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<document ID-DOI="10.1071/is21067" ID-GBIF-Dataset="8c0715c9-57f5-40db-8c48-99bf3719eedb" ID-Zenodo-Dep="6987297" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1660328621757" checkinUser="felipe" docAuthor="Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan &amp; Matos-Maraví, Pável" docDate="2022" docId="03E5E91565100538FF0DFA689A022EAE" docLanguage="en" docName="InvSyst.36.6.557-579.pdf" docOrigin="Invertebrate Systematics 36 (6)" docSource="http://dx.doi.org/10.1071/is21067" docTitle="Overbeckia papuana Klimeš &amp; Drescher &amp; Buchori &amp; Hidayat &amp; Nazarreta &amp; Potocký &amp; Rimandai &amp; Scheu &amp; Matos-Maraví 2022, sp. nov." docType="treatment" docVersion="4" lastPageNumber="571" masterDocId="FFDC916D651D0536FFBCFFF69F1B2B6D" masterDocTitle="Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae)" masterLastPageNumber="288" masterPageNumber="277" pageNumber="570" updateTime="1660340715462" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Klimeš, Petr</mods:namePart>
<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, BraniŠovská 31, CZ- 370 05 České Budějovice, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Drescher, Jochen</mods:namePart>
<mods:affiliation>Department of Animal Ecology, University of Göttingen, J. - F. - Blumenbach Institute for Zoology &amp; Anthropology, Untere Karspüle 2, D- 37073 Göttingen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Buchori, Damayanti</mods:namePart>
<mods:affiliation>Department of Plant Protection, Faculty of Agriculture, Bogor Agricultural University, Jl. Kamper, Kampus IPB Dramaga, Bogor 16680, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hidayat, Purnama</mods:namePart>
<mods:affiliation>Department of Plant Protection, Faculty of Agriculture, Bogor Agricultural University, Jl. Kamper, Kampus IPB Dramaga, Bogor 16680, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nazarreta, Rizky</mods:namePart>
<mods:affiliation>Department of Plant Protection, Faculty of Agriculture, Bogor Agricultural University, Jl. Kamper, Kampus IPB Dramaga, Bogor 16680, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Potocký, Pavel</mods:namePart>
<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, BraniŠovská 31, CZ- 370 05 České Budějovice, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rimandai, Maling</mods:namePart>
<mods:affiliation>New Guinea Binatang Research Center, PO Box 604, Madang, Papua New Guinea</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scheu, Stefan</mods:namePart>
<mods:affiliation>Department of Animal Ecology, University of Göttingen, J. - F. - Blumenbach Institute for Zoology &amp; Anthropology, Untere Karspüle 2, D- 37073 Göttingen, Germany &amp; Centre of Biodiversity and Sustainable Land Use, University of Göttingen, Büsgenweg 1, D- 37077 Göttingen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Matos-Maraví, Pável</mods:namePart>
<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, BraniŠovská 31, CZ- 370 05 České Budějovice, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Invertebrate Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-07-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>36</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>6</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>277</mods:start>
<mods:end>288</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1071/is21067</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1071/is21067</mods:identifier>
<mods:identifier type="GBIF-Dataset">8c0715c9-57f5-40db-8c48-99bf3719eedb</mods:identifier>
<mods:identifier type="Zenodo-Dep">6987297</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6987104" ID-Zenodo-Dep="6987104" LSID="urn:lsid:plazi:treatment:03E5E91565100538FF0DFA689A022EAE" httpUri="http://treatment.plazi.org/id/03E5E91565100538FF0DFA689A022EAE" lastPageId="14" lastPageNumber="571" pageId="13" pageNumber="570">
<subSubSection box="[177,690,1438,1469]" pageId="13" pageNumber="570" type="nomenclature">
<paragraph blockId="13.[177,690,1438,1469]" box="[177,690,1438,1469]" pageId="13" pageNumber="570">
<heading bold="true" box="[177,690,1438,1469]" centered="true" fontSize="12" level="2" pageId="13" pageNumber="570" reason="0">
<emphasis bold="true" box="[177,690,1438,1469]" pageId="13" pageNumber="570">
<taxonomicName authority="Klimes" authorityName="Klimeš &amp; Drescher &amp; Buchori &amp; Hidayat &amp; Nazarreta &amp; Potocký &amp; Rimandai &amp; Scheu &amp; Matos-Maraví" authorityYear="2022" box="[177,574,1438,1469]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="570" phylum="Arthropoda" rank="species" species="papuana" status="sp. nov.">
<emphasis bold="true" box="[177,474,1438,1469]" italics="true" pageId="13" pageNumber="570">Overbeckia papuana</emphasis>
Klimes
</taxonomicName>
,
<taxonomicNameLabel box="[589,690,1438,1468]" pageId="13" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[377,491,1507,1532]" pageId="13" pageNumber="570" type="description">
<paragraph blockId="13.[377,491,1507,1532]" box="[377,491,1507,1532]" pageId="13" pageNumber="570">
<heading box="[377,491,1507,1532]" centered="true" fontSize="10" level="4" pageId="13" pageNumber="570" reason="4">
(
<figureCitation box="[386,451,1507,1531]" captionStart="Fig" captionStartId="9.[314,351,1878,1901]" captionTargetBox="[314,1274,1129,1848]" captionTargetId="figure-2@9.[313,1274,1128,1848]" captionTargetPageId="9" captionText="Fig. 5. Overbeckia papuana sp. nov. holotype (worker) and its lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d)." figureDoi="http://doi.org/10.5281/zenodo.6987307" httpUri="https://zenodo.org/record/6987307/files/figure.png" pageId="13" pageNumber="570">Fig. 5</figureCitation>
,
<figureCitation box="[467,481,1507,1531]" captionStart="Fig" captionStartId="10.[154,191,1653,1676]" captionTargetBox="[154,1434,177,1622]" captionTargetId="figure-75@10.[154,1435,176,1623]" captionTargetPageId="10" captionText="Fig. 6. Overbeckia papuana sp. nov. paratypes (alates). Queen: lateral (a), dorsal (c) and frontal (e) views; male: lateral (b), dorsal (d) and frontal (f) views. Note that the petiole in queen (a) looks wider and blunter due to being bent to the right site (c)." figureDoi="http://doi.org/10.5281/zenodo.6987309" httpUri="https://zenodo.org/record/6987309/files/figure.png" pageId="13" pageNumber="570">6</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="570" type="etymology">
<paragraph blockId="13.[98,245,1599,1627]" box="[98,245,1599,1627]" pageId="13" pageNumber="570">
<heading bold="true" box="[98,245,1599,1627]" fontSize="11" level="3" pageId="13" pageNumber="570" reason="0">
<emphasis bold="true" box="[98,245,1599,1627]" pageId="13" pageNumber="570">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[98,770,1650,1738]" pageId="13" pageNumber="570">
Dedicated to all Papua New Guineans, the nation of the country from which the
<typeStatus box="[375,424,1683,1707]" pageId="13" pageNumber="570">type</typeStatus>
and nest series were collected (
<collectingCountry box="[105,329,1714,1738]" name="Papua New Guinea" pageId="13" pageNumber="570">Papua New Guinea</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="570" type="materials_examined">
<paragraph blockId="13.[98,771,1785,1931]" box="[98,356,1785,1813]" pageId="13" pageNumber="570">
<heading bold="true" box="[98,356,1785,1813]" fontSize="11" level="3" pageId="13" pageNumber="570" reason="0">
<emphasis bold="true" box="[98,356,1785,1813]" pageId="13" pageNumber="570">Material examined</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[98,771,1785,1931]" pageId="13" pageNumber="570">
<materialsCitation collectingDate="2007-03-08" collectionCode="ZMHB" collectorName="M. Rimandai" country="Papua New Guinea" latitude="-5.233333" location="Wanang village" longLatPrecision="1305" longitude="145.18333" pageId="13" pageNumber="570" specimenCode="CAMP021" specimenCount="1" specimenCount-worker="1" stateProvince="Madang province" typeStatus="holotype">
<emphasis bold="true" box="[98,187,1831,1850]" italics="true" pageId="13" pageNumber="570">
<typeStatus box="[98,183,1831,1850]" pageId="13" pageNumber="570">Holotype</typeStatus>
.
