222 lines
30 KiB
XML
222 lines
30 KiB
XML
<document id="A9A33D1A1FB73A563C54D39578CF8409" ID-DOI="10.21248/contrib.entomol.69.1.033-070" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742766" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638269617685" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2019" docId="03EE87A5FFFA3B7CFCBFFD5472179A7E" docLanguage="en" docName="ContribEntomol.69.1.33-70.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 69 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Drusilla penicillata Assing 2019, spec. nov." docType="treatment" docUuid="CAA17FAD-AE2A-496C-9D08-84D34358BDD5" docUuidSource="ZooBank" docVersion="3" lastPageNumber="60" masterDocId="FFD7FFDDFFE03B67FFC3FFDE77509866" masterDocTitle="On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="70" masterPageNumber="33" pageNumber="59" updateTime="1699437235520" updateUser="plazi">
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<mods:mods id="784521D6AEE22F454055024F2F44149A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0EA376AC1C8D7E16743ACE4E5E4918E4">
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<mods:title id="5EA5E21278C578A6E79B8487C27E2786">On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="5962A378B1AD38A4422271E2892A937A" type="personal">
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<mods:roleTerm id="A7CF151DB789EFA3476592B06685A66C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B48D1E0F0176A4DF64A71F510AD8E766">Assing, Volker</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F029A58A3F2399F6EC522B4576536A4C">text</mods:typeOfResource>
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<mods:title id="B977DEBAE304D613B02DA9F755EA52C1">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="4463AC20FB3F924886CA966FB4702EA7">
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<mods:date id="A885EEF7574A459BDD4FD31ACA5B1601">2019</mods:date>
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<mods:detail id="1717B30CE11CBAB111FEE6E14A6302D3" type="pubDate">
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<mods:number id="A2BBCBA084F6FF05738668370928CBED">2019-06-24</mods:number>
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</mods:detail>
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<mods:detail id="1A6A592D7AA49957A1DB2ABC57C187E7" type="volume">
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<mods:number id="87D12DCFD3E70A09D2B1D9BA34D8466A">69</mods:number>
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</mods:detail>
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<mods:detail id="FC16AC1587DB05F912C53EFBF738ECD8" type="issue">
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<mods:number id="F4549D6A9B65BF63DF482AB3C114DA8F">1</mods:number>
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</mods:detail>
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<mods:start id="2BEEB0767270E918C5C4E26774F0FDF3">33</mods:start>
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<mods:end id="5723BA4E5F4E7F227C04D94BD58C873C">70</mods:end>
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</mods:extent>
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<mods:classification id="7C68003B6348BEAE9CE3BCEAC0AC0B28">journal article</mods:classification>
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<mods:identifier id="507318B9439B096D0B5C8A0100D7E42C" type="DOI">10.21248/contrib.entomol.69.1.033-070</mods:identifier>
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<mods:identifier id="8A38F13D5B3FA25B67D307DC4FFE626E" type="ISSN">0005-805X</mods:identifier>
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<mods:identifier id="ED6B5151559AE0D75E5CA51385E94E99" type="Zenodo-Dep">5742766</mods:identifier>
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<mods:identifier id="37C9AB086FCEC4243943D83001E05B9D" type="ZooBank">B1F197EC-DB76-4BCC-8DBF-856436A81F9F</mods:identifier>
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</mods:mods>
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<treatment id="03EE87A5FFFA3B7CFCBFFD5472179A7E" LSID="urn:lsid:zoobank.org:act:CAA17FAD-AE2A-496C-9D08-84D34358BDD5" httpUri="http://treatment.plazi.org/id/03EE87A5FFFA3B7CFCBFFD5472179A7E" lastPageId="27" lastPageNumber="60" pageId="26" pageNumber="59">
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<subSubSection id="C35D6538FFFA3B7DFCBFFD5472C09AA3" pageId="26" pageNumber="59" type="nomenclature">
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<paragraph id="8BF836B3FFFA3B7DFCBFFD5473F99AC0" blockId="26.[892,1424,650,741]" box="[892,1193,650,678]" pageId="26" pageNumber="59">
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<heading id="D0B081DFFFFA3B7DFCBFFD5473F99AC0" box="[892,1193,650,678]" fontSize="12" level="2" pageId="26" pageNumber="59" reason="2">
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<taxonomicName id="4C474D30FFFA3B7DFCBFFD54736A9AC0" ID-CoL="8XZV8" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[892,1082,650,678]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="59" phylum="Arthropoda" rank="species" species="penicillata" status="sp. nov.">
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<emphasis id="B933EAA1FFFA3B7DFCBFFD54736A9AC0" box="[892,1082,650,678]" italics="true" pageId="26" pageNumber="59">Drusilla penicillata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20057DAFFFA3B7DFB83FD5473F99AC0" box="[1088,1193,650,678]" pageId="26" pageNumber="59" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BF836B3FFFA3B7DFCBFFD6F72C09AA3" blockId="26.[892,1424,650,741]" box="[892,1424,689,709]" pageId="26" pageNumber="59">
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<uri id="FFD63AB1FFFA3B7DFCBFFD6F72C09AA3" box="[892,1424,689,709]" pageId="26" pageNumber="59">
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urn:lsid:zoobank.org:act:
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<uuid id="FFE10C66FFFA3B7DFBE9FD6F72C09AA3" box="[1066,1424,689,709]" pageId="26" pageNumber="59">CAA17FAD-AE2A-496C-9D08-84D34358BDD5</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6538FFFA3B7DFCBFFD0F73759A83" box="[892,1061,721,741]" pageId="26" pageNumber="59" type="description">
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<paragraph id="8BF836B3FFFA3B7DFCBFFD0F73759A83" blockId="26.[892,1424,650,741]" box="[892,1061,721,741]" pageId="26" pageNumber="59">
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(
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<figureCitation id="137C2A36FFFA3B7DFC43FD0F74879A83" box="[896,983,721,741]" captionStart="Figs 35–52" captionStartId="7.[174,214,2006,2028]" captionTargetBox="[174,1421,255,1986]" captionTargetId="figure-119@7.[173,1421,254,1994]" captionTargetPageId="7" captionText="Figs 35–52: Drusilla bispinosa (35–43), D. shavrini (44–47), and D. penicillata (48–52): male forebody (35, 45, 49); antenna (36, 46, 51); male abdomen (37, 47, 52); anterior portion of male abdomen (38–39); posterior portion of male abdomen in dorsal and in lateral view (40–42); posterior portion of female abdomen (43); habitus (44, 48); head in dorso-lateral view (50). Scale bars: 35–49, 51–52: 1.0 mm; 50: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742772" httpUri="https://zenodo.org/record/5742772/files/figure.png" pageId="26" pageNumber="59">Figs 48–52</figureCitation>
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, 140–144)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6538FFFA3B7DFC95FCDE73899C7E" pageId="26" pageNumber="59" type="materials_examined">
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<paragraph id="8BF836B3FFFA3B7DFC95FCDE73899C7E" blockId="26.[854,1465,767,1048]" pageId="26" pageNumber="59">
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<emphasis id="B933EAA1FFFA3B7DFC95FCDE74BF9B71" bold="true" box="[854,1007,767,792]" pageId="26" pageNumber="59">Type material</emphasis>
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:
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<materialsCitation id="3B2F3CEEFFFA3B7DFC3DFCDE73089B1E" collectingDate="2018-03-27" collectorName="Marilog D. & Shavrin" country="Philippines" county="Epol Falls" latitude="7.45" location="Mindanao" longLatPrecision="1303" longitude="125.23333" municipality="Barangay Baganihan" pageId="26" pageNumber="59" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54FC8811FFFA3B7DFC3DFCDE73379B7E" box="[1022,1127,768,792]" pageId="26" pageNumber="59" type="holotype">Holotype</typeStatus>
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: “
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<collectingCountry id="F3507623FFFA3B7DFB56FCDE72639B7E" box="[1173,1331,768,792]" name="Philippines" pageId="26" pageNumber="59">PHILIPPINES</collectingCountry>
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:
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<location id="8E986068FFFA3B7DFAFCFCDE72E59B7E" LSID="urn:lsid:plazi:treatment:03EE87A5FFFA3B7CFCBFFD5472179A7E:8E986068FFFA3B7DFAFCFCDE72E59B7E" box="[1343,1461,768,792]" country="Philippines" county="Epol Falls" latitude="7.45" longLatPrecision="1303" longitude="125.23333" municipality="Barangay Baganihan" name="Mindanao" pageId="26" pageNumber="59">Mindanao</location>
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,
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<collectingMunicipality id="6B9CACC9FFFA3B7DFC95FCFE736C9B5E" box="[854,1084,800,824]" pageId="26" pageNumber="59">Barangay Baganihan</collectingMunicipality>
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,
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<collectorName id="26B25365FFFA3B7DFB8DFCFE739D9B51" box="[1102,1229,800,824]" pageId="26" pageNumber="59">Marilog D.</collectorName>
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,
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<collectingCounty id="62994E3FFFFA3B7DFB23FCFE72029B5E" box="[1248,1362,800,824]" pageId="26" pageNumber="59">Epol Falls</collectingCounty>
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,
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<geoCoordinate id="EE735074FFFA3B7DFAA6FCFE72E59B5E" box="[1381,1461,800,824]" degrees="7" direction="north" minutes="27" orientation="latitude" pageId="26" pageNumber="59" precision="925" value="7.45">7°27'N</geoCoordinate>
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,
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<geoCoordinate id="EE735074FFFA3B7DFC95FC9E74E59B3E" box="[854,949,832,856]" degrees="125" direction="east" minutes="14" orientation="longitude" pageId="26" pageNumber="59" precision="925" value="125.23333">125°14'E</geoCoordinate>
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, wet litter and debris near waterfall sift.,
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<date id="FFF91073FFFA3B7DFC95FCBE749D9B1E" box="[854,973,864,888]" pageId="26" pageNumber="59" value="2018-03-27">
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<collectingDate id="EFBDE99BFFFA3B7DFC95FCBE749D9B1E" box="[854,973,864,888]" pageId="26" pageNumber="59" value="2018-03-27">27.III.2018</collectingDate>
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</date>
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, leg.
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<collectorName id="26B25365FFFA3B7DFBC5FCBE73089B1E" box="[1030,1112,864,888]" pageId="26" pageNumber="59">Shavrin</collectorName>
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</materialsCitation>
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/
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<materialsCitation id="3B2F3CEEFFFA3B7DFBB0FCBE73859C7E" collectingDate="2018-03-26" collectingDateMax="2018-03-28" collectingDateMin="2018-03-26" collectionCode="V" collectorName="Marilog D. & Shavrin" country="Philippines" county="Eagles Ridge" latitude="7.75" location="Assing" longLatPrecision="1303" longitude="125.38333" municipality="Barangay Baganihan" pageId="26" pageNumber="59" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54FC8811FFFA3B7DFBB0FCBE73B89B1E" box="[1139,1256,864,888]" pageId="26" pageNumber="59" type="holotype">Holotypus</typeStatus>
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<taxonomicName id="4C474D30FFFA3B7DFACBFCBE742E9BFE" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="59" phylum="Arthropoda" rank="species" species="penicillata" status="sp. nov.">
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<emphasis id="B933EAA1FFFA3B7DFACBFCBE742E9BFE" italics="true" pageId="26" pageNumber="59">Drusilla penicillata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20057DAFFFA3B7DFC4AFC5E74929BF1" box="[905,962,896,919]" pageId="26" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
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, det.
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<collectionCode id="ED56AE76FFFA3B7DFBC6FC5E73479BF1" box="[1029,1047,896,919]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="59" type="Museum">V</collectionCode>
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||
.
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<location id="8E986068FFFA3B7DFBE3FC5E73379BFE" LSID="urn:lsid:plazi:treatment:03EE87A5FFFA3B7CFCBFFD5472179A7E:8E986068FFFA3B7DFBE3FC5E73379BFE" box="[1056,1127,896,920]" country="Philippines" county="Eagles Ridge" latitude="7.75" longLatPrecision="1303" longitude="125.38333" municipality="Barangay Baganihan" name="Assing" pageId="26" pageNumber="59">Assing</location>
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2018” (cAss).
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<typeStatus id="54FC8811FFFA3B7DFAD3FC5E722A9BFE" box="[1296,1402,896,920]" pageId="26" pageNumber="59" type="paratype">Paratypes</typeStatus>
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: 1, 1: same data as holotype; 1: “
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<collectingCountry id="F3507623FFFA3B7DFB07FC7E72329BDE" box="[1220,1378,928,952]" name="Philippines" pageId="26" pageNumber="59">PHILIPPINES</collectingCountry>
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||
:
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<location id="8E986068FFFA3B7DFAADFC7E74DB9BB1" LSID="urn:lsid:plazi:treatment:03EE87A5FFFA3B7CFCBFFD5472179A7E:8E986068FFFA3B7DFAADFC7E74DB9BB1" country="Philippines" county="Eagles Ridge" latitude="7.75" longLatPrecision="1303" longitude="125.38333" municipality="Barangay Baganihan" name="Mindanao" pageId="26" pageNumber="59">Mindanao</location>
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,
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<collectingMunicipality id="6B9CACC9FFFA3B7DFC5FFC1E73D09BBE" box="[924,1152,960,984]" pageId="26" pageNumber="59">Barangay Baganihan</collectingMunicipality>
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,
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<collectorName id="26B25365FFFA3B7DFB52FC1E725D9BB1" box="[1169,1293,960,984]" pageId="26" pageNumber="59">Marilog D.</collectorName>
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,
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<collectingCounty id="62994E3FFFFA3B7DFADCFC1E72E59BBE" box="[1311,1461,960,984]" pageId="26" pageNumber="59">Eagles Ridge</collectingCounty>
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||
,
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<geoCoordinate id="EE735074FFFA3B7DFC95FC3E74F09B9E" box="[854,928,992,1016]" degrees="7" direction="north" minutes="45" orientation="latitude" pageId="26" pageNumber="59" precision="925" value="7.75">7°45'N</geoCoordinate>
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,
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<geoCoordinate id="EE735074FFFA3B7DFC76FC3E73439B9E" box="[949,1043,992,1016]" degrees="125" direction="east" minutes="23" orientation="longitude" pageId="26" pageNumber="59" precision="925" value="125.38333">125°23'E</geoCoordinate>
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, secondary forest, wet litter sifted,
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<date id="FFF91073FFFA3B7DFC95FBDE74A49C7E" box="[854,1012,1024,1048]" pageId="26" pageNumber="59" value="2018-03-26" valueMax="2018-03-28" valueMin="2018-03-26">
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||
<collectingDate id="EFBDE99BFFFA3B7DFC95FBDE74A49C7E" box="[854,1012,1024,1048]" pageId="26" pageNumber="59" value="2018-03-26" valueMax="2018-03-28" valueMin="2018-03-26">26–28.III.2018</collectingDate>
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||
</date>
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||
, leg.
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<collectorName id="26B25365FFFA3B7DFBEAFBDE732A9C7E" box="[1065,1146,1024,1048]" pageId="26" pageNumber="59">Shavrin</collectorName>
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||
” (cAss)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35D6538FFFA3B7DFC95FB9E723C9CFE" pageId="26" pageNumber="59" type="etymology">
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||
<paragraph id="8BF836B3FFFA3B7DFC95FB9E723C9CFE" blockId="26.[854,1465,1088,1176]" pageId="26" pageNumber="59">
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||
<emphasis id="B933EAA1FFFA3B7DFC95FB9E749D9C3E" bold="true" box="[854,973,1088,1112]" pageId="26" pageNumber="59">Etymology</emphasis>
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: The specific epithet is an adjective derived from the Latin noun penicillus (brush) and alludes to the presence of a setiferous tubercle on the male head.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C35D6538FFFA3B7CFC95FB6173D8991E" lastPageId="27" lastPageNumber="60" pageId="26" pageNumber="59" type="description">
|
||
<paragraph id="8BF836B3FFFA3B7DFC95FB6173DA9DBE" blockId="26.[854,1465,1215,2104]" pageId="26" pageNumber="59">
|
||
<emphasis id="B933EAA1FFFA3B7DFC95FB6174889CB1" bold="true" box="[854,984,1215,1239]" pageId="26" pageNumber="59">Description</emphasis>
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: Relatively large species: body length
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<quantity id="4CBF9B56FFFA3B7DFC95FB3E74879C91" box="[854,983,1248,1272]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.7" metricValueMin="5.8" pageId="26" pageNumber="59" unit="mm" value="6.25" valueMax="6.7" valueMin="5.8">5.8–6.7 mm</quantity>
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; length of forebody 2.8–3.0 mm. Habitus as in
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<figureCitation id="137C2A36FFFA3B7DFCB0FADE74ED9D7E" box="[883,957,1280,1304]" captionStart="Figs 35–52" captionStartId="7.[174,214,2006,2028]" captionTargetBox="[174,1421,255,1986]" captionTargetId="figure-119@7.[173,1421,254,1994]" captionTargetPageId="7" captionText="Figs 35–52: Drusilla bispinosa (35–43), D. shavrini (44–47), and D. penicillata (48–52): male forebody (35, 45, 49); antenna (36, 46, 51); male abdomen (37, 47, 52); anterior portion of male abdomen (38–39); posterior portion of male abdomen in dorsal and in lateral view (40–42); posterior portion of female abdomen (43); habitus (44, 48); head in dorso-lateral view (50). Scale bars: 35–49, 51–52: 1.0 mm; 50: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742772" httpUri="https://zenodo.org/record/5742772/files/figure.png" pageId="26" pageNumber="59">Fig. 48</figureCitation>
|
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. Coloration: body pale-reddish with the posterior portions of paratergites III–VI weakly infuscate; legs with the tarsi reddish, the bases of the meso- and metafemora and all of the meso- and metatibiae yellow, and the femora (except for the bases of the meso- and metafemora) and the protibae pale-brown; antennae and maxillary palpi pale-reddish.
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</paragraph>
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<paragraph id="8BF836B3FFFA3B7DFC95FA3E74CD9F31" blockId="26.[854,1465,1215,2104]" pageId="26" pageNumber="59">
|
||
Head (
|
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<figureCitation id="137C2A36FFFA3B7DFC64FA3E74A29D9E" box="[935,1010,1504,1528]" captionStart="Figs 35–52" captionStartId="7.[174,214,2006,2028]" captionTargetBox="[174,1421,255,1986]" captionTargetId="figure-119@7.[173,1421,254,1994]" captionTargetPageId="7" captionText="Figs 35–52: Drusilla bispinosa (35–43), D. shavrini (44–47), and D. penicillata (48–52): male forebody (35, 45, 49); antenna (36, 46, 51); male abdomen (37, 47, 52); anterior portion of male abdomen (38–39); posterior portion of male abdomen in dorsal and in lateral view (40–42); posterior portion of female abdomen (43); habitus (44, 48); head in dorso-lateral view (50). Scale bars: 35–49, 51–52: 1.0 mm; 50: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742772" httpUri="https://zenodo.org/record/5742772/files/figure.png" pageId="26" pageNumber="59">Fig. 49</figureCitation>
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) sexually dimorphic, transverse, 1.05– 1.10 times as broad as long; punctation very fine and moderately sparse, punctures bearing long sub-erect to depressed setae directed mediad; median dorsal portion extensively impunctate; interstices without microreticulation. Eyes moderately large, approximately 1.5 times as long as distance from posterior margin of eye to posterior constriction in dorsal view. Antenna (
|
||
<figureCitation id="137C2A36FFFA3B7DFAC0F91E721E9EBE" box="[1283,1358,1728,1752]" captionStart="Figs 35–52" captionStartId="7.[174,214,2006,2028]" captionTargetBox="[174,1421,255,1986]" captionTargetId="figure-119@7.[173,1421,254,1994]" captionTargetPageId="7" captionText="Figs 35–52: Drusilla bispinosa (35–43), D. shavrini (44–47), and D. penicillata (48–52): male forebody (35, 45, 49); antenna (36, 46, 51); male abdomen (37, 47, 52); anterior portion of male abdomen (38–39); posterior portion of male abdomen in dorsal and in lateral view (40–42); posterior portion of female abdomen (43); habitus (44, 48); head in dorso-lateral view (50). Scale bars: 35–49, 51–52: 1.0 mm; 50: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742772" httpUri="https://zenodo.org/record/5742772/files/figure.png" pageId="26" pageNumber="59">Fig. 51</figureCitation>
|
||
)
|
||
<quantity id="4CBF9B56FFFA3B7DFAA1F91E72E39EB1" box="[1378,1459,1728,1752]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="26" pageNumber="59" unit="mm" value="2.9">2.9 mm</quantity>
|
||
long and slender; antennomeres IV–X oblong, IV twice as long as broad, V–X decreasingly oblong, X weakly oblong, and XI shorter than the combined length of IX and X.
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFFA3B7CFC95F8BE76BF995E" blockId="26.[854,1465,1215,2104]" lastBlockId="27.[174,785,256,1432]" lastPageId="27" lastPageNumber="60" pageId="26" pageNumber="59">
|
||
Pronotum (
|
||
<figureCitation id="137C2A36FFFA3B7DFC12F8BE734C9F1E" box="[977,1052,1888,1912]" captionStart="Figs 35–52" captionStartId="7.[174,214,2006,2028]" captionTargetBox="[174,1421,255,1986]" captionTargetId="figure-119@7.[173,1421,254,1994]" captionTargetPageId="7" captionText="Figs 35–52: Drusilla bispinosa (35–43), D. shavrini (44–47), and D. penicillata (48–52): male forebody (35, 45, 49); antenna (36, 46, 51); male abdomen (37, 47, 52); anterior portion of male abdomen (38–39); posterior portion of male abdomen in dorsal and in lateral view (40–42); posterior portion of female abdomen (43); habitus (44, 48); head in dorso-lateral view (50). Scale bars: 35–49, 51–52: 1.0 mm; 50: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742772" httpUri="https://zenodo.org/record/5742772/files/figure.png" pageId="26" pageNumber="59">Fig. 49</figureCitation>
|
||
) without sexual dimorphism, strongly convex in cross-section, approximately as broad as long and slightly broader than head, broadest at anterior angles; along midline with sharp and narrow sulcus posteriorly terminating in a deep impression; lateral margins indistinctly sinuate in dorsal view; posterior angles obtusely marked; punctation dense and distinct; with several long black setae in lateral portions; interstices without microsculpture.
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFFB3B7CFF6DFE9E75009A71" blockId="27.[174,785,256,1432]" pageId="27" pageNumber="60">
|
||
Elytra (
|
||
<figureCitation id="137C2A36FFFB3B7CFF38FE9E7616993E" box="[251,326,320,344]" captionStart="Figs 35–52" captionStartId="7.[174,214,2006,2028]" captionTargetBox="[174,1421,255,1986]" captionTargetId="figure-119@7.[173,1421,254,1994]" captionTargetPageId="7" captionText="Figs 35–52: Drusilla bispinosa (35–43), D. shavrini (44–47), and D. penicillata (48–52): male forebody (35, 45, 49); antenna (36, 46, 51); male abdomen (37, 47, 52); anterior portion of male abdomen (38–39); posterior portion of male abdomen in dorsal and in lateral view (40–42); posterior portion of female abdomen (43); habitus (44, 48); head in dorso-lateral view (50). Scale bars: 35–49, 51–52: 1.0 mm; 50: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742772" httpUri="https://zenodo.org/record/5742772/files/figure.png" pageId="27" pageNumber="60">Fig. 49</figureCitation>
|
||
) approximately 0.8 times as long as pronotum, strongly transverse (width combined); punctation very dense, coarse, and somewhat asperate, denser near scutellum than elsewhere; interstices reduced to narrow ridges, without microsculpture. Hind wings fully developed. Legs very long and slender; metatarsomere I longer than the combined length of III and IV.
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFFB3B7CFF6DFDFE75C89B1E" blockId="27.[174,785,256,1432]" pageId="27" pageNumber="60">
|
||
Abdomen (
|
||
<figureCitation id="137C2A36FFFB3B7CFEEDFDFE76299A5E" box="[302,377,544,568]" captionStart="Figs 35–52" captionStartId="7.[174,214,2006,2028]" captionTargetBox="[174,1421,255,1986]" captionTargetId="figure-119@7.[173,1421,254,1994]" captionTargetPageId="7" captionText="Figs 35–52: Drusilla bispinosa (35–43), D. shavrini (44–47), and D. penicillata (48–52): male forebody (35, 45, 49); antenna (36, 46, 51); male abdomen (37, 47, 52); anterior portion of male abdomen (38–39); posterior portion of male abdomen in dorsal and in lateral view (40–42); posterior portion of female abdomen (43); habitus (44, 48); head in dorso-lateral view (50). Scale bars: 35–49, 51–52: 1.0 mm; 50: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742772" httpUri="https://zenodo.org/record/5742772/files/figure.png" pageId="27" pageNumber="60">Fig. 52</figureCitation>
|
||
) approximately as broad as elytra; tergite III with moderately dense and distinct setiferous punctation; tergite IV with a variable number of setiferous setae at posterior margin and with a transverse row of setiferous setae across middle; tergites V–VII with setiferous punctures at posterior margins and with very sparse punctation elsewhere; tergites III–VI with shallow and very fine transverse microsculpture visible only at high magnification; tergites VII–VIII with shallow microsculpture composed of transverse meshes; posterior margin of tergite VII with palisade fringe.
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFFB3B7CFF00FC5E76DD9CBE" blockId="27.[174,785,256,1432]" pageId="27" pageNumber="60">
|
||
: head (
|
||
<figureCitation id="137C2A36FFFB3B7CFEDBFC5E76359BFE" box="[280,357,896,920]" captionStart="Figs 35–52" captionStartId="7.[174,214,2006,2028]" captionTargetBox="[174,1421,255,1986]" captionTargetId="figure-119@7.[173,1421,254,1994]" captionTargetPageId="7" captionText="Figs 35–52: Drusilla bispinosa (35–43), D. shavrini (44–47), and D. penicillata (48–52): male forebody (35, 45, 49); antenna (36, 46, 51); male abdomen (37, 47, 52); anterior portion of male abdomen (38–39); posterior portion of male abdomen in dorsal and in lateral view (40–42); posterior portion of female abdomen (43); habitus (44, 48); head in dorso-lateral view (50). Scale bars: 35–49, 51–52: 1.0 mm; 50: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742772" httpUri="https://zenodo.org/record/5742772/files/figure.png" pageId="27" pageNumber="60">Fig. 50</figureCitation>
|
||
) in the middle of dorsal surface with small tubercle bearing approximately 4–5 long pale setae; tergite VIII (Fig. 140) with posterior margin weakly concave in the middle, in anterior portion with numerous gland openings; sternite VIII much longer than tergite VIII, with convex posterior margin, in anterior portion with numerous gland openings; median lobe of aedeagus approximately
|
||
<quantity id="4CBF9B56FFFB3B7CFDBFFBBE759F9C11" box="[636,719,1120,1144]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="60" unit="mm" value="0.9">0.9 mm</quantity>
|
||
long and shaped as in Figs 141–142; ventral process apically abruptly narrowed in ventral view; parameres approximately
|
||
<quantity id="4CBF9B56FFFB3B7CFF3EFB1E76009CB1" box="[253,336,1216,1240]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="27" pageNumber="60" unit="mm" value="0.8">0.8 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFFB3B7CFF01FB3E75F39DFE" blockId="27.[174,785,256,1432]" pageId="27" pageNumber="60">: tergite VIII (Fig. 143) with moderately sparse gland openings anteriorly, posterior margin very weakly concave in the middle, nearly truncate; sternite VIII with moderately sparse gland openings anteriorly, posterior margin broadly convex, in the middle with short modified marginal setae; spermatheca as in Fig. 144.</paragraph>
|
||
<paragraph id="8BF836B3FFFB3B7CFF6DFA6173D8991E" blockId="27.[174,785,1471,2104]" lastBlockId="27.[816,1427,256,376]" pageId="27" pageNumber="60">
|
||
<emphasis id="B933EAA1FFFB3B7CFF6DFA6176D19DBE" bold="true" box="[174,385,1471,1496]" pageId="27" pageNumber="60">Comparative notes</emphasis>
|
||
: As can be inferred from the similar general morphology of the genitalia,
|
||
<taxonomicName id="4C474D30FFFB3B7CFD43FA3E745A9D9E" authorityName="Assing" authorityYear="2019" box="[640,778,1504,1528]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="60" phylum="Arthropoda" rank="species" species="penicillata">
|
||
<emphasis id="B933EAA1FFFB3B7CFD43FA3E745A9D9E" box="[640,778,1504,1528]" italics="true" pageId="27" pageNumber="60">D. penicillata</emphasis>
|
||
</taxonomicName>
|
||
belongs to the same species group as
|
||
<taxonomicName id="4C474D30FFFB3B7CFD99F9DE759E9E7E" authorityName="Assing" authorityYear="2019" box="[602,718,1536,1560]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="60" phylum="Arthropoda" rank="species" species="shavrini">
|
||
<emphasis id="B933EAA1FFFB3B7CFD99F9DE759E9E7E" box="[602,718,1536,1560]" italics="true" pageId="27" pageNumber="60">D. shavrini</emphasis>
|
||
</taxonomicName>
|
||
. It is distinguished from all other species previously recorded from the
|
||
<collectingCountry id="F3507623FFFB3B7CFEDBF99E76C09E3E" box="[280,400,1600,1624]" name="Philippines" pageId="27" pageNumber="60">Philippines</collectingCountry>
|
||
(see the comparative notes in the section on
|
||
<taxonomicName id="4C474D30FFFB3B7CFEE7F9BE76C89E1E" authorityName="Assing" authorityYear="2019" box="[292,408,1632,1656]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="60" phylum="Arthropoda" rank="species" species="shavrini">
|
||
<emphasis id="B933EAA1FFFB3B7CFEE7F9BE76C89E1E" box="[292,408,1632,1656]" italics="true" pageId="27" pageNumber="60">D. shavrini</emphasis>
|
||
</taxonomicName>
|
||
) and the new species described in the present paper by the coloration and larger body size alone. In addition, it is characterized by the brush-like tubercle on the male head and by the shapes of the median lobe of the aedeagus and of the spermatheca. Regarding the coloration and the general morphology of the genitalia,
|
||
<taxonomicName id="4C474D30FFFB3B7CFF16F8FE76309F5E" authorityName="Assing" authorityYear="2019" box="[213,352,1824,1848]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="60" phylum="Arthropoda" rank="species" species="penicillata">
|
||
<emphasis id="B933EAA1FFFB3B7CFF16F8FE76309F5E" box="[213,352,1824,1848]" italics="true" pageId="27" pageNumber="60">D. penicillata</emphasis>
|
||
</taxonomicName>
|
||
somewhat resembles
|
||
<taxonomicName id="4C474D30FFFB3B7CFD8EF8FE759E9F5E" authorityName="ASSING" authorityYear="2015" box="[589,718,1824,1848]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="60" phylum="Arthropoda" rank="species" species="perforans">
|
||
<emphasis id="B933EAA1FFFB3B7CFD8EF8FE759E9F5E" box="[589,718,1824,1848]" italics="true" pageId="27" pageNumber="60">D. perforans</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3507623FFFB3B7CFF6DF89E765C9F3E" box="[174,268,1856,1880]" name="Vietnam" pageId="27" pageNumber="60">Vietnam</collectingCountry>
|
||
, from which it differs by larger body size, the coloration of the antennae, legs, elytra, and abdomen (
|
||
<taxonomicName id="4C474D30FFFB3B7CFF74F85E76689FFE" authorityName="ASSING" authorityYear="2015" box="[183,312,1920,1944]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="60" phylum="Arthropoda" rank="species" species="perforans">
|
||
<emphasis id="B933EAA1FFFB3B7CFF74F85E76689FFE" box="[183,312,1920,1944]" italics="true" pageId="27" pageNumber="60">D. perforans</emphasis>
|
||
</taxonomicName>
|
||
: antennomeres III–X infuscate; legs yellow with the apices of the metafemora infuscate; elytra infuscate in postero-lateral portions; abdomen yellow with the posterior portions of paratergites III–VI, the posterior portion of tergite V, most of tergite VI, and the median portion of tergite VII blackish), by the modifications of the male head, by a larger and less slender pronotum without a broad median impression, with weakly sinuate lateral margins, and with much finer and non-granulose punctation, and by the genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D6538FFFB3B7CFCF3FE4172179A7E" pageId="27" pageNumber="60" type="distribution">
|
||
<paragraph id="8BF836B3FFFB3B7CFCF3FE4172179A7E" blockId="27.[816,1427,415,536]" pageId="27" pageNumber="60">
|
||
<emphasis id="B933EAA1FFFB3B7CFCF3FE4173F099D1" bold="true" box="[816,1184,415,440]" pageId="27" pageNumber="60">Distribution and natural history</emphasis>
|
||
: The
|
||
<typeStatus id="54FC8811FFFB3B7CFB26FE7E724399DE" box="[1253,1299,416,440]" pageId="27" pageNumber="60">type</typeStatus>
|
||
specimens were collected in two geographically close localities in the Central Mindanao,
|
||
<collectingCountry id="F3507623FFFB3B7CFC3CFE3E7329999E" box="[1023,1145,480,504]" name="Philippines" pageId="27" pageNumber="60">Philippines</collectingCountry>
|
||
, by sifting wet litter near a waterfall and in a a secondary broad-leaved forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |