treatments-xml/data/03/EE/87/03EE87F7FF8BDE43FF25D7E7FAA9AB0E.xml
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<document id="911D73BED1904461A96D68E1D302A946" ID-CLB-Dataset="31536" ID-DOI="10.11646/zootaxa.4349.1.1" ID-GBIF-Dataset="3e5cd781-3051-4527-bace-d222e83d8398" ID-ISSN="1175-5326" ID-Zenodo-Dep="1049190" ID-ZooBank="DDFDC23A-FB21-41E2-B38B-A0FD19F5BFAE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510732256065" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2017" docId="03EE87F7FF8BDE43FF25D7E7FAA9AB0E" docLanguage="en" docName="zootaxa.4349.1.1.pdf.imf" docOrigin="Zootaxa 4349 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cephennodes (Cephennodes) manangensis Jałoszyński 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="43" masterDocId="FFD7FF8FFFA3DE69FFB2D111FFADAD5E" masterDocTitle="Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="120" masterPageNumber="1" pageNumber="41" updateTime="1698484713136" updateUser="ExternalLinkService">
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<mods:title id="ED51C2BA8F435D206FB917C22430ADB6">Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="08B9E33C1615D3D63452ADE607A098A2">2017</mods:date>
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<treatment id="03EE87F7FF8BDE43FF25D7E7FAA9AB0E" ID-DOI="http://doi.org/10.5281/zenodo.6026732" ID-GBIF-Taxon="137306731" ID-Zenodo-Dep="6026732" LSID="urn:lsid:plazi:treatment:03EE87F7FF8BDE43FF25D7E7FAA9AB0E" httpUri="http://treatment.plazi.org/id/03EE87F7FF8BDE43FF25D7E7FAA9AB0E" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection id="C35D656AFF8BDE41FF25D7E7FDBEAA6C" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8BF836E1FF8BDE41FF25D7E7FD45AA4F" blockId="40.[151,744,1782,1842]" box="[151,744,1782,1809]" pageId="40" pageNumber="41">
<heading id="D0B0818DFF8BDE41FF25D7E7FD45AA4F" bold="true" box="[151,744,1782,1809]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<emphasis id="B933EAF3FF8BDE41FF25D7E7FD45AA4F" bold="true" box="[151,744,1782,1809]" pageId="40" pageNumber="41">
<taxonomicName id="4C474D62FF8BDE41FF25D7E7FD30AA4E" ID-CoL="SFH2" authority="Jałoszyński, 2017" authorityName="Jałoszyński" authorityYear="2017" box="[151,669,1782,1809]" class="Insecta" family="Staphylinidae" genus="Cephennodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="manangensis" status="sp. nov." subGenus="Cephennodes">
<emphasis id="B933EAF3FF8BDE41FF25D7E7FE97AA4E" bold="true" box="[151,314,1782,1808]" italics="true" pageId="40" pageNumber="41">Cephennodes</emphasis>
(
<emphasis id="B933EAF3FF8BDE41FEF9D7E7FE43AA4E" bold="true" box="[331,494,1782,1808]" italics="true" pageId="40" pageNumber="41">Cephennodes</emphasis>
)
<emphasis id="B933EAF3FF8BDE41FE4CD7E7FD30AA4E" bold="true" box="[510,669,1782,1808]" italics="true" pageId="40" pageNumber="41">manangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2005788FF8BDE41FD17D7E6FD45AA4F" box="[677,744,1783,1809]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF836E1FF8BDE41FF25D608FDBEAA6C" blockId="40.[151,744,1782,1842]" box="[151,531,1817,1842]" pageId="40" pageNumber="41">
(
<figureCitation id="137C2A64FF8BDE41FF2DD608FF58AA6C" box="[159,245,1817,1842]" captionStart="FIGURES 94 101" captionStartId="40.[151,264,1618,1641]" captionTargetBox="[159,1423,204,1583]" captionTargetId="figure@40.[151,1436,193,1597]" captionTargetPageId="40" captionText="FIGURES 94 101. Dorsal habitus of Cephennodes (s. str.) dolakhanus s sp. n. (94), C. (s. str.) manangensis sp. n. (95), C. (s. str.) martensi sp. n. (96), C. (s. str.) paramartensi sp. n. (97), C. (s. str.) monolaminatus sp. n. (98), C. (s. str.) thakanus sp. n. (99), C. (s. str.) annapurnaensis sp. n. (100), and C. (s. str.) parbatensis sp. n. (101)." httpUri="https://zenodo.org/record/1049224/files/figure.png" pageId="40" pageNumber="41">Figs 95</figureCitation>
,
