFE2FFE595DDFC12FA60A3F6" box="[1260,1432,943,968]" pageId="17" pageNumber="318" value="1967-09-21" valueMax="1967-09-22" valueMin="1967-09-21">21–22.IX.1967</date>
|
||
,
|
||
<collectionCode id="ED56AE2DFFE2FFE591A6FC68FF31A3D2" LSID="urn:lsid:biocol.org:col:12647" box="[151,201,981,1004]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12647" name="Belfast Natural History and Philosophical Society" pageId="17" pageNumber="318">BLT</collectionCode>
|
||
<collectionCode id="ED56AE2DFFE2FFE591E3FC69FF0CA3D2" box="[210,244,980,1004]" pageId="17" pageNumber="318">RS</collectionCode>
|
||
Hutton” (1 ex.
|
||
<collectionCode id="ED56AE2DFFE2FFE5909CFC69FDFBA3D2" LSID="urn:lsid:biocol.org:col:34871" box="[429,515,980,1004]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
|
||
).
|
||
<emphasis id="B933EAFAFFE2FFE59326FC69FDA2A3D3" bold="true" box="[535,602,980,1005]" pageId="17" pageNumber="318">
|
||
<collectingCountry id="F3507678FFE2FFE59326FC69FDADA3D3" box="[535,597,980,1005]" name="Peru" pageId="17" pageNumber="318">Perú</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
“Prov. Loreto, Rio Yarapa, Puerto Miguel”, “
|
||
<quantity id="4CBF9B0DFFE2FFE59559FC69FB4DA3D2" box="[1128,1205,980,1004]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="318" unit="m" value="200.0">200 m</quantity>
|
||
,
|
||
<date id="FFF91028FFE2FFE595F2FC69FA80A3D3" box="[1219,1400,980,1005]" pageId="17" pageNumber="318" value="1994-12-16" valueMax="1994-12-23" valueMin="1994-12-16">16–23.XII.1994</date>
|
||
, T. Hàcz & G. Holzinger” (3 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE590D5FC45FDC5A42E" box="[484,573,1016,1040]" pageId="17" pageNumber="318">HNMB</collectionCode>
|
||
); “Loreto, Yacumama Lodge,
|
||
<geoCoordinate id="EE73502FFFE2FFE592A1FC4AFC1FA42E" box="[912,999,1015,1040]" direction="west" orientation="longitude" pageId="17" pageNumber="318" precision="5555" value="-73.6">73.6°W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE73502FFFE2FFE592C2FC45FBCBA42E" box="[1011,1075,1016,1040]" direction="south" orientation="latitude" pageId="17" pageNumber="318" precision="5555" value="-4.8">4.8°S</geoCoordinate>
|
||
, nr. junction Rio Maranon - Rio Ucayali,
|
||
<date id="FFF91028FFE2FFE591CCFBA1FE55A40B" box="[253,429,1052,1077]" pageId="17" pageNumber="318" value="1994-08-06" valueMax="1994-08-20" valueMin="1994-08-06">6–20.VIII.1994</date>
|
||
, Skelly leg., small light in wood, coll. L. Hendrich” (8 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE5957CFBA1FB74A40A" box="[1101,1164,1052,1076]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="17" pageNumber="318">ZSM</collectionCode>
|
||
); “Colonia Calleria, Rio Calleria,
|
||
<quantity id="4CBF9B0DFFE2FFE59031FB82FEB1A466" box="[256,329,1087,1112]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="318" unit="km" value="15.0">15 km</quantity>
|
||
E Ecayali, muddy river shore,
|
||
<date id="FFF91028FFE2FFE59384FB82FCAFA466" box="[693,855,1087,1112]" pageId="17" pageNumber="318" value="1961-10-12" valueMax="1961-10-25" valueMin="1961-10-12">12–25.X.1961</date>
|
||
, B. Malkin” (2 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE59571FBFDFB6EA466" LSID="urn:lsid:biocol.org:col:34871" box="[1088,1174,1088,1112]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
|
||
); “Madre de Dios, Rio Tambopata Res., 30 air km SW Pto. Maldonado,
|
||
<quantity id="4CBF9B0DFFE2FFE593F6FBD9FCE9A442" box="[711,785,1124,1148]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="17" pageNumber="318" unit="m" value="290.0">290 m</quantity>
|
||
,
|
||
<date id="FFF91028FFE2FFE5922FFBD9FC56A443" box="[798,942,1124,1149]" pageId="17" pageNumber="318" value="1979-11-02" valueMax="1979-11-05" valueMin="1979-11-02">2–5.XI.1979</date>
|
||
, J. B. Heppner, subtropical moist forest” (3 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE591FBFB35FED8A49E" LSID="urn:lsid:biocol.org:col:34871" box="[202,288,1160,1184]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
|
||
).
