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<document id="F06205E5139DEC9EAE385C544D2A4C9E" ID-CLB-Dataset="46041" ID-DOI="10.5281/zenodo.279058" ID-GBIF-Dataset="aadbfc66-59f3-4ecb-af49-5788c22fa7e0" ID-ISSN="1175-5326" ID-Zenodo-Dep="279058" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460407254231" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="03EE87BA5A37FFE63C9078ACFA0AFAE2" docLanguage="en" docName="zt03093p034.pdf" docOrigin="Zootaxa 3093" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ochthebius" docType="key" docVersion="9" lastPageNumber="4" masterDocId="FFD7FFC25A35FFE53C077B5AFF96FFE5" masterDocTitle="New species and new collection records of aquatic beetles in the genus Ochthebius Leach from southern Africa (Coleoptera: Hydraenidae)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="3" updateTime="1698299945171" updateUser="ExternalLinkService">
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<mods:title id="F0760D375F24B21D5929C77DB1EBC4B0">New species and new collection records of aquatic beetles in the genus Ochthebius Leach from southern Africa (Coleoptera: Hydraenidae)</mods:title>
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<mods:date id="153114468B827F9C2DA8E6978A469ED5">2011</mods:date>
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<treatment id="03EE87BA5A37FFE63C9078ACFA0AFAE2" ID-DOI="http://doi.org/10.5281/zenodo.6194031" ID-GBIF-Taxon="119422226" ID-Zenodo-Dep="6194031" LSID="urn:lsid:plazi:treatment:03EE87BA5A37FFE63C9078ACFA0AFAE2" httpUri="http://treatment.plazi.org/id/03EE87BA5A37FFE63C9078ACFA0AFAE2" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="C35D65275A37FFE73C9078ACFC6EFBF5" box="[151,1016,1014,1040]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BF836AC5A37FFE73C9078ACFC6EFBF5" blockId="2.[151,1016,1014,1040]" box="[151,1016,1014,1040]" pageId="2" pageNumber="3">
<heading id="D0B081C05A37FFE73C9078ACFC6EFBF5" bold="true" box="[151,1016,1014,1040]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B933EABE5A37FFE73C9078ACFC6EFBF5" bold="true" box="[151,1016,1014,1040]" pageId="2" pageNumber="3">
Key to subgenera, groups and species of southern African
<taxonomicName id="4C474D2F5A37FFE73F6978ACFC6EFBF5" ID-CoL="8VRWR" box="[878,1016,1014,1040]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B933EABE5A37FFE73F6978ACFC6EFBF5" bold="true" box="[878,1016,1014,1040]" italics="true" pageId="2" pageNumber="3">Ochthebius</emphasis>
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1 Pronotum with sides of sclerotized part gradually rounded from anterior angles to beyond middle, then produced as minute point in front of excavate posterior angles; pronotal disc with distinct midlongitudinal sulcus, but lacking anterior and posterior foveae..............................................................................
<taxonomicName id="4C474D2F5A37FFE738D77F22FA1CFB69" authority="Stephens" authorityName="Stephens" box="[1232,1418,1143,1165]" class="Insecta" family="Hydraenidae" genus="Asiobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B933EABE5A37FFE738D77F22FABBFB68" box="[1232,1325,1144,1165]" italics="true" pageId="2" pageNumber="3">Asiobates</emphasis>
Stephens
</taxonomicName>
2
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A37FFE73C907FCFFA0AFB22" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907FCFFA0AFB22" pageId="2" pageNumber="3">
- Pronotum with sides of sclerotized part sinuately attenuate or gradually arcuate to basal angles; pronotal impressions various, but never with combination as stated above...................................................
<taxonomicName id="4C474D2F5A37FFE738EF7FE8FAC3FB22" ID-CoL="8VRWR" box="[1256,1365,1202,1223]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B933EABE5A37FFE738EF7FE8FAC3FB22" box="[1256,1365,1202,1223]" italics="true" pageId="2" pageNumber="3">Ochthebius</emphasis>
</taxonomicName>
(
<emphasis id="B933EABE5A37FFE739647FE8FA14FB22" box="[1379,1410,1202,1223]" italics="true" pageId="2" pageNumber="3">s.s.</emphasis>
) 3
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</paragraph>
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<keyStep id="30B32E095A37FFE73C907F95FA0DFAD8" pageId="2" pageNumber="3">
<paragraph id="8BF836AC5A37FFE73C907F95FA0DFAE7" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907F95FA0DFAE7" pageId="2" pageNumber="3">
2 Body more robust; size larger, ca.
<quantity id="4CBF9B495A37FFE73E467F95FD0AFB01" box="[577,668,1231,1252]" metricMagnitude="-3" metricUnit="m" metricValue="2.12" pageId="2" pageNumber="3" unit="mm" value="2.12">2.12 mm</quantity>
; pronotum very coarsely, deeply and densely punctate; elytra deeply striateimpressed (
<figureCitation id="137C2A295A37FFE73D487FB7FE02FAE7" box="[335,404,1261,1282]" captionStart="FIGURE 15" captionStartId="22.[151,250,1873,1896]" captionTargetBox="[232,1354,748,1818]" captionTargetId="figure@22.[223,1364,718,1852]" captionTargetPageId="22" captionText="FIGURE 15. Ochthebius bupunctus, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279068/files/figure.png" pageId="2" pageNumber="3">Fig. 15</figureCitation>
)..........................................................................
<taxonomicName id="4C474D2F5A37FFE7391B7FB7FA0DFAE7" ID-CoL="74GCS" box="[1308,1435,1261,1282]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bupunctus">
<emphasis id="B933EABE5A37FFE7391B7FB7FA0DFAE7" box="[1308,1435,1261,1282]" italics="true" pageId="2" pageNumber="3">O. bupunctus</emphasis>
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<keyLead id="30B695995A37FFE73C907E50FA0DFAD8" pageId="2" pageNumber="3">
- Body less convex, less robust; size smaller, ca.
<quantity id="4CBF9B495A37FFE73ECD7E50FCB1FAFA" box="[714,807,1290,1311]" metricMagnitude="-3" metricUnit="m" metricValue="1.98" pageId="2" pageNumber="3" unit="mm" value="1.98">1.98 mm</quantity>
; pronotum less densely punctate; elytra weakly or not striateimpressed..................................................................................
<taxonomicName id="4C474D2F5A37FFE739197E72FA0DFAD8" ID-CoL="74GCW" box="[1310,1435,1320,1341]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="andronius">
<emphasis id="B933EABE5A37FFE739197E72FA0DFAD8" box="[1310,1435,1320,1341]" italics="true" pageId="2" pageNumber="3">O. andronius</emphasis>
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<keyLead id="30B695995A37FFE73C907E1FFA0DFA71" pageId="2" pageNumber="3">
3 Head with frontoclypeal suture very weakly impressed; labrum subequal in length to clypeus, slightly inclined if at all; intercoxal ventrite and anterior part of 1st abdominal ventrite withdrawn under metaventrite, pubescent part of 1st ventrite forming part of coxal cavity; legs elongate, metafemora slender; inhabits rock pools in coastal splash zone.........
<taxonomicName id="4C474D2F5A37FFE738F07E25FADAFA71" ID-CoL="48F4Y" box="[1271,1356,1407,1428]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="capicola">Capicola</taxonomicName>
group 4
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A37FFE73C907EC7FA0AFA2A" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907EC7FA0AFA2A" pageId="2" pageNumber="3">- Head with frontoclypeal suture deeply impressed; labrum shorter than and inclined at distinct angle to clypeus; intercoxal ventrite visible; metacoxal cavities distinctly rimmed posteriorly................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A37FFE73C907E8DFA0DF9A0" pageId="2" pageNumber="3">
<paragraph id="8BF836AC5A37FFE73C907E8DFA0AF9EF" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907E8DFA0AF9EF" pageId="2" pageNumber="3">
4 Elytral punctures serial except random near suture basally, near humerus, and laterally beyond 8th series; pronotum with lateral lobes rugulose, discal interstices shining; large species (ca.
<quantity id="4CBF9B495A37FFE73F1D7EAFFCE7F9EF" box="[794,881,1525,1546]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="3" unit="mm" value="2.4">2.40 mm</quantity>
)....................................
<taxonomicName id="4C474D2F5A37FFE7392D7EAFFA0AF9EF" ID-CoL="48FBC" box="[1322,1436,1525,1546]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="rubripes">
<emphasis id="B933EABE5A37FFE7392D7EAFFA0AF9EF" box="[1322,1436,1525,1546]" italics="true" pageId="2" pageNumber="3">O. rubripes</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BF836AC5A37FFE73C907D48FA0DF9A0" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907D48FA0DF9A0" pageId="2" pageNumber="3">
- Elytral punctures completely random; pronotum uniformly finely microreticulate, dull; smaller species (ca.
<quantity id="4CBF9B495A37FFE738EE7D48FAD6F9C2" box="[1257,1344,1554,1575]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="2" pageNumber="3" unit="mm" value="1.92">1.92 mm</quantity>
)...................................................................................................
<taxonomicName id="4C474D2F5A37FFE7392D7D6AFA0DF9A0" ID-CoL="48F4Y" box="[1322,1435,1584,1605]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="capicola">
<emphasis id="B933EABE5A37FFE7392D7D6AFA0DF9A0" box="[1322,1435,1584,1605]" italics="true" pageId="2" pageNumber="3">O. capicola</emphasis>
</taxonomicName>
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</keyStep>
<keyStep id="30B32E095A37FFE73C907D17FA0AF95F" pageId="2" pageNumber="3">
<paragraph id="8BF836AC5A37FFE73C907D17FA0AF99A" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907D17FA0AF99A" pageId="2" pageNumber="3">
5 Dorsum black with distinctive white pubescence; pronotum with midlongitudinal sulcus; metaventrite uniformly clothed with very dense and very short hydrofuge pubescence.............................................
<taxonomicName id="4C474D2F5A37FFE738D37D30FADAF99A" ID-CoL="48F9P" box="[1236,1356,1642,1663]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="namibiensis">Namibiensis</taxonomicName>
group 6
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A37FFE73C907DDDFA0AF95F" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907DDDFA0AF95F" pageId="2" pageNumber="3">- Dorsum at least in part brownish or testaceous; pronotum with or without midlongitudinal sulcus; metaventrite posteromedially with setae less dense than and cuticle more shining than lateral area, sometimes strongly shining and nearly glabrous.. 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A37FFE73C907D98FA0AF88F" pageId="2" pageNumber="3">
<paragraph id="8BF836AC5A37FFE73C907D98FA0AF8F7" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907D98FA0AF8F7" pageId="2" pageNumber="3">6 Elytron without &quot;accessory series&quot; between 1st and 2nd series and between 9th and 10th series; explanate margin of elytron distinctly wider in front of than behind middle (in female about as wide as mesotibia); pronotal posterior foveae well-developed, microreticulate within............................................................................. 7</keyLead>
</paragraph>
<paragraph id="8BF836AC5A37FFE73C907C40FA0AF88F" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907C40FA0AF88F" pageId="2" pageNumber="3">- Elytron with &quot;accessory series&quot; between 1st and 2nd series (in basal 1/3) and another between 9th and 10th series (punctures sometimes subserially arranged); explanate margin of elytron very narrow in both sexes, widest at midlength; pronotal posterior foveae varying from obsolete to well-developed.......................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A37FFE73C907C28FA0AF805" pageId="2" pageNumber="3">
<paragraph id="8BF836AC5A37FFE73C907C28FA0DF840" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907C28FA0DF840" pageId="2" pageNumber="3">
7 Size larger (ca.
<quantity id="4CBF9B495A37FFE73D717C28FE59F862" box="[374,463,1906,1927]" metricMagnitude="-3" metricUnit="m" metricValue="1.87" pageId="2" pageNumber="3" unit="mm" value="1.87">1.87 mm</quantity>
); sides of elytra more rounded; pronotal anterior foveae more widely separated from posterior foveae, together circumscribing an elongate rectangle....................................................
<taxonomicName id="4C474D2F5A37FFE7390B7CCAFA0DF840" ID-CoL="48F9P" box="[1292,1435,1936,1957]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="B933EABE5A37FFE7390B7CCAFA0DF840" box="[1292,1435,1936,1957]" italics="true" pageId="2" pageNumber="3">O. namibiensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A37FFE73C907CF7FA0AF805" blockId="2.[151,1436,1085,2016]" pageId="2" pageNumber="3">
<keyLead id="30B695995A37FFE73C907CF7FA0AF805" pageId="2" pageNumber="3">
- Size smaller (ca.
<quantity id="4CBF9B495A37FFE73D807CF7FE76F827" box="[391,480,1965,1986]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" pageId="2" pageNumber="3" unit="mm" value="1.71">1.71 mm</quantity>
); sides of elytra less rounded; pronotal anterior foveae less widely separated from posterior foveae, together circumscribing a square (
<figureCitation id="137C2A295A37FFE73E2F7C90FDFDF83A" box="[552,619,1994,2015]" captionStart="FIGURE 11" captionStartId="17.[151,250,2000,2023]" captionTargetBox="[226,1357,661,1979]" captionTargetId="figure@17.[226,1360,661,1979]" captionTargetPageId="17" captionText="FIGURE 11. Ochthebius zulu, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279065/files/figure.png" pageId="2" pageNumber="3">Fig. 11</figureCitation>
).............................................................
<taxonomicName id="4C474D2F5A37FFE739517C91FA0AF805" ID-CoL="48FDJ" box="[1366,1436,1995,2016]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="zulu">
<emphasis id="B933EABE5A37FFE739517C91FA0AF805" box="[1366,1436,1995,2016]" italics="true" pageId="2" pageNumber="3">O. zulu</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A36FFE63C907BCDFA0DFEE0" pageId="3" pageNumber="4">
<paragraph id="8BF836AC5A36FFE63C907BCDFA0AFF2F" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907BCDFA0AFF2F" pageId="3" pageNumber="4">
8 Pronotal sides concavely attenuate from near anterior angles, hyaline border very wide, nearly as wide as subdigitiform, sharply rounded lateral lobe; pronotal posterior foveae well-developed...................................
<taxonomicName id="4C474D2F5A36FFE639297BEFFA0AFF2F" ID-CoL="6SFVL" box="[1326,1436,181,202]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="spatulus">
<emphasis id="B933EABE5A36FFE639297BEFFA0AFF2F" box="[1326,1436,181,202]" italics="true" pageId="3" pageNumber="4">O. spatulus</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BF836AC5A36FFE63C907B88FA0DFEE0" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907B88FA0DFEE0" pageId="3" pageNumber="4">
- Pronotal sides sinuately attenuate, concave from about middle to posterior angles, hyaline border much narrower than widely rounded lateral lobe; pronotal posterior foveae obsolete or weakly developed...........................
<taxonomicName id="4C474D2F5A36FFE6390B7BAAFA0DFEE0" ID-CoL="6SFTR" box="[1292,1435,240,261]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pagotrichus">
<emphasis id="B933EABE5A36FFE6390B7BAAFA0DFEE0" box="[1292,1435,240,261]" italics="true" pageId="3" pageNumber="4">O. pagotrichus</emphasis>
</taxonomicName>
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</keyStep>
<keyStep id="30B32E095A36FFE63C907A57FA0AFE9F" pageId="3" pageNumber="4">
<paragraph id="8BF836AC5A36FFE63C907A57FA0AFEB8" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907A57FA0AFEB8" pageId="3" pageNumber="4">
9 Elytral intervals 2, 3, 5, and 7 each with unilinear row of erect short fine setae; labrum in both sexes bearing non-spiniform setae, anterior margin apicomedially emarginate; pronotal foveae confluent medially with shallow midlongitudinal impression (location marked by microreticulation); (
<taxonomicName id="4C474D2F5A36FFE63E6F7A1DFD53FEB9" ID-CoL="48F65" box="[616,709,327,348]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="extremus">Extremus</taxonomicName>
group)............................................
<taxonomicName id="4C474D2F5A36FFE6392E7A12FA0AFEB8" ID-CoL="48F65" box="[1321,1436,328,349]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="extremus">
<emphasis id="B933EABE5A36FFE6392E7A12FA0AFEB8" box="[1321,1436,328,349]" italics="true" pageId="3" pageNumber="4">O. extremus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A36FFE63C907A3FFA0AFE9F" blockId="3.[151,1437,151,1287]" box="[151,1436,357,378]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907A3FFA0AFE9F" box="[151,1436,357,378]" pageId="3" pageNumber="4">- Elytral intervals without rows of erect setae; labrum various; pronotal impression various........................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A36FFE63C907AD8FA0DFE0A" pageId="3" pageNumber="4">
<paragraph id="8BF836AC5A36FFE63C907AD8FA0AFE50" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907AD8FA0AFE50" pageId="3" pageNumber="4">
10 Elytral punctures random; body form compressed; pronotal impressions very shallow; male labrum not strongly upturned, without spiniform setae; flightless species;
<taxonomicName id="4C474D2F5A36FFE63E877AC5FD4FFE51" ID-CoL="48F7M" box="[640,729,415,436]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="involatus">Involatus</taxonomicName>
group (
<figureCitation id="137C2A295A36FFE63F237AC5FCFFFE51" box="[804,873,415,436]" captionStart="FIGURE 13" captionStartId="19.[151,250,1865,1888]" captionTargetBox="[246,1315,434,1809]" captionTargetId="figure@19.[229,1357,407,1843]" captionTargetPageId="19" captionText="FIGURE 13. Ochthebius involatus, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279067/files/figure.png" pageId="3" pageNumber="4">Fig. 13</figureCitation>
)...................................
<taxonomicName id="4C474D2F5A36FFE639217AFAFA0AFE50" ID-CoL="48F7M" box="[1318,1436,416,437]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="involatus">
<emphasis id="B933EABE5A36FFE639217AFAFA0AFE50" box="[1318,1436,416,437]" italics="true" pageId="3" pageNumber="4">O. involatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A36FFE63C907AE7FA0DFE0A" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907AE7FA0DFE0A" pageId="3" pageNumber="4">
- Elytral punctures serial; labrum in male with anterior margin apicomedially reflexed, usually with short spiniform setae forming group on each side of middle,; pronotal impression various..................................
<taxonomicName id="4C474D2F5A36FFE638E67A80FAD7FE0A" ID-CoL="48FBF" box="[1249,1345,474,495]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="salinarius">Salinarius</taxonomicName>
group 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A36FFE63C907AADFA0AFD67" pageId="3" pageNumber="4">
<paragraph id="8BF836AC5A36FFE63C907AADFA0DFDCF" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907AADFA0DFDCF" pageId="3" pageNumber="4">11 Pronotum with midlongitudinal sulcus well-developed, even in middle; anterior and posterior foveae separate from or only very shallowly joined to sulcus; elytron testaceous with partial or complete brownish longitudinal lines through each series 12</keyLead>
</paragraph>
<paragraph id="8BF836AC5A36FFE63C907968FA0AFD67" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907968FA0AFD67" pageId="3" pageNumber="4">- Pronotum with midlongitudinal impression restricted to anterior and posterior, obsolete if at all present over median area; anterior and posterior foveae deeply confluent with respective parts of midlongitudinal impression; elytron without longitudinal lines of color, often with darker coloration at summit of posterior declivity....................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A36FFE63C9079D0FA0AFD3F" pageId="3" pageNumber="4">
<paragraph id="8BF836AC5A36FFE63C9079D0FA0AFD7A" blockId="3.[151,1437,151,1287]" box="[151,1436,650,671]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C9079D0FA0AFD7A" box="[151,1436,650,671]" pageId="3" pageNumber="4">
12 Elytra with complete, well formed longitudinal lines of color through each series.........................
<taxonomicName id="4C474D2F5A36FFE6391979D0FA0AFD7A" ID-CoL="48FBF" box="[1310,1436,650,671]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="salinarius">
<emphasis id="B933EABE5A36FFE6391979D0FA0AFD7A" box="[1310,1436,650,671]" italics="true" pageId="3" pageNumber="4">O. salinarius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A36FFE63C9079FDFA0AFD3F" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C9079FDFA0AFD3F" pageId="3" pageNumber="4">
- Each elytral puncture with a dark brown dot, some of which are contiguous, but not forming parallel-side dark lines (
<figureCitation id="137C2A295A36FFE6394479FDFAECFD59" box="[1347,1402,679,700]" captionStart="FIGURE 1" captionStartId="4.[151,250,1961,1984]" captionTargetBox="[256,1334,645,1923]" captionTargetId="figure@4.[226,1360,598,1935]" captionTargetPageId="4" captionText="FIGURE 1. Ochthebius endroedyi, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279059/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
)............................................................................................
<taxonomicName id="4C474D2F5A36FFE63918799FFA0AFD3F" ID-CoL="48F5Z" box="[1311,1436,709,730]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="endroedyi">
<emphasis id="B933EABE5A36FFE63918799FFA0AFD3F" box="[1311,1436,709,730]" italics="true" pageId="3" pageNumber="4">O. endroedyi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A36FFE63C9079B8FA0AFC89" pageId="3" pageNumber="4">
<paragraph id="8BF836AC5A36FFE63C9079B8FA0DFCD7" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C9079B8FA0DFCD7" pageId="3" pageNumber="4">
13 Pronotal reliefs finely sparsely punctate; metaventrite with small basomedian area less densely pubescent and more shining than remainder, which is microreticulate beneath hydrofuge pubescence; female with narrow median glabrous weakly shining area on ventrites 24 and anterior part of 5; size larger (ca.
<quantity id="4CBF9B495A36FFE63F1F7847FC33FCD7" box="[792,933,797,818]" metricMagnitude="-3" metricUnit="m" metricValue="1.975" metricValueMax="2.1" metricValueMin="1.85" pageId="3" pageNumber="4" unit="mm" value="1.975" valueMax="2.1" valueMin="1.85">1.852.10 mm</quantity>
).................................
<taxonomicName id="4C474D2F5A36FFE639317847FA0DFCD7" ID-CoL="6SG62" box="[1334,1435,797,818]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pedalis">
<emphasis id="B933EABE5A36FFE639317847FA0DFCD7" box="[1334,1435,797,818]" italics="true" pageId="3" pageNumber="4">O. pedalis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A36FFE63C907860FA0AFC89" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907860FA0AFC89" pageId="3" pageNumber="4">
- Pronotal reliefs with coarse, irregularly spaced punctures, some confluent; metaventrite pubescent laterally, disc strongly shining, glabrous except few setiferous punctures; smaller species (ca.
<quantity id="4CBF9B495A36FFE63F67780DFC7BFC89" box="[864,1005,855,876]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="3" pageNumber="4" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.601.80 mm</quantity>
)................................. 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A36FFE63C90782FFA0DFC07" pageId="3" pageNumber="4">
<paragraph id="8BF836AC5A36FFE63C90782FFA0AFC42" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C90782FFA0AFC42" pageId="3" pageNumber="4">
14 Color generally dark brown, elytra uniformly colored, without darker dot around each serial puncture; elytra with a granule at the anterior margin of each serial puncture (
<figureCitation id="137C2A295A36FFE63E8278C8FD28FC42" box="[645,702,914,935]" captionStart="FIGURE 7" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[241,1346,438,1834]" captionTargetId="figure@12.[228,1358,409,1845]" captionTargetPageId="12" captionText="FIGURE 7. Ochthebius granulinus, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279063/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
).................................................
<taxonomicName id="4C474D2F5A36FFE6391178C8FA0AFC42" ID-CoL="48F6R" box="[1302,1436,914,935]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="granulinus">
<emphasis id="B933EABE5A36FFE6391178C8FA0AFC42" box="[1302,1436,914,935]" italics="true" pageId="3" pageNumber="4">O. granulinus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A36FFE63C9078F5FA0DFC07" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C9078F5FA0DFC07" pageId="3" pageNumber="4">- Elytra lighter brown, with darker color around each serial puncture, caused by the underlying columellae; elytra without granules............................................................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A36FFE63C9078B0FA0AFBB2" pageId="3" pageNumber="4">
<paragraph id="8BF836AC5A36FFE63C9078B0FA0DFBF9" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C9078B0FA0DFBF9" pageId="3" pageNumber="4">15 Pronotal posterior foveae deeper laterally than medially; females with hydrofuge pubescence very sparse in midline of ventrites 35........................................................................................... 16</keyLead>
</paragraph>
<paragraph id="8BF836AC5A36FFE63C907F7FFA0AFBB2" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907F7FFA0AFBB2" pageId="3" pageNumber="4">- Pronotal posterior foveae markedly confluent, deeper medially than laterally; females with hydrofuge pubescence not significantly sparser in midline of ventrites 35.................................................................. 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A36FFE63C907F05FA0AFB28" pageId="3" pageNumber="4">
<paragraph id="8BF836AC5A36FFE63C907F05FA0AFB77" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907F05FA0AFB77" pageId="3" pageNumber="4">
16 Pronotal posterior foveae deeper and with more abrupt lateral margins; body less convex; elytra with less rounded sides and less acuminate apices; size larger (ca.
<quantity id="4CBF9B495A36FFE63E497F27FD31FB77" box="[590,679,1149,1170]" metricMagnitude="-3" metricUnit="m" metricValue="1.74" pageId="3" pageNumber="4" unit="mm" value="1.74">1.74 mm</quantity>
); aedeagus as illustrated (
<figureCitation id="137C2A295A36FFE63F9B7F27FC76FB77" box="[924,992,1149,1170]" captionStart="FIGURE 5" captionStartId="10.[151,250,1671,1694]" captionTargetBox="[246,1352,337,1626]" captionTargetId="figure@10.[226,1360,305,1650]" captionTargetPageId="10" captionText="FIGURE 5. Ochthebius anchorus, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279062/files/figure.png" pageId="3" pageNumber="4">Figs. 5</figureCitation>
,
<figureCitation id="137C2A295A36FFE63FEC7F27FC61FB77" box="[1003,1015,1149,1170]" captionStart="FIGURE 6" captionStartId="11.[151,250,1650,1673]" captionText="FIGURE 6. Ochthebius anchorus, aedeagus of holotype." pageId="3" pageNumber="4">6</figureCitation>
)........................
<taxonomicName id="4C474D2F5A36FFE639227F27FA0AFB77" ID-CoL="48F3R" box="[1317,1436,1149,1170]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="anchorus">
<emphasis id="B933EABE5A36FFE639227F27FA0AFB77" box="[1317,1436,1149,1170]" italics="true" pageId="3" pageNumber="4">O. anchorus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A36FFE63C907FC0FA0AFB28" blockId="3.[151,1437,151,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907FC0FA0AFB28" pageId="3" pageNumber="4">
- Pronotal posterior foveae shallower and with less abrupt lateral margins; body more convex; elytra with more rounded sides and more acuminate apices; size smaller (ca.
<quantity id="4CBF9B495A36FFE63E947FEDFD7AFB29" box="[659,748,1207,1228]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="3" pageNumber="4" unit="mm" value="1.66">1.66 mm</quantity>
);...............................................
<taxonomicName id="4C474D2F5A36FFE6392D7FE2FA0AFB28" ID-CoL="48FCC" box="[1322,1436,1208,1229]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="spinasus">
<emphasis id="B933EABE5A36FFE6392D7FE2FA0AFB28" box="[1322,1436,1208,1229]" italics="true" pageId="3" pageNumber="4">O. spinasus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B32E095A36FFE63C907F8FFA0AFAE2" pageId="3" pageNumber="4">
<paragraph id="8BF836AC5A36FFE63C907F8FFA0DFB0F" blockId="3.[151,1437,151,1287]" box="[151,1435,1237,1258]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907F8FFA0DFB0F" box="[151,1435,1237,1258]" pageId="3" pageNumber="4">
17 Metaventral glabrous area triangular; size larger (ca.
<quantity id="4CBF9B495A36FFE63EDA7F8FFCA0FB0F" box="[733,822,1237,1258]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" pageId="3" pageNumber="4" unit="mm" value="1.77">1.77 mm</quantity>
); aedeagus as illustrated (
<figureCitation id="137C2A295A36FFE6382B7F8FFBF9FB0F" box="[1068,1135,1237,1258]" captionStart="FIGURE 3" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[242,1328,493,1855]" captionTargetId="figure@7.[232,1355,448,1884]" captionTargetPageId="7" captionText="FIGURE 3. Ochthebius sitiensis, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279061/files/figure.png" pageId="3" pageNumber="4">Figs. 3</figureCitation>
,
<figureCitation id="137C2A295A36FFE6387D7F8FFB11FB0F" box="[1146,1159,1237,1258]" captionStart="FIGURE 4" captionStartId="8.[151,250,1624,1647]" captionText="FIGURE 4. Ochthebius sitiensis, aedeagus of holotype." pageId="3" pageNumber="4">4</figureCitation>
).............
<taxonomicName id="4C474D2F5A36FFE639367F8FFA0DFB0F" ID-CoL="48FC7" box="[1329,1435,1237,1258]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sitiensis">
<emphasis id="B933EABE5A36FFE639367F8FFA0DFB0F" box="[1329,1435,1237,1258]" italics="true" pageId="3" pageNumber="4">O. sitiensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF836AC5A36FFE63C907FA8FA0AFAE2" blockId="3.[151,1437,151,1287]" box="[151,1436,1266,1287]" pageId="3" pageNumber="4">
<keyLead id="30B695995A36FFE63C907FA8FA0AFAE2" box="[151,1436,1266,1287]" pageId="3" pageNumber="4">
- Metaventral glabrous area subrhomboidal; size smaller (ca.
<quantity id="4CBF9B495A36FFE63F197FA8FCE1FAE2" box="[798,887,1266,1287]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="3" pageNumber="4" unit="mm" value="1.67">1.67 mm</quantity>
); aedeagus as illustrated (
<figureCitation id="137C2A295A36FFE6386A7FA8FB27FAE2" box="[1133,1201,1266,1287]" captionStart="FIGURE 9" captionStartId="15.[151,250,1977,2000]" captionTargetBox="[263,1343,534,1927]" captionTargetId="figure@15.[233,1353,519,1955]" captionTargetPageId="15" captionText="FIGURE 9. Ochthebius bicomicus, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279064/files/figure.png" pageId="3" pageNumber="4">Figs. 9</figureCitation>
,
<figureCitation id="137C2A295A36FFE638BC7FA8FB42FAE2" box="[1211,1236,1266,1287]" captionStart="FIGURE 10" captionStartId="16.[151,250,1605,1628]" captionText="FIGURE 10. Ochthebius bicomicus, aedeagus of holotype." pageId="3" pageNumber="4">10</figureCitation>
).....
<taxonomicName id="4C474D2F5A36FFE6391A7FA8FA0AFAE2" ID-CoL="48F4M" box="[1309,1436,1266,1287]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bicomicus">
<emphasis id="B933EABE5A36FFE6391A7FA8FA0AFAE2" box="[1309,1436,1266,1287]" italics="true" pageId="3" pageNumber="4">O. bicomicus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>