<figureCitation id="137C2A64FF8BDE41FEB3D60BFEC8AA6C" box="[257,357,1817,1842]" captionStart="FIGURES 102 111" captionStartId="41.[151,264,1919,1942]" captionTargetBox="[151,1436,423,1895]" captionTargetId="figure@41.[151,1436,401,1908]" captionTargetPageId="41" captionText="FIGURES 102 111. Right metatibia of male in dorsal view. Cephennodes (s. str.) dolakhanus s sp. n. (102, 104), C. (s. str.) manangensis sp. n. (103, 105), C. (s. str.) martensi sp. n. (106), C. (s. str.) paramartensi sp. n. (107), C. (s. str.) monolaminatus sp. n. (108), C. (s. str.) thakanus sp. n. (109), C. (s. str.) annapurnaensis sp. n. (110), and C. (s. str.) parbatensis sp. n. (111). 104 111 not to the same scale." httpUri="https://zenodo.org/record/1049226/files/figure.png" pageId="40" pageNumber="41">103, 105</figureCitation>
,
<figureCitation id="137C2A64FF8BDE41FEC3D608FE78AA6C" box="[369,469,1817,1842]" captionStart="FIGURES 112 127" captionStartId="45.[151,264,1763,1786]" captionTargetBox="[158,1435,194,1728]" captionTargetId="figure@45.[151,1436,193,1742]" captionTargetPageId="45" captionText="FIGURES 112 127. Abdomen of male in right lateral (112, 116, 120, 124) and ventral (113, 117, 121, 125) views and aedeagus in ventral (114, 118, 122, 126) and lateral (115, 119, 123, 127) views. Cephennodes (s. str.) dolakhanus sp. n. (112 115), C. (s. str.) manangensis sp. n. (116 119), C. (s. str.) martensi sp. n. (120 123), C. (s. str.) paramartensi sp. n. (124 127). Abbreviations: aml, apex of median lobe; ap, apical projection." httpUri="https://zenodo.org/record/1049228/files/figure.png" pageId="40" pageNumber="41">116119</figureCitation>
,
<figureCitation id="137C2A64FF8BDE41FE6DD60BFDA7AA6C" box="[479,522,1818,1842]" captionStart="FIGURES 144 145" captionStartId="51.[151,264,1344,1367]" captionTargetBox="[151,1436,183,1326]" captionTargetId="figure@51.[151,1436,183,1334]" captionTargetPageId="51" captionText="FIGURES 144 145. Distribution of Cephennodes (s. str.) dolakhanus sp. n. (144, circle), C. (s. str.) manangensis sp. n. (144, triangle), C. (s. str.) martensi sp. n. (145, circle), C. (s. str.) paramartensi sp. n. (145, triangle)." httpUri="https://zenodo.org/record/1049232/files/figure.png" pageId="40" pageNumber="41">144</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35D656AFF8BDE41FF25D670FE23AA9C" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8BF836E1FF8BDE41FF25D670FE23AA9C" blockId="40.[151,1436,1889,2022]" pageId="40" pageNumber="41">
<emphasis id="B933EAF3FF8BDE41FF25D670FCA9AA24" bold="true" box="[151,772,1889,1914]" pageId="40" pageNumber="41">
<typeStatus id="54FC8843FF8BDE41FF25D670FF7FAA24" box="[151,210,1889,1914]" pageId="40" pageNumber="41">Type</typeStatus>
material.
<typeStatus id="54FC8843FF8BDE41FEFED670FE10AA24" box="[332,445,1889,1914]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3507671FF8BDE41FE7BD670FD8BAA24" box="[457,550,1889,1914]" name="Nepal" pageId="40" pageNumber="41">NEPAL</collectingCountry>
(Manang District)
</emphasis>
:
<specimenCount id="9D41FD68FF8BDE41FCA6D673FC86AA27" box="[788,811,1890,1913]" pageId="40" pageNumber="41" type="male"></specimenCount>
, one label: &quot;
<collectingCountry id="F3507671FF8BDE41FC06D673FBBFAA24" box="[948,1042,1890,1914]" name="Nepal" pageId="40" pageNumber="41">NEPAL</collectingCountry>
, Manang Distr. / above Bagarchap,
<quantity id="4CBF9B04FF8BDE41FF25D697FF41AAC3" box="[151,236,1926,1950]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="40" pageNumber="41" unit="m" value="2200.0">2200 m</quantity>
/
<date id="FFF91021FF8BDE41FEB0D697FE02AAC0" box="[258,431,1926,1950]" pageId="40" pageNumber="41" value="1980-04-12" valueMax="1980-04-13" valueMin="1980-04-12">12.13.IV.1980</date>
/ leg. J. Martens &amp;
<collectingRegion id="4983F803FF8BDE41FD3DD697FD0BAAC0" box="[655,678,1926,1950]" country="Nigeria" name="Rivers" pageId="40" pageNumber="41">A</collectingRegion>
. Ausobsky&quot; [white, printed] (
<collectionCode id="ED56AE24FF8BDE41FC41D697FBE8AAC0" box="[1011,1093,1926,1950]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="40" pageNumber="41">SMNS</collectionCode>
).
<emphasis id="B933EAF3FF8BDE41FBEAD694FB63AAC0" bold="true" box="[1112,1230,1925,1950]" pageId="40" pageNumber="41">
<typeStatus id="54FC8843FF8BDE41FBEAD694FB64AAC0" box="[1112,1225,1925,1950]" pageId="40" pageNumber="41" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D41FD68FF8BDE41FB67D697FAAFAAC3" box="[1237,1282,1926,1950]" pageId="40" pageNumber="41" type="male">1 ♂</specimenCount>
, same data as for
<typeStatus id="54FC8843FF8BDE41FF0CD6B8FE8DAA9C" box="[190,288,1961,1986]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED56AE24FF8BDE41FE82D6BBFE2FAA9C" box="[304,386,1962,1986]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="40" pageNumber="41">SMNS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35D656AFF8BDE40FF75D6DCFD94AC43" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="8BF836E1FF8BDE40FF75D6DCFD94AC43" blockId="40.[151,1436,1889,2022]" lastBlockId="41.[151,1437,151,357]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<emphasis id="B933EAF3FF8BDE41FF75D6DCFEECAAB8" bold="true" box="[199,321,1997,2022]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
BL
<quantity id="4CBF9B04FF8BDE41FEC0D6DCFDBAAABB" box="[370,535,1997,2022]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="40" pageNumber="41" unit="mm" value="1.275" valueMax="1.3" valueMin="1.25">1.251.30 mm</quantity>
; body relatively slender; distal portion of metatibia strongly thickened, with oval subapical porous field much longer than broad and lacking one long bristle within short setal fringe of apical margin; abdomen in ventral view with sternite III bearing broadly rounded median lobe; apical portion of median lobe of aedeagus evenly rounded, its longitudinal axis forming about 45°-angle with long axis of aedeagus; apical projection with sinuate distal margin.
</paragraph>
</subSubSection>
<subSubSection id="C35D656AFF8ADE43FF75D036FE33A8C3" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="8BF836E1FF8ADE40FF75D036FB6BAC3A" blockId="41.[151,1437,151,357]" pageId="41" pageNumber="42">
<emphasis id="B933EAF3FF8ADE40FF75D036FEF4AC1E" bold="true" box="[199,345,295,320]" pageId="41" pageNumber="42">Description.</emphasis>
Body of male (
<figureCitation id="137C2A64FF8ADE40FDB4D036FDFAAC1E" box="[518,599,295,320]" captionStart="FIGURES 94 101" captionStartId="40.[151,264,1618,1641]" captionTargetBox="[159,1423,204,1583]" captionTargetId="figure@40.[151,1436,193,1597]" captionTargetPageId="40" captionText="FIGURES 94 101. Dorsal habitus of Cephennodes (s. str.) dolakhanus s sp. n. (94), C. (s. str.) manangensis sp. n. (95), C. (s. str.) martensi sp. n. (96), C. (s. str.) paramartensi sp. n. (97), C. (s. str.) monolaminatus sp. n. (98), C. (s. str.) thakanus sp. n. (99), C. (s. str.) annapurnaensis sp. n. (100), and C. (s. str.) parbatensis sp. n. (101)." httpUri="https://zenodo.org/record/1049224/files/figure.png" pageId="41" pageNumber="42">Fig. 95</figureCitation>
) moderately dark brown, with shallow constriction between pronotum and elytra, moderately convex, covered with light brown setae; BL
<quantity id="4CBF9B04FF8ADE40FCE9D05DFC52AC3A" box="[859,1023,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="41" pageNumber="42" unit="mm" value="1.275" valueMax="1.3" valueMin="1.25">1.251.30 mm</quantity>
(mean
<quantity id="4CBF9B04FF8ADE40FBE0D05DFB17AC3A" box="[1106,1210,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="41" pageNumber="42" unit="mm" value="1.28">1.28 mm</quantity>
).
</paragraph>
<caption id="DF386669FF8ADE40FF25D66EFE63AAAC" httpUri="https://zenodo.org/record/1049226/files/figure.png" pageId="41" pageNumber="42" targetBox="[151,1436,423,1895]" targetPageId="41">
<paragraph id="8BF836E1FF8ADE40FF25D66EFE63AAAC" blockId="41.[151,1436,1919,2034]" pageId="41" pageNumber="42">
<emphasis id="B933EAF3FF8ADE40FF25D66EFEC6AACA" bold="true" box="[151,363,1919,1942]" pageId="41" pageNumber="42">FIGURES 102111.</emphasis>
Right metatibia of male in dorsal view.
<taxonomicName id="4C474D62FF8ADE40FCBFD66EFBCCAACB" authorityName="Jałoszyński" authorityYear="2017" box="[781,1121,1919,1941]" class="Insecta" family="Staphylinidae" genus="Cephennodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dolakhanus">
<emphasis id="B933EAF3FF8ADE40FCBFD66EFC3BAACB" box="[781,918,1919,1941]" italics="true" pageId="41" pageNumber="42">Cephennodes</emphasis>
(s. str.)
<emphasis id="B933EAF3FF8ADE40FC5ED66EFBCCAACB" box="[1004,1121,1919,1941]" italics="true" pageId="41" pageNumber="42">dolakhanus</emphasis>
</taxonomicName>
<emphasis id="B933EAF3FF8ADE40FBDBD690FBDEAAC8" box="[1129,1139,1921,1942]" italics="true" pageId="41" pageNumber="42">s</emphasis>
<emphasis id="B933EAF3FF8ADE40FBC9D691FB18AACB" bold="true" box="[1147,1205,1920,1941]" pageId="41" pageNumber="42">sp. n.</emphasis>
(102, 104),
<taxonomicName id="4C474D62FF8ADE40FA84D691FEB6AAEA" authority="Jałoszyński, 2017" authorityName="Jałoszyński" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Cephennodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="manangensis" status="sp. nov.">
<emphasis id="B933EAF3FF8ADE40FA84D691FAE3AACB" box="[1334,1358,1920,1941]" italics="true" pageId="41" pageNumber="42">C.</emphasis>
(s. str.)
<emphasis id="B933EAF3FF8ADE40FF25D68EFEB6AAEA" box="[151,283,1951,1972]" italics="true" pageId="41" pageNumber="42">manangensis</emphasis>
</taxonomicName>
<emphasis id="B933EAF3FF8ADE40FE93D68EFEF7AAEA" bold="true" box="[289,346,1951,1972]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A2005788FF8ADE40FE93D68EFEF7AAEA" box="[289,346,1951,1972]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(103, 105),
<taxonomicName id="4C474D62FF8ADE40FE67D68EFD3AAAEA" authority="Jałoszyński, 2017" authorityName="Jałoszyński" authorityYear="2017" box="[469,663,1950,1972]" class="Insecta" family="Staphylinidae" genus="Cephennodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="B933EAF3FF8ADE40FE67D68EFE40AAEA" box="[469,493,1951,1972]" italics="true" pageId="41" pageNumber="42">C.</emphasis>
(s. str.)
<emphasis id="B933EAF3FF8ADE40FD8CD68EFD3AAAEA" box="[574,663,1951,1972]" italics="true" pageId="41" pageNumber="42">martensi</emphasis>
</taxonomicName>
<emphasis id="B933EAF3FF8ADE40FD2ED68EFD78AAEA" bold="true" box="[668,725,1951,1972]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A2005788FF8ADE40FD2ED68EFD78AAEA" box="[668,725,1951,1972]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(106),
<taxonomicName id="4C474D62FF8ADE40FCACD68EFBA2AAEA" authority="Jałoszyński, 2017" authorityName="Jałoszyński" authorityYear="2017" box="[798,1039,1950,1972]" class="Insecta" family="Staphylinidae" genus="Cephennodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="paramartensi" status="sp. nov.">
<emphasis id="B933EAF3FF8ADE40FCACD68EFC9BAAEA" box="[798,822,1951,1972]" italics="true" pageId="41" pageNumber="42">C.</emphasis>
(s. str.)
<emphasis id="B933EAF3FF8ADE40FC35D68EFBA2AAEA" box="[903,1039,1951,1972]" italics="true" pageId="41" pageNumber="42">paramartensi</emphasis>
</taxonomicName>
<emphasis id="B933EAF3FF8ADE40FBA7D68EFBE3AAEA" bold="true" box="[1045,1102,1951,1972]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A2005788FF8ADE40FBA7D68EFBE3AAEA" box="[1045,1102,1951,1972]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(107),
<taxonomicName id="4C474D62FF8ADE40FB25D68EFA31AAEA" authority="Jałoszyński, 2017" authorityName="Jałoszyński" authorityYear="2017" box="[1175,1436,1950,1972]" class="Insecta" family="Staphylinidae" genus="Cephennodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="monolaminatus" status="sp. nov.">
<emphasis id="B933EAF3FF8ADE40FB25D68EFB02AAEA" box="[1175,1199,1951,1972]" italics="true" pageId="41" pageNumber="42">C.</emphasis>
(s. str.)
<emphasis id="B933EAF3FF8ADE40FAB2D68FFA31AAEA" box="[1280,1436,1950,1972]" italics="true" pageId="41" pageNumber="42">monolaminatus</emphasis>
</taxonomicName>
<emphasis id="B933EAF3FF8ADE40FF25D6ACFF7CAA8C" bold="true" box="[151,209,1981,2002]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A2005788FF8ADE40FF25D6ACFF7CAA8C" box="[151,209,1981,2002]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(108),
<taxonomicName id="4C474D62FF8ADE40FEAED6AFFE49AA8D" authority="Jałoszyński, 2017" authorityName="Jałoszyński" authorityYear="2017" box="[284,484,1981,2003]" class="Insecta" family="Staphylinidae" genus="Cephennodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="thakanus" status="sp. nov.">
<emphasis id="B933EAF3FF8ADE40FEAED6AFFE99AA8D" box="[284,308,1982,2003]" italics="true" pageId="41" pageNumber="42">C.</emphasis>
(s. str.)
<emphasis id="B933EAF3FF8ADE40FE3AD6ACFE49AA8D" box="[392,484,1981,2003]" italics="true" pageId="41" pageNumber="42">thakanus</emphasis>
</taxonomicName>
<emphasis id="B933EAF3FF8ADE40FE59D6ACFD88AA8C" bold="true" box="[491,549,1981,2002]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A2005788FF8ADE40FE59D6ACFD88AA8C" box="[491,549,1981,2002]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(109),
<taxonomicName id="4C474D62FF8ADE40FDC3D6AFFCD3AA8D" authority="Jałoszyński, 2017" authorityName="Jałoszyński" authorityYear="2017" box="[625,894,1981,2003]" class="Insecta" family="Staphylinidae" genus="Cephennodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="annapurnaensis" status="sp. nov.">
<emphasis id="B933EAF3FF8ADE40FDC3D6AFFD25AA8D" box="[625,648,1982,2003]" italics="true" pageId="41" pageNumber="42">C.</emphasis>
(s. str.)
<emphasis id="B933EAF3FF8ADE40FD6FD6AFFCD3AA8D" box="[733,894,1982,2003]" italics="true" pageId="41" pageNumber="42">annapurnaensis</emphasis>
</taxonomicName>
<emphasis id="B933EAF3FF8ADE40FC37D6ACFC12AA8C" bold="true" box="[901,959,1981,2002]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A2005788FF8ADE40FC37D6ACFC12AA8C" box="[901,959,1981,2002]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(110), and
<taxonomicName id="4C474D62FF8ADE40FB84D6AFFAB5AA8D" authority="Jałoszyński, 2017" authorityName="Jałoszyński" authorityYear="2017" box="[1078,1304,1981,2003]" class="Insecta" family="Staphylinidae" genus="Cephennodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="parbatensis" status="sp. nov.">
<emphasis id="B933EAF3FF8ADE40FB84D6AFFBE0AA8D" box="[1078,1101,1982,2003]" italics="true" pageId="41" pageNumber="42">C.</emphasis>
(s. str.)
<emphasis id="B933EAF3FF8ADE40FB10D6ACFAB5AA8D" box="[1186,1304,1981,2003]" italics="true" pageId="41" pageNumber="42">parbatensis</emphasis>
</taxonomicName>
<emphasis id="B933EAF3FF8ADE40FAADD6ACFAF4AA8C" bold="true" box="[1311,1369,1981,2002]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A2005788FF8ADE40FAADD6ACFAF4AA8C" box="[1311,1369,1981,2002]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(111). 104111 not to the same scale.
</paragraph>
</caption>
<paragraph id="8BF836E1FF89DE43FF75D186FE38AF46" blockId="42.[151,1437,151,1616]" pageId="42" pageNumber="43">
Head subtrapezoidal, HL
<quantity id="4CBF9B04FF89DE43FE58D189FDFCADF1" box="[490,593,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="42" pageNumber="43" unit="mm" value="0.18">0.18 mm</quantity>
, HW
<quantity id="4CBF9B04FF89DE43FD26D189FC95ADF1" box="[660,824,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.7999999999999998" pageId="42" pageNumber="43" unit="mm" value="0.19" valueMax="0.2" valueMin="0.18">0.180.20 mm</quantity>
(mean
<quantity id="4CBF9B04FF89DE43FC3CD189FC5AADEE" box="[910,1015,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="42" pageNumber="43" unit="mm" value="0.19">0.19 mm</quantity>
); vertex broadly rounded, with a pair of tiny tubercles; frons flattened; supraantennal tubercles distinct but weakly elevated; eyes large and strongly convex, coarsely faceted. Punctures on vertex and frons distinct, small but sharply marked and densely, evenly distributed, separated by spaces as wide as 12 × diameters of puncture; setae short, moderately dense, suberect. Antennae short, AnL
<quantity id="4CBF9B04FF89DE43FE10D036FDBDAC61" box="[418,528,295,320]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="42" pageNumber="43" unit="mm" value="0.63">0.63 mm</quantity>
, compact, pentamerous club indistinctly delimited and moderately strongly thickened; antennomere I twice as long as broad; II slightly shorter and narrower than I, 1.8 × as long as broad; III IV each slightly narrower than II, subquadrate; V as broad as IV but slightly longer, 1.2 × as long as broad; VI as broad as V but slightly shorter, subquadrate; VII slightly broader and longer than VI, as long as broad; VIII slightly broader but minimally shorter than VII, slightly transverse; IX much larger than VIII, as long as broad; X distinctly larger than IX, slightly broader than long; XI distinctly broader than X, slightly longer than IXX together, about 2.2 × as long as broad.
</paragraph>
<paragraph id="8BF836E1FF89DE43FF75D334FBF0AFAE" blockId="42.[151,1437,151,1616]" pageId="42" pageNumber="43">
Pronotum in dorsal view nearly semicircular, broadest slightly in front of middle, PL
<quantity id="4CBF9B04FF89DE43FB15D335FAE0AF62" box="[1191,1357,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" metricValueMax="4.0" metricValueMin="3.8" pageId="42" pageNumber="43" unit="mm" value="0.39" valueMax="0.4" valueMin="0.38">0.380.40 mm</quantity>
(mean
<quantity id="4CBF9B04FF89DE43FF25D359FEA9AF3E" box="[151,260,584,608]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="42" pageNumber="43" unit="mm" value="0.39">0.39 mm</quantity>
), PW
<quantity id="4CBF9B04FF89DE43FEE2D356FE55AF01" box="[336,504,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" metricValueMax="6.0" metricValueMin="5.8" pageId="42" pageNumber="43" unit="mm" value="0.59" valueMax="0.6" valueMin="0.58">0.580.60 mm</quantity>
(mean
<quantity id="4CBF9B04FF89DE43FDE7D356FD6CAF3E" box="[597,705,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="42" pageNumber="43" unit="mm" value="0.59">0.59 mm</quantity>
); anterior margin broadly rounded; lateral margins indistinctly microserrate, rounded in anterior half, then nearly straight and slightly convergent toward obtuse-angled hind corners; posterior margin distinctly bisinuate; lateral carinae narrow, fused with lateral margins; antebasal pits small and shallow, each located much closer to posterior than to lateral pronotal margin. Punctures on disc much smaller and shallower than those on head, but very dense; setae dense, short, suberect.
</paragraph>
<paragraph id="8BF836E1FF89DE43FF75D3EDFCEDAE96" blockId="42.[151,1437,151,1616]" pageId="42" pageNumber="43">
Elytra short, oval, broadest distinctly anterior to middle, EL
<quantity id="4CBF9B04FF89DE43FCDED3EDFBA2AE4A" box="[876,1039,764,788]" metricMagnitude="-4" metricUnit="m" metricValue="7.1499999999999995" metricValueMax="7.3" metricValueMin="7.0" pageId="42" pageNumber="43" unit="mm" value="0.715" valueMax="0.73" valueMin="0.7">0.700.73 mm</quantity>
(mean
<quantity id="4CBF9B04FF89DE43FBD0D3EDFB67AE4A" box="[1122,1226,764,788]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="42" pageNumber="43" unit="mm" value="0.71">0.71 mm</quantity>
), EW
<quantity id="4CBF9B04FF89DE43FAA2D3EDFADBAE4A" box="[1296,1398,764,789]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="42" pageNumber="43" unit="mm" value="0.63">0.63 mm</quantity>
, EI 1.121.16. Humeral calli weakly elevated but distinct; subhumeral lines moderately distinct, equal to about 0.3 EL, each developed as moderately sharply marked stepwise border between more convex humeral region and less convex adsutural area; basal pit on each elytron large and connected to short, elongate impression extending posteromesally; apices of elytra separately rounded. Punctures very small, sharply marked, dense, slightly more distinct than those on pronotum; setae short, dense, suberect.
</paragraph>
<paragraph id="8BF836E1FF89DE43FF75D2C5FE7EAEB3" blockId="42.[151,1437,151,1616]" box="[199,467,980,1005]" pageId="42" pageNumber="43">Hind wings not studied.</paragraph>
<paragraph id="8BF836E1FF89DE43FF75D2E6FEF2A96B" blockId="42.[151,1437,151,1616]" pageId="42" pageNumber="43">Metaventrite with indistinctly delimited postmesocoxal impressions, remaining surface covered with very fine punctures, glossy.</paragraph>
<paragraph id="8BF836E1FF89DE43FF75D52EFB17A9FE" blockId="42.[151,1437,151,1616]" pageId="42" pageNumber="43">
Hind legs with strongly modified tibiae (
<figureCitation id="137C2A64FF89DE43FD2AD52EFC94A906" box="[664,825,1087,1112]" captionStart="FIGURES 102 111" captionStartId="41.[151,264,1919,1942]" captionTargetBox="[151,1436,423,1895]" captionTargetId="figure@41.[151,1436,401,1908]" captionTargetPageId="41" captionText="FIGURES 102 111. Right metatibia of male in dorsal view. Cephennodes (s. str.) dolakhanus s sp. n. (102, 104), C. (s. str.) manangensis sp. n. (103, 105), C. (s. str.) martensi sp. n. (106), C. (s. str.) paramartensi sp. n. (107), C. (s. str.) monolaminatus sp. n. (108), C. (s. str.) thakanus sp. n. (109), C. (s. str.) annapurnaensis sp. n. (110), and C. (s. str.) parbatensis sp. n. (111). 104 111 not to the same scale." httpUri="https://zenodo.org/record/1049226/files/figure.png" pageId="42" pageNumber="43">Figs 103, 105</figureCitation>
); in dorsal view lateral (external) tibial margin nearly straight in proximal half and evenly convex in distal half, spatulate portion of tibia gradually narrowed distally; dorsodistal porous field much longer than broad; distal fringe of setae lacking one long bristle.
</paragraph>
<paragraph id="8BF836E1FF89DE43FF75D5BDFCBFA9B6" blockId="42.[151,1437,151,1616]" pageId="42" pageNumber="43">
Abdomen (
<figureCitation id="137C2A64FF89DE43FEF6D5BDFE73A99B" box="[324,478,1196,1221]" captionStart="FIGURES 112 127" captionStartId="45.[151,264,1763,1786]" captionTargetBox="[158,1435,194,1728]" captionTargetId="figure@45.[151,1436,193,1742]" captionTargetPageId="45" captionText="FIGURES 112 127. Abdomen of male in right lateral (112, 116, 120, 124) and ventral (113, 117, 121, 125) views and aedeagus in ventral (114, 118, 122, 126) and lateral (115, 119, 123, 127) views. Cephennodes (s. str.) dolakhanus sp. n. (112 115), C. (s. str.) manangensis sp. n. (116 119), C. (s. str.) martensi sp. n. (120 123), C. (s. str.) paramartensi sp. n. (124 127). Abbreviations: aml, apex of median lobe; ap, apical projection." httpUri="https://zenodo.org/record/1049228/files/figure.png" pageId="42" pageNumber="43">Figs 116117</figureCitation>
) with sternites II and III distinctly projected in lateral view (
<figureCitation id="137C2A64FF89DE43FB32D5BDFB4CA99B" box="[1152,1249,1196,1221]" captionStart="FIGURES 112 127" captionStartId="45.[151,264,1763,1786]" captionTargetBox="[158,1435,194,1728]" captionTargetId="figure@45.[151,1436,193,1742]" captionTargetPageId="45" captionText="FIGURES 112 127. Abdomen of male in right lateral (112, 116, 120, 124) and ventral (113, 117, 121, 125) views and aedeagus in ventral (114, 118, 122, 126) and lateral (115, 119, 123, 127) views. Cephennodes (s. str.) dolakhanus sp. n. (112 115), C. (s. str.) manangensis sp. n. (116 119), C. (s. str.) martensi sp. n. (120 123), C. (s. str.) paramartensi sp. n. (124 127). Abbreviations: aml, apex of median lobe; ap, apical projection." httpUri="https://zenodo.org/record/1049228/files/figure.png" pageId="42" pageNumber="43">Fig. 116</figureCitation>
); in ventral view sternite III with broadly rounded median lobe (
<figureCitation id="137C2A64FF89DE43FD16D5DEFCABA9B6" box="[676,774,1231,1256]" captionStart="FIGURES 112 127" captionStartId="45.[151,264,1763,1786]" captionTargetBox="[158,1435,194,1728]" captionTargetId="figure@45.[151,1436,193,1742]" captionTargetPageId="45" captionText="FIGURES 112 127. Abdomen of male in right lateral (112, 116, 120, 124) and ventral (113, 117, 121, 125) views and aedeagus in ventral (114, 118, 122, 126) and lateral (115, 119, 123, 127) views. Cephennodes (s. str.) dolakhanus sp. n. (112 115), C. (s. str.) manangensis sp. n. (116 119), C. (s. str.) martensi sp. n. (120 123), C. (s. str.) paramartensi sp. n. (124 127). Abbreviations: aml, apex of median lobe; ap, apical projection." httpUri="https://zenodo.org/record/1049228/files/figure.png" pageId="42" pageNumber="43">Fig. 117</figureCitation>
).
</paragraph>
<paragraph id="8BF836E1FF89DE43FF75D5E5FF67A826" blockId="42.[151,1437,151,1616]" pageId="42" pageNumber="43">
Aedeagus (
<figureCitation id="137C2A64FF89DE43FEF7D5E5FE4CA852" box="[325,481,1268,1293]" captionStart="FIGURES 112 127" captionStartId="45.[151,264,1763,1786]" captionTargetBox="[158,1435,194,1728]" captionTargetId="figure@45.[151,1436,193,1742]" captionTargetPageId="45" captionText="FIGURES 112 127. Abdomen of male in right lateral (112, 116, 120, 124) and ventral (113, 117, 121, 125) views and aedeagus in ventral (114, 118, 122, 126) and lateral (115, 119, 123, 127) views. Cephennodes (s. str.) dolakhanus sp. n. (112 115), C. (s. str.) manangensis sp. n. (116 119), C. (s. str.) martensi sp. n. (120 123), C. (s. str.) paramartensi sp. n. (124 127). Abbreviations: aml, apex of median lobe; ap, apical projection." httpUri="https://zenodo.org/record/1049228/files/figure.png" pageId="42" pageNumber="43">Figs 118119</figureCitation>
) moderately stout; AeL
<quantity id="4CBF9B04FF89DE43FD4AD5E5FCF2A852" box="[760,863,1268,1292]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="42" pageNumber="43" unit="mm" value="0.18">0.18 mm</quantity>
, apical portion of median lobe evenly rounded, its longitudinal axis forming about 45°-angle with the long axis of aedeagus; apical projection with sinuate distal margin; parameres long and slender, not exceeding apex of median lobe, each bearing one apical and one subapical seta.
</paragraph>
<paragraph id="8BF836E1FF89DE43FF75D495FE33A8C3" blockId="42.[151,1437,151,1616]" box="[199,414,1412,1437]" pageId="42" pageNumber="43">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C35D656AFF89DE43FF75D4B6FD2FA89E" box="[199,642,1447,1472]" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8BF836E1FF89DE43FF75D4B6FD2FA89E" blockId="42.[151,1437,151,1616]" box="[199,642,1447,1472]" pageId="42" pageNumber="43">
<emphasis id="B933EAF3FF89DE43FF75D4B6FEF4A89E" bold="true" box="[199,345,1447,1472]" pageId="42" pageNumber="43">Distribution</emphasis>
(
<figureCitation id="137C2A64FF89DE43FEDAD4B6FE66A89E" box="[360,459,1447,1472]" captionStart="FIGURES 144 145" captionStartId="51.[151,264,1344,1367]" captionTargetBox="[151,1436,183,1326]" captionTargetId="figure@51.[151,1436,183,1334]" captionTargetPageId="51" captionText="FIGURES 144 145. Distribution of Cephennodes (s. str.) dolakhanus sp. n. (144, circle), C. (s. str.) manangensis sp. n. (144, triangle), C. (s. str.) martensi sp. n. (145, circle), C. (s. str.) paramartensi sp. n. (145, triangle)." httpUri="https://zenodo.org/record/1049232/files/figure.png" pageId="42" pageNumber="43">Fig. 144</figureCitation>
).
<collectingRegion id="4983F803FF89DE43FE6CD4B6FD9DA89E" box="[478,560,1447,1472]" country="Sri Lanka" name="Central" pageId="42" pageNumber="43">Central</collectingRegion>
<collectingCountry id="F3507671FF89DE43FD85D4B6FDD3A89E" box="[567,638,1447,1472]" name="Nepal" pageId="42" pageNumber="43">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D656AFF89DE43FF75D4DDFCDCA8BB" box="[199,881,1484,1509]" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8BF836E1FF89DE43FF75D4DDFCDCA8BB" blockId="42.[151,1437,151,1616]" box="[199,881,1484,1509]" pageId="42" pageNumber="43">
<emphasis id="B933EAF3FF89DE43FF75D4DDFEE3A8BB" bold="true" box="[199,334,1484,1509]" pageId="42" pageNumber="43">Etymology.</emphasis>
Locotypical; after the Manang District in
<collectingCountry id="F3507671FF89DE43FC95D4DDFCC3A8BB" box="[807,878,1484,1509]" name="Nepal" pageId="42" pageNumber="43">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D656AFF89DE43FF75D4FEFAA9AB0E" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BF836E1FF89DE43FF75D4FEFAA9AB0E" blockId="42.[151,1437,151,1616]" pageId="42" pageNumber="43">
<emphasis id="B933EAF3FF89DE43FF75D4FEFE96AB56" bold="true" box="[199,315,1519,1544]" pageId="42" pageNumber="43">Remarks.</emphasis>
This species can be identified on the basis of its small and slender body, the strongly elongate porous field on metatibia, the gradually narrowed spatulate portion of metatibia, and the moderately slender aedeagus with rounded apex of median lobe strongly curved outward from the long axis of aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>