|
||
<emphasis id="B933EAFAFFE2FFE59008FB3AFE4DA49E" bold="true" box="[313,437,1159,1184]" pageId="17" pageNumber="318">
|
||
<collectingCountry id="F3507678FFE2FFE59008FB3AFE48A49E" box="[313,432,1159,1184]" name="Paraguay" pageId="17" pageNumber="318">Paraguay</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
“Jejuí-Mí, light trap,
|
||
<date id="FFF91028FFE2FFE593F4FB35FCA9A49E" box="[709,849,1160,1184]" pageId="17" pageNumber="318" value="2003-12-18">18.XII.2003</date>
|
||
, leg. M.C. Michat” (2 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE59595FB35FAF8A49E" box="[1188,1280,1160,1184]" httpUri="http://grbio.org/cool/654h-avhc" name="Museo de Historia Natural de la Ciudad de Mexico" pageId="17" pageNumber="318">MCMC</collectionCode>
|
||
); “Dep. Alto
|
||
<collectingCountry id="F3507678FFE2FFE591A6FB10FEF8A4FB" box="[151,256,1197,1221]" name="Paraguay" pageId="17" pageNumber="318">Paraguay</collectingCountry>
|
||
, Campo Grande Farm, light trap,
|
||
<date id="FFF91028FFE2FFE59347FB11FD0CA4FA" box="[630,756,1196,1220]" pageId="17" pageNumber="318" value="2002-12-07">7.XII.2002</date>
|
||
, leg. M.C. Michat” (2 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE5952BFB11FB8EA4FA" box="[1050,1142,1196,1220]" httpUri="http://grbio.org/cool/654h-avhc" name="Museo de Historia Natural de la Ciudad de Mexico" pageId="17" pageNumber="318">MCMC</collectionCode>
|
||
); “Dep. Alto
|
||
<collectingCountry id="F3507678FFE2FFE5943BFB10FA8BA4FB" box="[1290,1395,1197,1221]" name="Paraguay" pageId="17" pageNumber="318">Paraguay</collectingCountry>
|
||
, rd. to Fortín Patria,
|
||
<date id="FFF91028FFE2FFE59064FB6DFE2BA4D6" box="[341,467,1232,1256]" pageId="17" pageNumber="318" value="2002-12-03">3.XII.2002</date>
|
||
, light trap, leg. M.C. Michat” (4 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE592B5FB6DFC18A4D6" box="[900,992,1232,1256]" httpUri="http://grbio.org/cool/654h-avhc" name="Museo de Historia Natural de la Ciudad de Mexico" pageId="17" pageNumber="318">MCMC</collectionCode>
|
||
); “Dep. Alto
|
||
<collectingCountry id="F3507678FFE2FFE5954DFB6CFB1DA4D7" box="[1148,1253,1233,1257]" name="Paraguay" pageId="17" pageNumber="318">Paraguay</collectingCountry>
|
||
, Defensores del Chaco N.P., Fortín Madrejón,
|
||
<date id="FFF91028FFE2FFE590D9FB49FD93A532" box="[488,619,1268,1292]" pageId="17" pageNumber="318" value="2002-11-23">23.XI.2002</date>
|
||
, light trap, leg. M.C. Michat” (2 exs
|
||
<collectionCode id="ED56AE2DFFE2FFE59539FB49FB9CA532" box="[1032,1124,1268,1292]" httpUri="http://grbio.org/cool/654h-avhc" name="Museo de Historia Natural de la Ciudad de Mexico" pageId="17" pageNumber="318">MCMC</collectionCode>
|
||
); “Dep. Cordillera, Arroyos y Esteros,
|
||
<date id="FFF91028FFE2FFE59022FAA5FE7FA50E" box="[275,391,1304,1328]" pageId="17" pageNumber="318" value="1991-09-08">8.IX.1991</date>
|
||
, Ulf Drechsel leg., Coll. Hendrich” (1 ex.
|
||
<collectionCode id="ED56AE2DFFE2FFE5924AFAA5FC41A50E" box="[891,953,1304,1328]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="17" pageNumber="318">ZSM</collectionCode>
|
||
); “Dep. Central, Asuncion, Rio
|
||
<collectingCountry id="F3507678FFE2FFE5941EFAA4FA60A50F" box="[1327,1432,1305,1329]" name="Paraguay" pageId="17" pageNumber="318">Paraguay</collectingCountry>
|
||
,
|
||
<date id="FFF91028FFE2FFE591A6FA81FEF3A56A" box="[151,267,1340,1364]" pageId="17" pageNumber="318" value="1990-11-02">2.XI.1990</date>
|
||
, Ulf Drechsel leg., Coll. Hendrich” (1 ex.
|
||
<collectionCode id="ED56AE2DFFE2FFE593DDFA81FCD2A56A" box="[748,810,1340,1364]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="17" pageNumber="318">ZSM</collectionCode>
|
||
).
|
||
<emphasis id="B933EAFAFFE2FFE5920CFA81FC43A56B" bold="true" box="[829,955,1340,1365]" pageId="17" pageNumber="318">
|
||
<collectingCountry id="F3507678FFE2FFE5920CFA81FC4EA56B" box="[829,950,1340,1365]" name="Suriname" pageId="17" pageNumber="318">Suriname</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
“Paramaribo, Guest House of Museum, at light,
|
||
<date id="FFF91028FFE2FFE591E9FAE2FE76A546" box="[216,398,1375,1400]" pageId="17" pageNumber="318" value="1969-07-19" valueMax="1969-07-20" valueMin="1969-07-19">
|
||
19–20.
|
||
<collectionCode id="ED56AE2DFFE2FFE59014FAE2FEB7A546" box="[293,335,1375,1400]" pageId="17" pageNumber="318">VII</collectionCode>
|
||
.1969
|
||
</date>
|
||
, leg N. Nieser” (1 ex.
|
||
<collectionCode id="ED56AE2DFFE2FFE593B9FADDFD2DA546" box="[648,725,1376,1400]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="318">NMW</collectionCode>
|
||
); idem except for “
|
||
<date id="FFF91028FFE2FFE59298FAE2FB90A546" box="[937,1128,1375,1400]" pageId="17" pageNumber="318" value="1969-08-21" valueMax="1969-08-25" valueMin="1969-08-21">21–25.VIII.1969</date>
|
||
” (1 ex.
|
||
<collectionCode id="ED56AE2DFFE2FFE595F0FADDFAF6A546" box="[1217,1294,1376,1400]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="318">NMW</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D6563FFE2FFE591F6FA39FB18A612" pageId="17" pageNumber="318" type="diagnosis">
|
||
<paragraph id="8BF836E8FFE2FFE591F6FA39FB18A612" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
|
||
<emphasis id="B933EAFAFFE2FFE591F6FA39FEB9A5A3" bold="true" box="[199,321,1412,1437]" pageId="17" pageNumber="318">Diagnosis.</emphasis>
|
||
Recognisable from most species of
|
||
<taxonomicName id="4C474D6BFFE2FFE593D6FA38FC8CA5A2" box="[743,884,1413,1436]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B933EAFAFFE2FFE593D6FA38FC8CA5A2" box="[743,884,1413,1436]" italics="true" pageId="17" pageNumber="318">Laccomimus</emphasis>
|
||
</taxonomicName>
|
||
by aedeagal characters only, and from the very close
|
||
<taxonomicName id="4C474D6BFFE2FFE591E9FA14FEB4A5FE" box="[216,332,1448,1472]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE2FFE591E9FA14FEB4A5FE" box="[216,332,1448,1472]" italics="true" pageId="17" pageNumber="318">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
(see above) by aedeagal characters and, frequently, by the more distinct pattern on elytra. Median lobe differentiated into basal and distal portion, divided by an angle or a curve (
|
||
<figureCitation id="137C2A6DFFE2FFE59529FA71FB52A5DA" box="[1048,1194,1484,1509]" captionStart-0="FIGURES 94 – 97" captionStart-1="FIGURES 98 – 102" captionStartId-0="43.[151,264,1919,1941]" captionStartId-1="44.[151,264,1877,1899]" captionTargetBox-0="[174,1408,200,1884]" captionTargetBox-1="[162,1422,201,1834]" captionTargetId-0="figure@43.[164,1418,193,1889]" captionTargetId-1="figure@44.[151,1436,193,1856]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURES 94 – 97. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 94) holotype, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 95) likely aberrant specimen from type locality, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 96) Loreto, Perú, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 97) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere." captionText-1="FIGURES 98 – 102. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 98) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere; 99) Beni, Bolivia, a) median lobe, b) magnification of apex of median lobe; 100) Santa Rosa, Bolivia, median lobe; 101) Esmeraldas, Ecuador, a) median lobe, b) left paramere; 102) Albrook, Panamá, a) median lobe, b) magnification of apex of median lobe, c) left paramere." httpUri-0="https://zenodo.org/record/244316/files/figure.png" httpUri-1="https://zenodo.org/record/244317/files/figure.png" pageId="17" pageNumber="318">Figs 94–102</figureCitation>
|
||
), with apex ending in small button-like expansion. The rounded tip of the prosternal process, which is very different from the acuminate tip of
|
||
<taxonomicName id="4C474D6BFFE2FFE591EBF9A8FEB4A612" box="[218,332,1556,1580]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE2FFE591EBF9A8FEB4A612" box="[218,332,1556,1580]" italics="true" pageId="17" pageNumber="318">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
, is constant in this species. The average size is smaller with respect to
|
||
<taxonomicName id="4C474D6BFFE2FFE59557F9A8FB21A612" box="[1126,1241,1556,1580]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE2FFE59557F9A8FB21A612" box="[1126,1241,1556,1580]" italics="true" pageId="17" pageNumber="318">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D6563FFE2FFE691F6F98AFA93A122" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318" type="description">
|
||
<paragraph id="8BF836E8FFE2FFE591F6F98AFD0CA6A6" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
|
||
<emphasis id="B933EAFAFFE2FFE591F6F98AFEA1A66E" bold="true" box="[199,345,1591,1616]" pageId="17" pageNumber="318">Description.</emphasis>
|
||
<emphasis id="B933EAFAFFE2FFE5906EF985FE40A66E" box="[351,440,1592,1616]" italics="true" pageId="17" pageNumber="318">Habitus</emphasis>
|
||
(
|
||
<figureCitation id="137C2A6DFFE2FFE590F6F98AFDE4A66E" box="[455,540,1591,1616]" captionStart="FIGURES 59 – 66" captionStartId="38.[151,264,1953,1975]" captionTargetBox="[161,1426,197,1927]" captionTargetId="figure@38.[151,1435,193,1932]" captionTargetPageId="38" captionText="FIGURES 59 – 66. Subschematic body shape and colour patterns in Laccomimus species: 59) L. distinctus; 60) L. malkini, a) typical specimen from Colombia, b) specimen from Piracicaba, Brazil; 61) L. variegatus (Perú); 62) L. amazonas; 63) L. pumilio (Florida, USA); 64) L. alvarengi, a – b) variation in colouration patterns; 65) L. bordoni; 66) L. spangleri." httpUri="https://zenodo.org/record/244311/files/figure.png" pageId="17" pageNumber="318">Fig. 64</figureCitation>