</emphasis>
<specimenCount box="[193,261,1831,1850]" pageId="13" pageNumber="570" type="worker">Worker</specimenCount>
(
<collectionCode box="[274,339,1832,1851]" pageId="13" pageNumber="570">ZMHB</collectionCode>
): Morphospecies
<specimenCode box="[500,599,1832,1851]" pageId="13" pageNumber="570">CAMP021</specimenCode>
/Acc.N. HP0177/ Tree number code WS4D0868/Secondary forest/
<location LSID="urn:lsid:plazi:treatment:03E5E91565100538FF0DFA689A022EAE:8E930ED86510053BFDE0F8B49DE12C38" box="[604,762,1858,1877]" country="Papua New Guinea" latitude="-5.233333" longLatPrecision="1305" longitude="145.18333" name="Wanang village" pageId="13" pageNumber="570" stateProvince="Madang province">Wanang village</location>
/
<collectingRegion box="[98,263,1885,1904]" country="Papua New Guinea" name="Madang" pageId="13" pageNumber="570">Madang province</collectingRegion>
/
<collectingCountry box="[270,460,1884,1904]" name="Papua New Guinea" pageId="13" pageNumber="570">Papua New Guinea</collectingCountry>
/
<date box="[468,554,1884,1904]" pageId="13" pageNumber="570" value="2007-03-08">
<collectingDate box="[468,554,1884,1904]" pageId="13" pageNumber="570" value="2007-03-08">8.iii.2007</collectingDate>
</date>
/
<geoCoordinate box="[562,632,1884,1904]" degrees="05" direction="south" minutes="14" orientation="latitude" pageId="13" pageNumber="570" precision="925" value="-5.233333">05°14S</geoCoordinate>
,
<geoCoordinate box="[644,728,1884,1904]" degrees="145" direction="east" minutes="11" orientation="longitude" pageId="13" pageNumber="570" precision="925" value="145.18333">145°11E</geoCoordinate>
/leg.
<collectorName box="[98,220,1912,1931]" pageId="13" pageNumber="570">M. Rimandai</collectorName>
.
</materialsCitation>
</paragraph>
<paragraph blockId="13.[818,1491,178,704]" pageId="13" pageNumber="570">
<materialsCitation pageId="13" pageNumber="570">
<emphasis bold="true" box="[850,947,178,197]" italics="true" pageId="13" pageNumber="570">
<typeStatus box="[850,943,178,197]" pageId="13" pageNumber="570">Paratypes</typeStatus>
.
</emphasis>
Five workers (1w
<collectionCode box="[1128,1191,179,198]" pageId="13" pageNumber="570">ZMHB</collectionCode>
, 1w
<collectionCode box="[1236,1286,178,197]" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="13" pageNumber="570" type="Other institutional type">IECA</collectionCode>
, 2w
<collectionCode box="[1333,1386,178,197]" country="Papua New Guinea" httpUri="http://grbio.org/cool/7mi7-dyvm" name="National Agricultural Insect Collection" pageId="13" pageNumber="570">NAIC</collectionCode>
, 1w
<collectionCode box="[1432,1481,178,197]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="13" pageNumber="570" type="Museum">MZB</collectionCode>
),
<specimenCount box="[818,906,205,224]" pageId="13" pageNumber="570" type="male">one male</specimenCount>
(
<collectionCode box="[920,986,205,224]" pageId="13" pageNumber="570">ZMHB</collectionCode>
),
</materialsCitation>
<materialsCitation pageId="13" pageNumber="570">
<specimenCount box="[1004,1103,205,224]" pageId="13" pageNumber="570" type="queen">one queen</specimenCount>
without wings (
<collectionCode box="[1262,1329,205,224]" pageId="13" pageNumber="570">ZMHB</collectionCode>
): From the same nest series as
<typeStatus box="[949,1032,232,251]" pageId="13" pageNumber="570">holotype</typeStatus>
(the same data).
</materialsCitation>
</paragraph>
<paragraph blockId="13.[818,1491,178,704]" pageId="13" pageNumber="570">
<emphasis bold="true" box="[850,1339,258,277]" pageId="13" pageNumber="570">
<emphasis bold="true" box="[850,1135,258,277]" italics="true" pageId="13" pageNumber="570">Additional material examined.</emphasis>
<materialsCitation pageId="13" pageNumber="570">
<collectingCountry box="[1142,1335,258,277]" name="Papua New Guinea" pageId="13" pageNumber="570">Papua New Guinea</collectingCountry>
: Other
<specimenCount pageId="13" pageNumber="570" type="worker">30 workers</specimenCount>
and
<specimenCount box="[892,966,285,304]" pageId="13" pageNumber="570" type="male">2 males</specimenCount>
, including brood, kept in one vial with ethanol in
<collectionCode box="[1438,1487,285,304]" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="13" pageNumber="570" type="Other institutional type">IECA</collectionCode>
, all from the same nest as the
<typeStatus box="[1117,1157,312,331]" pageId="13" pageNumber="570">type</typeStatus>
specimens (same data).
</materialsCitation>
</emphasis>
<materialsCitation pageId="13" pageNumber="570">
<emphasis bold="true" box="[1400,1490,311,330]" pageId="13" pageNumber="570">
<collectingCountry box="[1400,1490,311,330]" name="Australia" pageId="13" pageNumber="570">Australia</collectingCountry>
</emphasis>
(photos):
<specimenCount box="[922,1021,338,357]" pageId="13" pageNumber="570" type="worker">1 worker</specimenCount>
(
<collectionCode box="[1049,1113,338,357]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="13" pageNumber="570" type="Museum">SAMA</collectionCode>
):
<taxonomicName baseAuthorityName="Eisner and Wilson" baseAuthorityYear="1952" box="[1149,1263,338,357]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1149,1263,338,357]" italics="true" pageId="13" pageNumber="570">Camponotus</emphasis>
</taxonomicName>
/Cairns NQ/on tree/ Parkland/
<date box="[915,1012,365,384]" pageId="13" pageNumber="570" value="1975-08-02">2.viii.1975</date>
/leg.
<collectionCode box="[1065,1080,365,384]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="13" pageNumber="570" type="Herbarium">B</collectionCode>
.
<collectionCode box="[1084,1099,365,384]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="13" pageNumber="570" type="Herbarium">B</collectionCode>
. Lowery (frontal and lateral photo on p.
<quantity box="[834,895,391,410]" metricMagnitude="0" metricUnit="m" metricValue="2.921" pageId="13" pageNumber="570" unit="in" value="115.0">115 in</quantity>
<bibRefCitation author="McArthur A" box="[903,1062,391,411]" pageId="13" pageNumber="570" refId="ref18953" refString="McArthur A (2012) ' A guide to Colobopsis ants of the world. ' (South Australian Museum: Adelaide, SA, Australia)" type="book" year="2012">McArthur (2012)</bibRefCitation>
and
<collectionCode box="[1110,1127,392,411]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="13" pageNumber="570" type="Herbarium">A</collectionCode>
. McArthur, pers. comm. for coll. information; previously misidentified by the original author for
<taxonomicName authorityName="Forel" authorityYear="1895" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="570" phylum="Arthropoda" rank="species" species="janeti">
<emphasis bold="true" italics="true" pageId="13" pageNumber="570">Camponotus janeti</emphasis>
</taxonomicName>
).
</materialsCitation>
<materialsCitation pageId="13" pageNumber="570">
<specimenCount box="[907,1015,445,464]" pageId="13" pageNumber="570" type="worker">3 workers</specimenCount>
(
<collectionCode box="[1045,1104,445,464]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="13" pageNumber="570" type="Museum">WAM</collectionCode>
): CASENT0872736, CASENT0872753, CASENT0872754 (AntWeb codes;
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1157,1260,471,490]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1157,1260,471,490]" italics="true" pageId="13" pageNumber="570">Overbeckia</emphasis>
</taxonomicName>
wam01):
<collectingRegion box="[1372,1483,472,491]" country="Australia" name="Queensland" pageId="13" pageNumber="570">Queensland</collectingRegion>
/ Mossman/
<date box="[921,1015,498,517]" pageId="13" pageNumber="570" value="2002-09-18">18.ix.2002</date>
/leg,
<collectionCode box="[1067,1084,499,518]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="570">R</collectionCode>
.
<collectingCountry box="[1098,1163,498,517]" name="Jordan" pageId="13" pageNumber="570">Jordan</collectingCountry>
/nest in dead twig in living tree behind sugar mill (thicket near Mossman
<collectionCode box="[1263,1281,525,544]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="570">R</collectionCode>
.)/det.
<collectionCode box="[1351,1367,525,544]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="13" pageNumber="570" type="Herbarium">B</collectionCode>
.
<collectionCode box="[1370,1386,525,544]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="13" pageNumber="570" type="Herbarium">E</collectionCode>
. Heterick,
<date box="[818,888,552,571]" pageId="13" pageNumber="570" value="2018-02">ii.2018</date>
.
</materialsCitation>
<materialsCitation pageId="13" pageNumber="570">
<specimenCount box="[915,1019,552,571]" pageId="13" pageNumber="570" type="worker">1 worker</specimenCount>
(
<collectionCode box="[1050,1110,552,571]" pageId="13" pageNumber="570">CASC</collectionCode>
): CASENT0887749 (AntWeb code;
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[823,926,578,597]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[823,926,578,597]" italics="true" pageId="13" pageNumber="570">Overbeckia</emphasis>
</taxonomicName>
au01):
<collectingRegion box="[1019,1131,578,597]" country="Australia" name="Queensland" pageId="13" pageNumber="570">Queensland</collectingRegion>
/Mt Peters/
<date box="[1255,1333,578,597]" pageId="13" pageNumber="570" value="2014-05-01">
1.
<emphasis bold="true" box="[1266,1282,578,597]" italics="true" pageId="13" pageNumber="570">v.</emphasis>
2014
</date>
/leg,
<collectionCode box="[1388,1410,579,598]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="13" pageNumber="570" type="Herbarium">M</collectionCode>
. Collis/ det.
<collectionCode box="[861,879,605,624]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="13" pageNumber="570" type="Herbarium">A</collectionCode>
. Andersen
<date box="[985,1089,605,624]" pageId="13" pageNumber="570" value="2020-11-30">30.xi.2020</date>
.
</materialsCitation>
<materialsCitation latitude="-11.633333" longitude="142.85" pageId="13" pageNumber="570">
<emphasis bold="true" box="[1100,1192,605,624]" pageId="13" pageNumber="570">
<collectingCountry box="[1100,1192,605,624]" name="Australia" pageId="13" pageNumber="570">Australia</collectingCountry>
</emphasis>
(physical specimen):
<specimenCount box="[1403,1490,605,624]" pageId="13" pageNumber="570" type="worker">1 worker</specimenCount>
(
<collectionCode box="[824,891,632,651]" pageId="13" pageNumber="570">ZMHB</collectionCode>
): n#15720/
<collectionCode box="[1002,1065,631,650]" pageId="13" pageNumber="570">AUST</collectionCode>
Qld/Captain Billy Landing/rainforest, ex. dead hanging twig/
<geoCoordinate box="[1004,1076,657,677]" degrees="11" direction="south" minutes="38" orientation="latitude" pageId="13" pageNumber="570" precision="925" value="-11.633333">11°38S</geoCoordinate>
,
<geoCoordinate box="[1085,1173,657,677]" degrees="142" direction="east" minutes="51" orientation="longitude" pageId="13" pageNumber="570" precision="925" value="142.85">142°51E</geoCoordinate>
/leg.
<collectionCode box="[1220,1235,659,678]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="13" pageNumber="570" type="Herbarium">P</collectionCode>
. Ward (the same collection series as CASENT0882197 at AntWeb;
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1194,1297,685,704]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1194,1297,685,704]" italics="true" pageId="13" pageNumber="570">Overbeckia</emphasis>
</taxonomicName>
au01).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="570" type="diagnosis">
<paragraph blockId="13.[818,948,747,775]" box="[818,948,747,775]" pageId="13" pageNumber="570">
<heading bold="true" box="[818,948,747,775]" fontSize="11" level="3" pageId="13" pageNumber="570" reason="0">
<emphasis bold="true" box="[818,948,747,775]" pageId="13" pageNumber="570">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[818,1490,798,1079]" pageId="13" pageNumber="570">
General appearance of worker and queen as in
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1333,1482,798,823]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="570" phylum="Arthropoda" rank="species" species="subclavata">
<emphasis bold="true" box="[1333,1482,798,823]" italics="true" pageId="13" pageNumber="570">O. subclavata</emphasis>
</taxonomicName>
, but hairier on the head, with erect hairs on dorsum of petiole, thorax and abdomen; a few hairs also present on distal part of scape. Head integument less punctate than in
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[818,972,926,950]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="570" phylum="Arthropoda" rank="species" species="subclavata">
<emphasis bold="true" box="[818,972,926,950]" italics="true" pageId="13" pageNumber="570">O. subclavata</emphasis>
</taxonomicName>
, with many short-erect hairs extending past head margins in frontal view; tiny appressed hairs in high densities all over the head including genae; frontal carinae significantly approaching one another anteriorly (
<collectionCode box="[824,882,1055,1079]" pageId="13" pageNumber="570">FCDI</collectionCode>
&gt; 190) (
<figureCitation box="[987,1064,1054,1079]" captionStart="Fig" captionStartId="9.[314,351,1878,1901]" captionTargetBox="[314,1274,1129,1848]" captionTargetId="figure-2@9.[313,1274,1128,1848]" captionTargetPageId="9" captionText="Fig. 5. Overbeckia papuana sp. nov. holotype (worker) and its lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d)." figureDoi="http://doi.org/10.5281/zenodo.6987307" httpUri="https://zenodo.org/record/6987307/files/figure.png" pageId="13" pageNumber="570">
Fig. 5
<emphasis bold="true" box="[1051,1064,1054,1078]" italics="true" pageId="13" pageNumber="570">d</emphasis>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="571" pageId="13" pageNumber="570" type="description">
<paragraph blockId="13.[818,1122,1121,1149]" box="[818,1122,1121,1149]" pageId="13" pageNumber="570">
<heading bold="true" box="[818,1122,1121,1149]" fontSize="11" level="3" pageId="13" pageNumber="570" reason="0">
<emphasis bold="true" box="[818,1122,1121,1149]" pageId="13" pageNumber="570">Description of worker</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[818,1490,1171,1933]" pageId="13" pageNumber="570">
<emphasis bold="true" box="[818,982,1171,1195]" italics="true" pageId="13" pageNumber="570">Morphometrics</emphasis>
.
<typeStatus box="[1014,1120,1172,1196]" pageId="13" pageNumber="570">Holotype</typeStatus>
, HP0177 nest series (
<specimenCount pageId="13" pageNumber="570" type="generic" typeStatus="paratypes">five paratypes</specimenCount>
; total
<emphasis bold="true" box="[1012,1030,1204,1227]" italics="true" pageId="13" pageNumber="570">
<collectionCode box="[1012,1030,1204,1227]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="13" pageNumber="570" type="Herbarium">N</collectionCode>
</emphasis>
= 6, min.max):
<collectionCode box="[1242,1274,1204,1227]" country="USA" lsid="urn:lsid:biocol.org:col:13245" name="Houghton Lake Wildlife Research Station" pageId="13" pageNumber="570" type="Herbarium">HL</collectionCode>
1.08 (1.041.41), HW 1.10 (1.051.43), IOD 0.82 (0.791.10),
<collectionCode box="[1392,1421,1236,1260]" country="Romania" lsid="urn:lsid:biocol.org:col:15579" name="Babes-Bolyai University" pageId="13" pageNumber="570" type="Herbarium">CL</collectionCode>
0.17 (0.140.23), CW 0.41 (0.390.55),
<collectionCode box="[1238,1274,1268,1291]" country="France" name="Musee de Lectoure" pageId="13" pageNumber="570">ML</collectionCode>
0.44 (0.360.57), EL 0.28 (0.260.34), EW 0.21 (0.190.27), FCDp 0.37 (0.350.52), FCDa 0.18 (0.150.27),
<collectionCode box="[1251,1278,1332,1356]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="13" pageNumber="570" type="Herbarium">SL</collectionCode>
1.05 (1.021.24), SWmax 0.14 (0.120.18), SWmin 0.06 (0.060.09),
<collectionCode box="[1396,1433,1364,1387]" country="China" lsid="urn:lsid:biocol.org:col:14766" name="Wolong Nature Reserve" pageId="13" pageNumber="570" type="Herbarium">WL</collectionCode>
1.58 (1.522.02),
<collectionCode box="[968,1006,1396,1419]" country="Thailand" name="Paleontological Collections" pageId="13" pageNumber="570">PW</collectionCode>
0.77 (0.691.03), HFL 1.09 (0.921.24), HFT 0.24 (0.240.30), HTL 0.94 (0.821.05), PetW 0.39 (0.370.47), PetL 0.17 (0.170.24),
<collectionCode box="[1243,1299,1460,1484]" pageId="13" pageNumber="570">FCDI</collectionCode>
208 (193236),
<collectionCode box="[818,842,1492,1516]" country="USA" lsid="urn:lsid:biocol.org:col:15344" name="Carnegie Institution of Washington" pageId="13" pageNumber="570" type="Herbarium">CI</collectionCode>
102 (99105),
<collectionCode box="[1020,1042,1492,1516]" country="Argentina" name="Museo Botánico (SI)" pageId="13" pageNumber="570" type="Herbarium">SI</collectionCode>
95 (87101),
<collectionCode box="[1206,1237,1492,1515]" country="Italy" lsid="urn:lsid:biocol.org:col:14381" name="Università degli Studi di Milano" pageId="13" pageNumber="570" type="Herbarium">MI</collectionCode>
40 (3043).
</paragraph>
<paragraph blockId="13.[818,1490,1171,1933]" pageId="13" pageNumber="570">
<emphasis bold="true" box="[850,907,1524,1548]" italics="true" pageId="13" pageNumber="570">Head</emphasis>
. General morphology as for the genus (see above). Frontal carinae relatively more approaching one another anteriorly above the clypeal margin compared to those of the other species (
<collectionCode box="[1020,1078,1620,1644]" pageId="13" pageNumber="570">FCDI</collectionCode>
= 208), with rather straight carinae curvature all along (only slightly convex laterally adjacent to the antennal sockets at frontal view;
<figureCitation box="[1270,1347,1684,1708]" captionStart="Fig" captionStartId="9.[314,351,1878,1901]" captionTargetBox="[314,1274,1129,1848]" captionTargetId="figure-2@9.[313,1274,1128,1848]" captionTargetPageId="9" captionText="Fig. 5. Overbeckia papuana sp. nov. holotype (worker) and its lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d)." figureDoi="http://doi.org/10.5281/zenodo.6987307" httpUri="https://zenodo.org/record/6987307/files/figure.png" pageId="13" pageNumber="570">
Fig. 5
<emphasis bold="true" box="[1334,1347,1684,1708]" italics="true" pageId="13" pageNumber="570">d</emphasis>
</figureCitation>
).
</paragraph>
<paragraph blockId="13.[818,1490,1171,1933]" box="[850,1490,1716,1741]" pageId="13" pageNumber="570">
<emphasis bold="true" box="[850,1131,1716,1740]" italics="true" pageId="13" pageNumber="570">Mesosoma and metasoma</emphasis>
. As for the genus (see above).
</paragraph>
<paragraph blockId="13.[818,1490,1171,1933]" lastBlockId="14.[98,770,180,844]" lastPageId="14" lastPageNumber="571" pageId="13" pageNumber="570">
<emphasis bold="true" box="[850,930,1747,1771]" italics="true" pageId="13" pageNumber="570">Pilosity</emphasis>
. Tiny appressed hairs (approximately twice the length of ommatidium or smaller) present in high densities over the head, mesosoma and abdomen; relatively long erect hairs (length ≥ SWmin) present on clypeus, head integument and all over abdomen; these longer hairs are also present on genae and occipital corners of the head and on frons (extending past head margins from the frontal view;
<figureCitation box="[98,175,211,236]" captionStart="Fig" captionStartId="9.[314,351,1878,1901]" captionTargetBox="[314,1274,1129,1848]" captionTargetId="figure-2@9.[313,1274,1128,1848]" captionTargetPageId="9" captionText="Fig. 5. Overbeckia papuana sp. nov. holotype (worker) and its lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d)." figureDoi="http://doi.org/10.5281/zenodo.6987307" httpUri="https://zenodo.org/record/6987307/files/figure.png" pageId="14" pageNumber="571">
Fig. 5
<emphasis bold="true" box="[163,175,211,235]" italics="true" pageId="14" pageNumber="571">b</emphasis>
</figureCitation>
); scape with relatively short, appressed hairs (as in
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[98,246,244,268]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="subclavata">
<emphasis bold="true" box="[98,246,244,268]" italics="true" pageId="14" pageNumber="571">O. subclavata</emphasis>
</taxonomicName>
), with a few erect hairs not limited to the apex but present ventrally along the anterior half of the scape. These hairs shorter and less numerous than in
<taxonomicName authorityName="Klimeš &amp; Drescher &amp; Buchori &amp; Hidayat &amp; Nazarreta &amp; Potocký &amp; Rimandai &amp; Scheu &amp; Matos-Maraví" authorityYear="2022" box="[622,769,307,332]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="jambiensis">
<emphasis bold="true" box="[622,769,307,332]" italics="true" pageId="14" pageNumber="571">O. jambiensis</emphasis>
</taxonomicName>
(
<figureCitation box="[107,188,340,364]" captionStart="Fig" captionStartId="9.[314,351,1878,1901]" captionTargetBox="[314,1274,1129,1848]" captionTargetId="figure-2@9.[313,1274,1128,1848]" captionTargetPageId="9" captionText="Fig. 5. Overbeckia papuana sp. nov. holotype (worker) and its lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d)." figureDoi="http://doi.org/10.5281/zenodo.6987307" httpUri="https://zenodo.org/record/6987307/files/figure.png" pageId="14" pageNumber="571">
Fig. 5
<emphasis bold="true" box="[176,188,340,364]" italics="true" pageId="14" pageNumber="571">b</emphasis>
</figureCitation>
). Approximately six erect hairs growing dorsally from apical margin of petiole.
</paragraph>
<paragraph blockId="14.[98,770,180,844]" pageId="14" pageNumber="571">
<emphasis bold="true" box="[130,232,403,427]" italics="true" pageId="14" pageNumber="571">Sculpture</emphasis>
. Less dotted, rather covered by short lines joining each other to a cell formation with fine punctation over the glossy cuticle of the head (
<figureCitation box="[507,584,468,492]" captionStart="Fig" captionStartId="9.[314,351,1878,1901]" captionTargetBox="[314,1274,1129,1848]" captionTargetId="figure-2@9.[313,1274,1128,1848]" captionTargetPageId="9" captionText="Fig. 5. Overbeckia papuana sp. nov. holotype (worker) and its lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d)." figureDoi="http://doi.org/10.5281/zenodo.6987307" httpUri="https://zenodo.org/record/6987307/files/figure.png" pageId="14" pageNumber="571">
Fig. 5
<emphasis bold="true" box="[571,584,468,492]" italics="true" pageId="14" pageNumber="571">d</emphasis>
</figureCitation>
) and mesosoma dorsolaterally, with most of these cells like pits at smaller (&lt;80×) magnification (
<figureCitation box="[381,489,532,556]" captionStart="Fig" captionStartId="9.[314,351,1878,1901]" captionTargetBox="[314,1274,1129,1848]" captionTargetId="figure-2@9.[313,1274,1128,1848]" captionTargetPageId="9" captionText="Fig. 5. Overbeckia papuana sp. nov. holotype (worker) and its lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d)." figureDoi="http://doi.org/10.5281/zenodo.6987307" httpUri="https://zenodo.org/record/6987307/files/figure.png" pageId="14" pageNumber="571">
Fig. 5
<emphasis bold="true" box="[448,460,532,556]" italics="true" pageId="14" pageNumber="571">b</emphasis>
,
<emphasis bold="true" box="[478,489,532,555]" italics="true" pageId="14" pageNumber="571">c</emphasis>
</figureCitation>
). Lateral sites of thorax with soft lineation below mesothoracic spiracle towards mid and hind coxa. The lineation is also present on frontal coxa and petiole, but smoother.
</paragraph>
<paragraph blockId="14.[98,770,180,844]" pageId="14" pageNumber="571">
<emphasis bold="true" box="[130,260,660,684]" italics="true" pageId="14" pageNumber="571">Colouration</emphasis>
. Head, mesosoma and frontal coxa black; scape dark brown except at apex, where lighter brown; funiculus of antennae brown except the lighter apex; petiole dark brown; abdomen and hind tibiae brown, remaining appendages rather light brown, with mid and hind coxae, trochanters and palps pale yellow (
<figureCitation box="[498,603,820,844]" captionStart="Fig" captionStartId="9.[314,351,1878,1901]" captionTargetBox="[314,1274,1129,1848]" captionTargetId="figure-2@9.[313,1274,1128,1848]" captionTargetPageId="9" captionText="Fig. 5. Overbeckia papuana sp. nov. holotype (worker) and its lateral (a), frontal (b) and dorsal (c) views. Detail of head sculpture and shape of frontal carinae (d)." figureDoi="http://doi.org/10.5281/zenodo.6987307" httpUri="https://zenodo.org/record/6987307/files/figure.png" pageId="14" pageNumber="571">
Fig. 5
<emphasis bold="true" box="[563,576,820,843]" italics="true" pageId="14" pageNumber="571">a</emphasis>
,
<emphasis bold="true" box="[592,603,820,843]" italics="true" pageId="14" pageNumber="571">c</emphasis>
</figureCitation>
).
</paragraph>
<paragraph blockId="14.[98,387,897,925]" box="[98,387,897,925]" pageId="14" pageNumber="571">
<heading bold="true" box="[98,387,897,925]" fontSize="11" level="3" pageId="14" pageNumber="571" reason="0">
<emphasis bold="true" box="[98,387,897,925]" pageId="14" pageNumber="571">Description of queen</emphasis>
</heading>
</paragraph>
<paragraph blockId="14.[98,770,948,1676]" pageId="14" pageNumber="571">
<emphasis bold="true" box="[98,262,948,972]" italics="true" pageId="14" pageNumber="571">Morphometrics</emphasis>
.
<typeStatus box="[294,393,949,973]" pageId="14" pageNumber="571">Paratype</typeStatus>
HP0177 nest series:
<collectionCode box="[667,699,949,972]" country="USA" lsid="urn:lsid:biocol.org:col:13245" name="Houghton Lake Wildlife Research Station" pageId="14" pageNumber="571" type="Herbarium">HL</collectionCode>
1.56, HW 1.57, IOD 1.22,
<collectionCode box="[339,368,980,1004]" country="Romania" lsid="urn:lsid:biocol.org:col:15579" name="Babes-Bolyai University" pageId="14" pageNumber="571" type="Herbarium">CL</collectionCode>
0.32, CW 0.59,
<collectionCode box="[562,598,980,1003]" country="France" name="Musee de Lectoure" pageId="14" pageNumber="571">ML</collectionCode>
0.63, EL 0.49, EW 0.37, FCDp 0.50, FCDa 0.24,
<collectionCode box="[506,533,1012,1036]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="14" pageNumber="571" type="Herbarium">SL</collectionCode>
1.23, SWmax 0.16, SWmin 0.09,
<collectionCode box="[253,290,1045,1068]" country="China" lsid="urn:lsid:biocol.org:col:14766" name="Wolong Nature Reserve" pageId="14" pageNumber="571" type="Herbarium">WL</collectionCode>
2.51,
<collectionCode box="[365,403,1045,1068]" country="Thailand" name="Paleontological Collections" pageId="14" pageNumber="571">PW</collectionCode>
1.14, HFL 1.22, HFT 0.31, HTL 1.27, PetW 0.55, PetL 0.35,
<collectionCode box="[414,470,1076,1100]" pageId="14" pageNumber="571">FCDI</collectionCode>
208,
<collectionCode box="[535,559,1076,1100]" country="USA" lsid="urn:lsid:biocol.org:col:15344" name="Carnegie Institution of Washington" pageId="14" pageNumber="571" type="Herbarium">CI</collectionCode>
101,
<collectionCode box="[624,646,1076,1100]" country="Argentina" name="Museo Botánico (SI)" pageId="14" pageNumber="571" type="Herbarium">SI</collectionCode>
78,
<collectionCode box="[696,727,1076,1099]" country="Italy" lsid="urn:lsid:biocol.org:col:14381" name="Università degli Studi di Milano" pageId="14" pageNumber="571" type="Herbarium">MI</collectionCode>
40.
</paragraph>
<paragraph blockId="14.[98,770,948,1676]" pageId="14" pageNumber="571">
<emphasis bold="true" box="[130,474,1108,1132]" italics="true" pageId="14" pageNumber="571">Head, mesosoma and metasoma</emphasis>
. Similar to
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[615,762,1108,1132]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="subclavata">
<emphasis bold="true" box="[615,762,1108,1132]" italics="true" pageId="14" pageNumber="571">O. subclavata</emphasis>
</taxonomicName>
, except frontal carinae significantly approaching one another anteriorly (
<collectionCode box="[217,274,1172,1196]" pageId="14" pageNumber="571">FCDI</collectionCode>
= 208;
<figureCitation box="[363,434,1172,1196]" captionStart="Fig" captionStartId="10.[154,191,1653,1676]" captionTargetBox="[154,1434,177,1622]" captionTargetId="figure-75@10.[154,1435,176,1623]" captionTargetPageId="10" captionText="Fig. 6. Overbeckia papuana sp. nov. paratypes (alates). Queen: lateral (a), dorsal (c) and frontal (e) views; male: lateral (b), dorsal (d) and frontal (f) views. Note that the petiole in queen (a) looks wider and blunter due to being bent to the right site (c)." figureDoi="http://doi.org/10.5281/zenodo.6987309" httpUri="https://zenodo.org/record/6987309/files/figure.png" pageId="14" pageNumber="571">
Fig. 6
<emphasis bold="true" box="[423,434,1172,1195]" italics="true" pageId="14" pageNumber="571">e</emphasis>
</figureCitation>
).
</paragraph>
<paragraph blockId="14.[98,770,948,1676]" pageId="14" pageNumber="571">
<emphasis bold="true" box="[130,210,1204,1228]" italics="true" pageId="14" pageNumber="571">Pilosity</emphasis>
. Tiny appressed hairs on scape surface, with three to four longer erect hairs on the lateral sites of the first half of the scape (hair length ≥1 ocellus); presence of many relatively long, erect hairs on the head (length ≥1 ocellus) on genae, above the eyes and over the occipital corners of the head (
<figureCitation box="[335,410,1364,1388]" captionStart="Fig" captionStartId="10.[154,191,1653,1676]" captionTargetBox="[154,1434,177,1622]" captionTargetId="figure-75@10.[154,1435,176,1623]" captionTargetPageId="10" captionText="Fig. 6. Overbeckia papuana sp. nov. paratypes (alates). Queen: lateral (a), dorsal (c) and frontal (e) views; male: lateral (b), dorsal (d) and frontal (f) views. Note that the petiole in queen (a) looks wider and blunter due to being bent to the right site (c)." figureDoi="http://doi.org/10.5281/zenodo.6987309" httpUri="https://zenodo.org/record/6987309/files/figure.png" pageId="14" pageNumber="571">
Fig. 6
<emphasis bold="true" box="[399,410,1364,1387]" italics="true" pageId="14" pageNumber="571">e</emphasis>
</figureCitation>
), and on thorax dorsum, apical tip of petiole, and over both ventral and dorsal sites of all five abdominal tergites.
</paragraph>
<paragraph blockId="14.[98,770,948,1676]" pageId="14" pageNumber="571">
<emphasis bold="true" box="[130,232,1459,1483]" italics="true" pageId="14" pageNumber="571">Sculpture</emphasis>
. Fine punctation over the head, and on mesosoma dorsum and propodeum; and soft lineation on lateral sites of mesosoma (
<figureCitation box="[319,397,1524,1548]" captionStart="Fig" captionStartId="10.[154,191,1653,1676]" captionTargetBox="[154,1434,177,1622]" captionTargetId="figure-75@10.[154,1435,176,1623]" captionTargetPageId="10" captionText="Fig. 6. Overbeckia papuana sp. nov. paratypes (alates). Queen: lateral (a), dorsal (c) and frontal (e) views; male: lateral (b), dorsal (d) and frontal (f) views. Note that the petiole in queen (a) looks wider and blunter due to being bent to the right site (c)." figureDoi="http://doi.org/10.5281/zenodo.6987309" httpUri="https://zenodo.org/record/6987309/files/figure.png" pageId="14" pageNumber="571">
Fig. 6
<emphasis bold="true" box="[384,397,1524,1547]" italics="true" pageId="14" pageNumber="571">a</emphasis>
</figureCitation>
).
</paragraph>
<paragraph blockId="14.[98,770,948,1676]" pageId="14" pageNumber="571">
<emphasis bold="true" box="[130,260,1555,1579]" italics="true" pageId="14" pageNumber="571">Colouration</emphasis>
. Black head, thorax, petiole and frontal coxae; dark brown abdomen and appendages except middle and hind coxae, trochanters, palps and apical tips of antennae and tarsi that are lighter brown (
<figureCitation box="[522,600,1652,1676]" captionStart="Fig" captionStartId="10.[154,191,1653,1676]" captionTargetBox="[154,1434,177,1622]" captionTargetId="figure-75@10.[154,1435,176,1623]" captionTargetPageId="10" captionText="Fig. 6. Overbeckia papuana sp. nov. paratypes (alates). Queen: lateral (a), dorsal (c) and frontal (e) views; male: lateral (b), dorsal (d) and frontal (f) views. Note that the petiole in queen (a) looks wider and blunter due to being bent to the right site (c)." figureDoi="http://doi.org/10.5281/zenodo.6987309" httpUri="https://zenodo.org/record/6987309/files/figure.png" pageId="14" pageNumber="571">
Fig. 6
<emphasis bold="true" box="[587,600,1652,1675]" italics="true" pageId="14" pageNumber="571">a</emphasis>
</figureCitation>
).
</paragraph>
<paragraph blockId="14.[98,371,1729,1757]" box="[98,371,1729,1757]" pageId="14" pageNumber="571">
<heading bold="true" box="[98,371,1729,1757]" fontSize="11" level="3" pageId="14" pageNumber="571" reason="0">
<emphasis bold="true" box="[98,371,1729,1757]" pageId="14" pageNumber="571">Description of male</emphasis>
</heading>
</paragraph>
<paragraph blockId="14.[98,770,1780,1932]" pageId="14" pageNumber="571">
<emphasis bold="true" box="[98,262,1780,1804]" italics="true" pageId="14" pageNumber="571">Morphometrics</emphasis>
.
<typeStatus box="[294,393,1780,1804]" pageId="14" pageNumber="571">Paratype</typeStatus>
HP0177 nest series:
<collectionCode box="[667,699,1780,1803]" country="USA" lsid="urn:lsid:biocol.org:col:13245" name="Houghton Lake Wildlife Research Station" pageId="14" pageNumber="571" type="Herbarium">HL</collectionCode>
0.84, HW 1.00, IOD 0.66,
<collectionCode box="[339,368,1812,1836]" country="Romania" lsid="urn:lsid:biocol.org:col:15579" name="Babes-Bolyai University" pageId="14" pageNumber="571" type="Herbarium">CL</collectionCode>
0.12, CW 0.35,
<collectionCode box="[562,598,1813,1836]" country="France" name="Musee de Lectoure" pageId="14" pageNumber="571">ML</collectionCode>
0.31, EL 0.34, EW 0.29, FCDp 0.27, FCDa 0.10,
<collectionCode box="[506,533,1844,1868]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="14" pageNumber="571" type="Herbarium">SL</collectionCode>
0.67, SWmax 0.10, SWmin 0.06,
<collectionCode box="[253,290,1876,1899]" country="China" lsid="urn:lsid:biocol.org:col:14766" name="Wolong Nature Reserve" pageId="14" pageNumber="571" type="Herbarium">WL</collectionCode>
1.71,
<collectionCode box="[365,403,1876,1899]" country="Thailand" name="Paleontological Collections" pageId="14" pageNumber="571">PW</collectionCode>
0.96, HFL 1.31, HFT 0.18, HTL 0.99, PetW 0.33, PetL 0.21,
<collectionCode box="[414,470,1908,1932]" pageId="14" pageNumber="571">FCDI</collectionCode>
255,
<collectionCode box="[535,559,1908,1932]" country="USA" lsid="urn:lsid:biocol.org:col:15344" name="Carnegie Institution of Washington" pageId="14" pageNumber="571" type="Herbarium">CI</collectionCode>
119,
<collectionCode box="[624,646,1908,1932]" country="Argentina" name="Museo Botánico (SI)" pageId="14" pageNumber="571" type="Herbarium">SI</collectionCode>
67,
<collectionCode box="[696,727,1909,1932]" country="Italy" lsid="urn:lsid:biocol.org:col:14381" name="Università degli Studi di Milano" pageId="14" pageNumber="571" type="Herbarium">MI</collectionCode>
37.
</paragraph>
<paragraph blockId="14.[818,1490,180,652]" pageId="14" pageNumber="571">
Structures and diagnosis (
<figureCitation box="[1158,1302,180,204]" captionStart="Fig" captionStartId="10.[154,191,1653,1676]" captionTargetBox="[154,1434,177,1622]" captionTargetId="figure-75@10.[154,1435,176,1623]" captionTargetPageId="10" captionText="Fig. 6. Overbeckia papuana sp. nov. paratypes (alates). Queen: lateral (a), dorsal (c) and frontal (e) views; male: lateral (b), dorsal (d) and frontal (f) views. Note that the petiole in queen (a) looks wider and blunter due to being bent to the right site (c)." figureDoi="http://doi.org/10.5281/zenodo.6987309" httpUri="https://zenodo.org/record/6987309/files/figure.png" pageId="14" pageNumber="571">
Fig. 6
<emphasis bold="true" box="[1227,1239,180,204]" italics="true" pageId="14" pageNumber="571">b</emphasis>
,
<emphasis bold="true" box="[1260,1302,180,204]" italics="true" pageId="14" pageNumber="571">d, f</emphasis>
</figureCitation>
). No apparent large distinction when compared to the male of
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1333,1482,211,236]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="subclavata">
<emphasis bold="true" box="[1333,1482,211,236]" italics="true" pageId="14" pageNumber="571">O. subclavata</emphasis>
</taxonomicName>
, except for the following minor differences: The specimen is ~10% larger than the measured male of
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1284,1436,276,300]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="subclavata">
<emphasis bold="true" box="[1284,1436,276,300]" italics="true" pageId="14" pageNumber="571">O. subclavata</emphasis>
</taxonomicName>
, but morphological measures otherwise considerably overlapping between the two species males. Frontal carinae relatively more approaching one another anteriorly than in
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1290,1439,372,396]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="subclavata">
<emphasis bold="true" box="[1290,1439,372,396]" italics="true" pageId="14" pageNumber="571">O. subclavata</emphasis>
</taxonomicName>
, but this difference not as distinct as between the two species workers (
<collectionCode box="[926,984,436,460]" pageId="14" pageNumber="571">FCDI</collectionCode>
= 255
<emphasis bold="true" box="[1074,1086,436,459]" italics="true" pageId="14" pageNumber="571">v</emphasis>
.
<quantity box="[1106,1183,436,460]" metricMagnitude="0" metricUnit="m" metricValue="5.816599999999999" pageId="14" pageNumber="571" unit="in" value="229.0">229 in</quantity>
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1196,1351,436,460]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="subclavata">
<emphasis bold="true" box="[1196,1351,436,460]" italics="true" pageId="14" pageNumber="571">O. subclavata</emphasis>
</taxonomicName>
male). The hairs on clypeus more numerous than on clypeus of
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="subclavata">
<emphasis bold="true" italics="true" pageId="14" pageNumber="571">O. subclavata</emphasis>
</taxonomicName>
; presence of many more small hairs on frons and above ocelli (length near 1/2 of ocellus) over the occipital region (
<figureCitation box="[909,984,564,588]" captionStart="Fig" captionStartId="10.[154,191,1653,1676]" captionTargetBox="[154,1434,177,1622]" captionTargetId="figure-75@10.[154,1435,176,1623]" captionTargetPageId="10" captionText="Fig. 6. Overbeckia papuana sp. nov. paratypes (alates). Queen: lateral (a), dorsal (c) and frontal (e) views; male: lateral (b), dorsal (d) and frontal (f) views. Note that the petiole in queen (a) looks wider and blunter due to being bent to the right site (c)." figureDoi="http://doi.org/10.5281/zenodo.6987309" httpUri="https://zenodo.org/record/6987309/files/figure.png" pageId="14" pageNumber="571">
Fig. 6
<emphasis bold="true" box="[976,984,564,588]" italics="true" pageId="14" pageNumber="571">f</emphasis>
</figureCitation>
). These small, dense hairs are also present dorsolaterally on thorax (on mesoscutum and mesoscutellar disc;
<figureCitation box="[875,978,628,652]" captionStart="Fig" captionStartId="10.[154,191,1653,1676]" captionTargetBox="[154,1434,177,1622]" captionTargetId="figure-75@10.[154,1435,176,1623]" captionTargetPageId="10" captionText="Fig. 6. Overbeckia papuana sp. nov. paratypes (alates). Queen: lateral (a), dorsal (c) and frontal (e) views; male: lateral (b), dorsal (d) and frontal (f) views. Note that the petiole in queen (a) looks wider and blunter due to being bent to the right site (c)." figureDoi="http://doi.org/10.5281/zenodo.6987309" httpUri="https://zenodo.org/record/6987309/files/figure.png" pageId="14" pageNumber="571">
Fig. 6
<emphasis bold="true" box="[938,950,628,652]" italics="true" pageId="14" pageNumber="571">b</emphasis>
,
<emphasis bold="true" box="[965,978,628,652]" italics="true" pageId="14" pageNumber="571">d</emphasis>
</figureCitation>
). Colouration as in
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1199,1348,628,652]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="subclavata">
<emphasis bold="true" box="[1199,1348,628,652]" italics="true" pageId="14" pageNumber="571">O. subclavata</emphasis>
</taxonomicName>
male.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="571" type="distribution">
<paragraph blockId="14.[818,940,695,723]" box="[818,940,695,723]" pageId="14" pageNumber="571">
<heading bold="true" box="[818,940,695,723]" fontSize="11" level="3" pageId="14" pageNumber="571" reason="0">
<emphasis bold="true" box="[818,940,695,723]" pageId="14" pageNumber="571">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="14.[818,1490,747,1475]" pageId="14" pageNumber="571">
The arboreal nest of
<taxonomicName authority="Klimeš &amp; Drescher &amp; Buchori &amp; Hidayat &amp; Nazarreta &amp; Potocký &amp; Rimandai &amp; Scheu &amp; Matos-Maraví, 2022" authorityName="Klimeš &amp; Drescher &amp; Buchori &amp; Hidayat &amp; Nazarreta &amp; Potocký &amp; Rimandai &amp; Scheu &amp; Matos-Maraví" authorityYear="2022" box="[1037,1162,747,771]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="papuana" status="sp. nov.">
<emphasis bold="true" box="[1037,1162,747,771]" italics="true" pageId="14" pageNumber="571">O. papuana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1168,1252,747,770]" pageId="14" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
in
<collectingCountry box="[1283,1490,747,771]" name="Papua New Guinea" pageId="14" pageNumber="571">Papua New Guinea</collectingCountry>
was collected at
<quantity box="[999,1075,779,803]" metricMagnitude="1" metricUnit="m" metricValue="1.64" pageId="14" pageNumber="571" unit="m" value="16.4">16.4 m</quantity>
in a dead vine growing along the tree trunk of
<taxonomicName authority="(F. Muell.) Kosterm." authorityName="Kosterm." baseAuthorityName="F. Muell." box="[929,1490,810,835]" class="Magnoliopsida" family="Malvaceae" genus="Trichospermum" kingdom="Plantae" order="Malvales" pageId="14" pageNumber="571" phylum="Tracheophyta" rank="species" species="pleiostigma">
<emphasis bold="true" box="[929,1237,810,834]" italics="true" pageId="14" pageNumber="571">Trichospermum pleiostigma</emphasis>
(F. Muell.) Kosterm.
</taxonomicName>
(diameter at breast height =
<quantity box="[1144,1236,843,867]" metricMagnitude="-1" metricUnit="m" metricValue="1.63" pageId="14" pageNumber="571" unit="cm" value="16.3">16.3 cm</quantity>
). The nest consisted of less than one hundred workers, with one wingless gyne,
<specimenCount pageId="14" pageNumber="571" type="male">three males</specimenCount>
, and multiple larvae and pupae enclosed in cocoons.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="571" type="materials_examined">
<paragraph blockId="14.[818,1490,747,1475]" pageId="14" pageNumber="571">
The material from
<collectingCountry box="[1060,1161,939,963]" name="Australia" pageId="14" pageNumber="571">Australia</collectingCountry>
was examined and measured mainly from photographs from the following sources: (i) the book by
<bibRefCitation author="McArthur A" box="[923,1119,1003,1027]" pageId="14" pageNumber="571" refId="ref18953" refString="McArthur A (2012) ' A guide to Colobopsis ants of the world. ' (South Australian Museum: Adelaide, SA, Australia)" type="book" year="2012">McArthur (2012)</bibRefCitation>
, where a worker of
<taxonomicName authorityName="Viehmeyer" authorityYear="1916" box="[1368,1490,1002,1026]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1368,1490,1002,1026]" italics="true" pageId="14" pageNumber="571">Overbeckia</emphasis>
</taxonomicName>
was incorrectly assigned to
<taxonomicName authorityName="Forel" authorityYear="1895" box="[1176,1393,1035,1059]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="janeti">
<emphasis bold="true" box="[1176,1393,1035,1059]" italics="true" pageId="14" pageNumber="571">Camponotus janeti</emphasis>
</taxonomicName>
(minor worker fits
<emphasis bold="true" box="[952,1086,1066,1090]" italics="true" pageId="14" pageNumber="571">
<collectionCode box="[952,973,1066,1090]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="14" pageNumber="571" type="Herbarium">O</collectionCode>
. papuana
</emphasis>
, whereas major worker presumably fits
<taxonomicName authorityName="Forel" authorityYear="1895" box="[865,962,1099,1123]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="janeti">
<emphasis bold="true" box="[865,962,1099,1123]" italics="true" pageId="14" pageNumber="571">C. janeti</emphasis>
</taxonomicName>
; there is uncertainty as to whether the two individuals are from different localities and pins or are on a single pin at
<collectionCode box="[977,1053,1163,1187]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="14" pageNumber="571" type="Museum">SAMA</collectionCode>
); (ii) the photos on AntWeb of the
<specimenCount pageId="14" pageNumber="571" type="generic">three specimens</specimenCount>
in
<bibRefCitation author="Heterick BE" box="[977,1159,1195,1219]" pageId="14" pageNumber="571" pagination="163 - 166" refId="ref18158" refString="Heterick BE (2019) First record of the formicine genus Overbeckia (Hymenoptera: Formicidae) from Australia. Myrmecological News 29, 163 - 166." type="journal article" year="2019">Heterick (2019)</bibRefCitation>
(see above). The Australian specimens appear to be larger in body size based on the measures from the photos (
<specimenCount box="[1188,1304,1259,1283]" pageId="14" pageNumber="571" type="worker">4 workers</specimenCount>
measured from photos,
<collectionCode box="[908,945,1291,1314]" country="China" lsid="urn:lsid:biocol.org:col:14766" name="Wolong Nature Reserve" pageId="14" pageNumber="571" type="Herbarium">WL</collectionCode>
&gt;
<quantity box="[980,1080,1290,1315]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="14" pageNumber="571" unit="mm" value="1.95">1.95 mm</quantity>
) than the
<collectingCountry box="[1197,1412,1291,1315]" name="Papua New Guinea" pageId="14" pageNumber="571">Papua New Guinea</collectingCountry>
(
<collectingCountry box="[1427,1482,1291,1315]" name="Papua New Guinea" pageId="14" pageNumber="571">PNG</collectingCountry>
) nest series (6w measured,
<collectionCode box="[1127,1164,1323,1346]" country="China" lsid="urn:lsid:biocol.org:col:14766" name="Wolong Nature Reserve" pageId="14" pageNumber="571" type="Herbarium">WL</collectionCode>
&lt;1.74). However, we treat these as one species because the Australian single specimen sent by
<collectionCode box="[936,954,1387,1410]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="14" pageNumber="571" type="Herbarium">P</collectionCode>
. Ward for direct examination (n#15720) matches
<taxonomicName authorityName="Klimeš &amp; Drescher &amp; Buchori &amp; Hidayat &amp; Nazarreta &amp; Potocký &amp; Rimandai &amp; Scheu &amp; Matos-Maraví" authorityYear="2022" box="[923,1056,1419,1443]" class="Insecta" family="Formicidae" genus="Overbeckia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="571" phylum="Arthropoda" rank="species" species="papuana">
<emphasis bold="true" box="[923,1056,1419,1443]" italics="true" pageId="14" pageNumber="571">O. papuana</emphasis>
</taxonomicName>
<typeStatus box="[1071,1130,1419,1443]" pageId="14" pageNumber="571">types</typeStatus>
in the body size (
<collectionCode box="[1357,1397,1419,1443]" country="China" lsid="urn:lsid:biocol.org:col:14766" name="Wolong Nature Reserve" pageId="14" pageNumber="571" type="Herbarium">WL</collectionCode>
= 1.64; Supplementary
<tableCitation box="[995,1094,1451,1475]" captionStart="Table 1" captionStartId="5.[98,162,181,204]" captionTargetPageId="5" captionText="Table 1. Estimated crown group divergence ages of the tribe Camponotini and the seven non-monotypic genera." httpUri="http://table.plazi.org/id/DF33088B65180533FFDEFF439BC22BA1" pageId="14" pageNumber="571" tableUuid="DF33088B65180533FFDEFF439BC22BA1">Table S1</tableCitation>
) and morphology.
</paragraph>
</subSubSection>
</treatment>
</document>