|
||
). TL:
|
||
<quantity id="4CBF9B0DFFE2FFE59350F98AFD01A66E" box="[609,761,1591,1616]" metricMagnitude="-3" metricUnit="m" metricValue="2.075" metricValueMax="2.25" metricValueMin="1.9" pageId="17" pageNumber="318" unit="mm" value="2.075" valueMax="2.25" valueMin="1.9">1.9–2.25 mm</quantity>
|
||
; MW:
|
||
<quantity id="4CBF9B0DFFE2FFE59277F98AFC24A66E" box="[838,988,1591,1616]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.2" metricValueMin="0.95" pageId="17" pageNumber="318" unit="mm" value="1.075" valueMax="1.2" valueMin="0.95">0.95–1.2 mm</quantity>
|
||
(
|
||
<typeStatus id="54FC884AFFE2FFE592DAF98AFBABA66E" box="[1003,1107,1591,1616]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
|
||
: TL:
|
||
<quantity id="4CBF9B0DFFE2FFE595BEF985FB10A66E" box="[1167,1256,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="318" unit="mm" value="2.2">2.2 mm</quantity>
|
||
; MW:
|
||
<quantity id="4CBF9B0DFFE2FFE59404F985FA77A66E" box="[1333,1423,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="318" unit="mm" value="1.1">1.1 mm</quantity>
|
||
); A-TL:
|
||
<quantity id="4CBF9B0DFFE2FFE591D4F9E1FEC6A64A" box="[229,318,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="17" pageNumber="318" unit="mm" value="2.1">2.1 mm</quantity>
|
||
; A-MW:
|
||
<quantity id="4CBF9B0DFFE2FFE59096F9E1FE05A64A" box="[423,509,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="318" unit="mm" value="1.1">1.1 mm</quantity>
|
||
(NUM: 47 exs); very similar to
|
||
<taxonomicName id="4C474D6BFFE2FFE59254F9E0FC20A64A" box="[869,984,1628,1652]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE2FFE59254F9E0FC20A64A" box="[869,984,1628,1652]" italics="true" pageId="17" pageNumber="318">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
except for aedeagal characters, shape of prosternal process and, frequently, elytral colouration.
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFE2FFE591F6F919FAB3A716" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
|
||
<emphasis id="B933EAFAFFE2FFE591F6F919FEEDA682" box="[199,277,1700,1724]" italics="true" pageId="17" pageNumber="318">Colour</emphasis>
|
||
. Variable. Pale patterns of elytra normally (but not always) more sharply defined on darker background and often more extended with respect to
|
||
<taxonomicName id="4C474D6BFFE2FFE5935DF974FD19A6DE" box="[620,737,1736,1760]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE2FFE5935DF974FD19A6DE" box="[620,737,1736,1760]" italics="true" pageId="17" pageNumber="318">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
. Most commonly pale subbasal band of elytra fragmented in semicircular spots, although strong variability occurs (see below). Pronotum normally with narrower dark transversal band, more evidently bicolour; head, antennae, mouthparts, legs, and underside as in
|
||
<taxonomicName id="4C474D6BFFE2FFE595E0F8ACFABCA716" box="[1233,1348,1808,1832]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE2FFE595E0F8ACFABCA716" box="[1233,1348,1808,1832]" italics="true" pageId="17" pageNumber="318">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFE2FFE591F6F889FB80A74E" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
|
||
<emphasis id="B933EAFAFFE2FFE591F6F889FE18A772" box="[199,480,1844,1868]" italics="true" pageId="17" pageNumber="318">Structures and sculpture</emphasis>
|
||
. Mostly as in
|
||
<taxonomicName id="4C474D6BFFE2FFE593A0F888FCF0A772" box="[657,776,1844,1868]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE2FFE593A0F888FCF0A772" box="[657,776,1844,1868]" italics="true" pageId="17" pageNumber="318">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
, but prosternal process rounded at tip and elevated, not carinate (
|
||
<figureCitation id="137C2A6DFFE2FFE591CFF8EAFEAEA74E" box="[254,342,1879,1904]" captionStart="FIGURES 26 – 31" captionStartId="33.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@33.[151,1436,193,1932]" captionTargetPageId="33" captionText="FIGURES 26 – 31. SEMs of head, prosternum and prosternal process in Laccomimus species: 26) L. amazonas (male), oblique lateral view; 27) idem, at higher magnification; 28) L. alvarengi (female) ventral view; 29) L. bordoni (female), ventral view; 30) idem, oblique lateral view; 31) idem, at higher magnification." httpUri="https://zenodo.org/record/244306/files/figure.png" pageId="17" pageNumber="318">Figs 28</figureCitation>
|
||
; 69); iridescence on pronotum and mainly on elytra often more intense.
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFE2FFE691F6F8C1FAE3A0C6" blockId="17.[151,1437,151,2013]" lastBlockId="18.[151,1437,151,1112]" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318">
|
||
<emphasis id="B933EAFAFFE2FFE591F6F8C1FEF9A7AA" box="[199,257,1916,1940]" italics="true" pageId="17" pageNumber="318">Male</emphasis>
|
||
. Last abdominal ventrite as in
|
||
<emphasis id="B933EAFAFFE2FFE59369F8C0FD2BA7AA" box="[600,723,1916,1940]" italics="true" pageId="17" pageNumber="318">
|
||
<taxonomicName id="4C474D6BFFE2FFE59369F8C0FD36A7AA" box="[600,718,1916,1940]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pumilio">L. pumilio</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Outer claw of forelegs scimitar-shaped, variable in shape, often with denticles on ventral margin (
|
||
<figureCitation id="137C2A6DFFE2FFE5933FF81DFD68A786" box="[526,656,1952,1977]" captionStart="FIGURES 67 – 83" captionStartId="39.[151,264,1798,1820]" captionTargetBox="[159,1430,201,1748]" captionTargetId="figure@39.[151,1436,193,1777]" captionTargetPageId="39" captionText="FIGURES 67 – 83. Prosternal process and claws in Laccomimus species. 67 – 69) P rosternal process: 67) L. pumilio (Florida, USA); 68) L. pumilio (Cuba); 69) L. alvarengi (Perú). 70 – 83) M ale left protarsal claws: 70) L. bordoni (Paraguay); 71) L. malkini (Madre de Dios, Perú); 72) L. spangleri (holotype); 73) L. distinctus (holotype); 74) L. bolivari (Trinidad); 75) L. amazonas (holotype); 76) L. variegatus (Colonia Calleria, Perú); 77) L. alvarengi (holotype); 78) L. alvarengi (Loreto, Perú); 79) L. alvarengi (Guayaramerin, Bolivia); 80) L. alvarengi (Esmeraldas, Ecuador); 81) L. alvarengi (Albrook, Panamá); 82) L. pumilio (Cayo District, Belize); 83) L. pumilio (Cayamas, Cuba)." httpUri="https://zenodo.org/record/244312/files/figure.png" pageId="17" pageNumber="318">Figs 77–81</figureCitation>
|
||
). Median lobe of aedeagus in dorsal view as in
|
||
<figureCitation id="137C2A6DFFE2FFE59593F81DFB07A786" box="[1186,1279,1952,1977]" captionStart="FIGURES 107 – 115" captionStartId="46.[151,264,1951,1973]" captionTargetBox="[184,1390,196,1925]" captionTargetId="figure@46.[170,1417,193,1930]" captionTargetPageId="46" captionText="FIGURES 107 – 115. Median lobe of aedeagus in Laccomimus species, dorsal view: 107) L. distinctus; 108) L. malkini; 109) L. variegatus; 110) L. amazonas; 111) L. alvarengi; 112) L. improvidus; 113) L. bordoni; 114) L. spangleri; 115) L. bolivari." httpUri="https://zenodo.org/record/244319/files/figure.png" pageId="17" pageNumber="318">Fig. 111</figureCitation>
|
||
; same as in
|
||
<taxonomicName id="4C474D6BFFE2FFE594B4F81CFF15A7E2" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE2FFE594B4F81CFF15A7E2" italics="true" pageId="17" pageNumber="318">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
(see description above). In left lateral view variable (
|
||
<figureCitation id="137C2A6DFFE2FFE59274F879FC65A7E2" box="[837,925,1988,2013]" captionStart="FIGURES 94 – 97" captionStartId="43.[151,264,1919,1941]" captionTargetBox="[174,1408,200,1884]" captionTargetId="figure@43.[164,1418,193,1889]" captionTargetPageId="43" captionText="FIGURES 94 – 97. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 94) holotype, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 95) likely aberrant specimen from type locality, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 96) Loreto, Perú, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 97) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere." httpUri="https://zenodo.org/record/244316/files/figure.png" pageId="17" pageNumber="318">Figs 94</figureCitation>
|
||
a; 95a, 96a; 97a; 98a; 99a; 100a; 101a; 102a): in
|
||
<typeStatus id="54FC884AFFE1FFE69185FF2AFEEEA08E" box="[180,278,151,176]" pageId="18" pageNumber="319" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="137C2A6DFFE1FFE69017FF2AFE80A08E" box="[294,376,151,176]" captionStart="FIGURES 94 – 97" captionStartId="43.[151,264,1919,1941]" captionTargetBox="[174,1408,200,1884]" captionTargetId="figure@43.[164,1418,193,1889]" captionTargetPageId="43" captionText="FIGURES 94 – 97. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 94) holotype, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 95) likely aberrant specimen from type locality, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 96) Loreto, Perú, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 97) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere." httpUri="https://zenodo.org/record/244316/files/figure.png" pageId="18" pageNumber="319">Fig. 94</figureCitation>
|
||
a) base and distal portion forming a curve instead of an angle, distal portion less elongate with respect to
|
||
<taxonomicName id="4C474D6BFFE1FFE69025FF00FE74A0EA" box="[276,396,188,212]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE69025FF00FE74A0EA" box="[276,396,188,212]" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
and more uniformly wide along its length, although visibly sinuate on apical half. Left paramere (
|
||
<figureCitation id="137C2A6DFFE1FFE6903FFF62FE9DA0C6" box="[270,357,223,248]" captionStart="FIGURES 94 – 97" captionStartId="43.[151,264,1919,1941]" captionTargetBox="[174,1408,200,1884]" captionTargetId="figure@43.[164,1418,193,1889]" captionTargetPageId="43" captionText="FIGURES 94 – 97. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 94) holotype, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 95) likely aberrant specimen from type locality, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 96) Loreto, Perú, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 97) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere." httpUri="https://zenodo.org/record/244316/files/figure.png" pageId="18" pageNumber="319">Figs 94</figureCitation>
|
||
c; 95c; 96c; 97c; 98c; 101c; 102c) strongly sinuate apically, expanded and thickened.
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFE1FFE691F6FEB9FA93A122" blockId="18.[151,1437,151,1112]" box="[199,1387,260,285]" pageId="18" pageNumber="319">
|
||
<emphasis id="B933EAFAFFE1FFE691F6FEB9FEE3A122" box="[199,283,260,284]" italics="true" pageId="18" pageNumber="319">Female</emphasis>
|
||
. Often duller than males, with more intense iridescence. Genital pieces (
|
||
<figureCitation id="137C2A6DFFE1FFE69578FEB9FB5FA122" box="[1097,1191,260,285]" captionStart="FIGURES 116 – 121" captionStartId="47.[151,264,1829,1851]" captionTargetBox="[229,1344,199,1783]" captionTargetId="figure@47.[198,1370,193,1789]" captionTargetPageId="47" captionText="FIGURES 116 – 121. Female genitalia and cladogram. 116 – 120) Female genitalia in Laccomimus species, left lateral view: 116) L. distinctus (Paraguay), a) left gonocoxa, b) ramus; 117) L. malkini (Perú), a) left gonocoxa, b) ramus; 118) L. alvarengi (Brazil), a) left gonocoxa, b) ramus; 119) L. variegatus (Perú), a) left gonocoxa, b) ramus; 120) L. bordoni (Bolivia), ramus (arrow indicates lateral proximal process of ramus). 121) Single most parsimonious cladogram rooted between ingroup and outgroup, with character changes mapped for each clade. Solid rectangles indicate unique character state transformations; open rectangles indicate homoplasious character state transformations. Support values are indicated on each branch as Bremer / Bootstrap." httpUri="https://zenodo.org/record/244320/files/figure.png" pageId="18" pageNumber="319">Fig. 118</figureCitation>
|
||
) as in
|
||
<taxonomicName id="4C474D6BFFE1FFE695C0FEB8FA9DA122" box="[1265,1381,260,284]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE695C0FEB8FA9DA122" box="[1265,1381,260,284]" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D6563FFE1FFE691F6FE9AFB82A1B6" pageId="18" pageNumber="319" type="distribution">
|
||
<paragraph id="8BF836E8FFE1FFE691F6FE9AFB82A1B6" blockId="18.[151,1437,151,1112]" pageId="18" pageNumber="319">
|
||
<emphasis id="B933EAFAFFE1FFE691F6FE9AFE98A17E" bold="true" box="[199,352,295,320]" pageId="18" pageNumber="319">Distribution.</emphasis>
|
||
<taxonomicName id="4C474D6BFFE1FFE6905AFE95FD96A17E" box="[363,622,296,320]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="alvarengi">
|
||
<emphasis id="B933EAFAFFE1FFE6905AFE95FD96A17E" box="[363,622,296,320]" italics="true" pageId="18" pageNumber="319">Laccomimus alvarengi</emphasis>
|
||
</taxonomicName>
|
||
is one of the most widespread and variable member of
|
||
<taxonomicName id="4C474D6BFFE1FFE69437FE95FA6BA101" box="[1286,1427,296,319]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B933EAFAFFE1FFE69437FE95FA6BA101" box="[1286,1427,296,319]" italics="true" pageId="18" pageNumber="319">Laccomimus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="F3507678FFE1FFE691A6FEF1FEF1A15B" box="[151,265,332,357]" name="Argentina" pageId="18" pageNumber="319">Argentina</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507678FFE1FFE69024FEF1FE92A15B" box="[277,362,332,357]" name="Bolivia" pageId="18" pageNumber="319">Bolivia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507678FFE1FFE69046FEF1FE46A15B" box="[375,446,332,357]" name="Brazil" pageId="18" pageNumber="319">Brazil</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507678FFE1FFE690FAFEF1FDD2A15B" box="[459,554,332,357]" name="Ecuador" pageId="18" pageNumber="319">Ecuador</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507678FFE1FFE69306FEF1FD6BA15A" box="[567,659,332,356]" name="Panama" pageId="18" pageNumber="319">Panamá</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507678FFE1FFE693AEFEF0FCF0A15B" box="[671,776,333,357]" name="Paraguay" pageId="18" pageNumber="319">Paraguay</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507678FFE1FFE69224FEF1FCB3A15A" box="[789,843,332,356]" name="Peru" pageId="18" pageNumber="319">Perú</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507678FFE1FFE69266FEF1FC3AA15B" box="[855,962,332,357]" name="Suriname" pageId="18" pageNumber="319">Suriname</collectingCountry>
|
||
(
|
||
<figureCitation id="137C2A6DFFE1FFE692E3FEF1FBC2A15B" box="[978,1082,332,357]" captionStart="FIGURES 122 – 125" captionStartId="48.[151,264,1871,1893]" captionTargetBox="[157,1432,197,1844]" captionTargetId="figure@48.[151,1436,193,1850]" captionTargetPageId="48" captionText="FIGURES 122 – 125. Distribution maps of Laccomimus species: 122) L. distinctus; 123) L. malkini; 124) L. variegatus (open circles), L. spinosus (single pentagon in Brazil), L. amazonas (single star in Venezuela); 125) L. alvarengi in South America." httpUri="https://zenodo.org/record/244321/files/figure.png" pageId="18" pageNumber="319">Figs 125</figureCitation>
|
||
; 128). It has not been recorded from
|
||
<collectingCountry id="F3507678FFE1FFE691E5FED2FEB2A1B6" box="[212,330,367,392]" name="Venezuela" pageId="18" pageNumber="319">Venezuela</collectingCountry>
|
||
, however, in spite of VAIP’s intensive collections during the last decade.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D6563FFE1FFE691F6FE29FB1BA40A" pageId="18" pageNumber="319" type="etymology">
|
||
<paragraph id="8BF836E8FFE1FFE691F6FE29FF37A1CA" blockId="18.[151,1437,151,1112]" pageId="18" pageNumber="319">
|
||
<emphasis id="B933EAFAFFE1FFE691F6FE29FEB6A193" bold="true" box="[199,334,404,429]" pageId="18" pageNumber="319">Etymology.</emphasis>
|
||
Frank N. Young and Paul J. Spangler named,
|
||
<emphasis id="B933EAFAFFE1FFE69262FE28FC40A192" box="[851,952,404,428]" italics="true" pageId="18" pageNumber="319">in litteris</emphasis>
|
||
, this taxon “
|
||
<taxonomicName id="4C474D6BFFE1FFE69576FE28FABEA192" box="[1095,1350,404,428]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="alvarengi" status="sp. nov.">
|
||
<emphasis id="B933EAFAFFE1FFE69576FE28FABEA192" box="[1095,1350,404,428]" italics="true" pageId="18" pageNumber="319">Laccomimus alvarengi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2005781FFE1FFE6947CFE28FA72A192" box="[1357,1418,404,428]" pageId="18" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
|
||
”, in honour of the collector Moacir Alvarenga (cf.
|
||
<bibRefCitation id="EFD64B19FFE1FFE693F4FE05FCAAA1EE" author="Young" box="[709,850,440,464]" pageId="18" pageNumber="319" refString="Young, F. N. (1974) Review of the predaceous water beetles of genus Anodocheilus (Coleoptera: Dytiscidae: Hydroporinae). Occasional Papers of the Museum of Zoology of the University of Michigan, 670, 1 - 28." type="journal article" year="1974">Young 1974</bibRefCitation>
|
||
: 2, 3). The specific epithet is a noun in the genitive case.
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFE1FFE691F6FE42FB1BA40A" blockId="18.[151,1437,151,1112]" pageId="18" pageNumber="319">
|
||
<emphasis id="B933EAFAFFE1FFE691F6FE42FE4BA226" bold="true" box="[199,435,511,536]" pageId="18" pageNumber="319">Notes on variability</emphasis>
|
||
.
|
||
<taxonomicName id="4C474D6BFFE1FFE690F3FDBCFDB3A226" box="[450,587,512,536]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="alvarengi">
|
||
<emphasis id="B933EAFAFFE1FFE690F3FDBCFDB3A226" box="[450,587,512,536]" italics="true" pageId="18" pageNumber="319">L. alvarengi</emphasis>
|
||
</taxonomicName>
|
||
is more variable than
|
||
<taxonomicName id="4C474D6BFFE1FFE6927DFDBCFC38A226" box="[844,960,512,536]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE6927DFDBCFC38A226" box="[844,960,512,536]" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
. Except for the striking differences in the prosternal process, morphological separation of the two species is often difficult. The visibly sinuate and expanded apex of the left paramere and the less broadened distal portion of the median lobe seem to be constant diagnostic characters of
|
||
<taxonomicName id="4C474D6BFFE1FFE69003FDD0FE44A2BA" box="[306,444,620,644]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="alvarengi">
|
||
<emphasis id="B933EAFAFFE1FFE69003FDD0FE44A2BA" box="[306,444,620,644]" italics="true" pageId="18" pageNumber="319">L. alvarengi</emphasis>
|
||
</taxonomicName>
|
||
with respect to
|
||
<taxonomicName id="4C474D6BFFE1FFE69346FDD0FD14A2BA" box="[631,748,620,644]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE69346FDD0FD14A2BA" box="[631,748,620,644]" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
. In other features the median lobe is very variable, and this variability has been observed in different populations. Intermediate forms between the
|
||
<typeStatus id="54FC884AFFE1FFE6956EFD32FB39A296" box="[1119,1217,655,680]" pageId="18" pageNumber="319" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C474D6BFFE1FFE695D4FD2CFA95A296" box="[1253,1389,656,680]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="alvarengi">
|
||
<emphasis id="B933EAFAFFE1FFE695D4FD2CFA95A296" box="[1253,1389,656,680]" italics="true" pageId="18" pageNumber="319">L. alvarengi</emphasis>
|
||
</taxonomicName>
|
||
and Florida specimens of
|
||
<taxonomicName id="4C474D6BFFE1FFE690BAFD08FE06A2F2" box="[395,510,692,716]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE690BAFD08FE06A2F2" box="[395,510,692,716]" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
have been observed frequently, even in exemplars from the same locality (see for example
|
||
<figureCitation id="137C2A6DFFE1FFE691CCFD6AFEA9A2CE" box="[253,337,727,752]" captionStart="FIGURES 94 – 97" captionStartId="43.[151,264,1919,1941]" captionTargetBox="[174,1408,200,1884]" captionTargetId="figure@43.[164,1418,193,1889]" captionTargetPageId="43" captionText="FIGURES 94 – 97. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 94) holotype, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 95) likely aberrant specimen from type locality, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 96) Loreto, Perú, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 97) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere." httpUri="https://zenodo.org/record/244316/files/figure.png" pageId="18" pageNumber="319">Fig. 97</figureCitation>
|
||
). In most cases, however, the aedeagus is distinctive with respect to
|
||
<taxonomicName id="4C474D6BFFE1FFE69563FD64FB3DA2CE" box="[1106,1221,728,752]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE69563FD64FB3DA2CE" box="[1106,1221,728,752]" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
, particularly in the angulated separation between the base and distal portion and the somewhat widened distal portion. An aberrant median lobe was found dissected from a specimen collected at the
|
||
<typeStatus id="54FC884AFFE1FFE692B8FC9DFC41A306" box="[905,953,800,824]" pageId="18" pageNumber="319">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="137C2A6DFFE1FFE69514FCA2FB82A306" box="[1061,1146,799,824]" captionStart="FIGURES 94 – 97" captionStartId="43.[151,264,1919,1941]" captionTargetBox="[174,1408,200,1884]" captionTargetId="figure@43.[164,1418,193,1889]" captionTargetPageId="43" captionText="FIGURES 94 – 97. Male genitalia in Laccomimus alvarengi from different localities, left lateral view: 94) holotype, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 95) likely aberrant specimen from type locality, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 96) Loreto, Perú, a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 97) Mbrucuyá National Park, Argentina, a) median lobe, b) left paramere." httpUri="https://zenodo.org/record/244316/files/figure.png" pageId="18" pageNumber="319">Fig. 95</figureCitation>
|
||
). Colouration is also very variable. Normally the elytral patterns in
|
||
<taxonomicName id="4C474D6BFFE1FFE6935FFCF8FD00A362" box="[622,760,836,860]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="alvarengi">
|
||
<emphasis id="B933EAFAFFE1FFE6935FFCF8FD00A362" box="[622,760,836,860]" italics="true" pageId="18" pageNumber="319">L. alvarengi</emphasis>
|
||
</taxonomicName>
|
||
are more defined than in
|
||
<taxonomicName id="4C474D6BFFE1FFE69510FCF8FB6DA362" box="[1057,1173,836,860]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE69510FCF8FB6DA362" box="[1057,1173,836,860]" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
, and some populations exhibit very bright elytral patterns, including large and complete pale subbasal bands (
|
||
<figureCitation id="137C2A6DFFE1FFE69593FCDAFB01A3BE" box="[1186,1273,871,896]" captionStart="FIGURES 59 – 66" captionStartId="38.[151,264,1953,1975]" captionTargetBox="[161,1426,197,1927]" captionTargetId="figure@38.[151,1435,193,1932]" captionTargetPageId="38" captionText="FIGURES 59 – 66. Subschematic body shape and colour patterns in Laccomimus species: 59) L. distinctus; 60) L. malkini, a) typical specimen from Colombia, b) specimen from Piracicaba, Brazil; 61) L. variegatus (Perú); 62) L. amazonas; 63) L. pumilio (Florida, USA); 64) L. alvarengi, a – b) variation in colouration patterns; 65) L. bordoni; 66) L. spangleri." httpUri="https://zenodo.org/record/244311/files/figure.png" pageId="18" pageNumber="319">Fig. 64</figureCitation>
|
||
a, b).We have observed, however, specimens (especially from
|
||
<collectingCountry id="F3507678FFE1FFE693FDFC31FCEAA39B" box="[716,786,908,933]" name="Brazil" pageId="18" pageNumber="319">Brazil</collectingCountry>
|
||
) with obscure or reduced pale markings, similar to
|
||
<taxonomicName id="4C474D6BFFE1FFE694B4FC30FF14A3F6" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE694B4FC30FF14A3F6" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
. Uniformly dark-chestnut elytra occur in few specimens from
|
||
<collectingCountry id="F3507678FFE1FFE69280FC12FC0DA3F6" box="[945,1013,943,968]" name="Brazil" pageId="18" pageNumber="319">Brazil</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3507678FFE1FFE6951FFC0DFB60A3F6" box="[1070,1176,944,968]" name="Paraguay" pageId="18" pageNumber="319">Paraguay</collectingCountry>
|
||
. Since the shape of the prosternal process unambiguously separates
|
||
<taxonomicName id="4C474D6BFFE1FFE693BBFC68FCEAA3D2" box="[650,786,980,1004]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="alvarengi">
|
||
<emphasis id="B933EAFAFFE1FFE693BBFC68FCEAA3D2" box="[650,786,980,1004]" italics="true" pageId="18" pageNumber="319">L. alvarengi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C474D6BFFE1FFE69266FC68FC32A3D2" box="[855,970,980,1004]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE69266FC68FC32A3D2" box="[855,970,980,1004]" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
, and a visibly sinuate and expanded apex of the left paramere and a less broadened distal portion of the median lobe are also apparently constant differences, we decided to describe
|
||
<taxonomicName id="4C474D6BFFE1FFE690ACFBA0FDDDA40A" box="[413,549,1052,1076]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="alvarengi">
|
||
<emphasis id="B933EAFAFFE1FFE690ACFBA0FDDDA40A" box="[413,549,1052,1076]" italics="true" pageId="18" pageNumber="319">L. alvarengi</emphasis>
|
||
</taxonomicName>
|
||
as a distinct species and not as a subspecies within
|
||
<taxonomicName id="4C474D6BFFE1FFE69558FBA0FB23A40A" box="[1129,1243,1052,1076]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE69558FBA0FB23A40A" box="[1129,1243,1052,1076]" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D6563FFE1FFE691F6FB82FDCAA469" box="[199,562,1087,1112]" pageId="18" pageNumber="319" type="discussion">
|
||
<paragraph id="8BF836E8FFE1FFE691F6FB82FDCAA469" blockId="18.[151,1437,151,1112]" box="[199,562,1087,1112]" pageId="18" pageNumber="319">
|
||
<emphasis id="B933EAFAFFE1FFE691F6FB82FEC3A466" bold="true" box="[199,315,1087,1112]" pageId="18" pageNumber="319">Remarks.</emphasis>
|
||
See under
|
||
<taxonomicName id="4C474D6BFFE1FFE69089FBFCFDD2A466" box="[440,554,1088,1112]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE1FFE69089FBFCFDD2A466" box="[440,554,1088,1112]" italics="true" pageId="18" pageNumber="319">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |