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<document id="1C858C737C6993284732016CAA9E51DD" ID-DOI="10.11646/zootaxa.3637.1.5" ID-GBIF-Dataset="ef40fc0f-2189-4ae6-8d9d-caa4e0bb1bbe" ID-ISSN="1175-5326" ID-Zenodo-Dep="283594" ID-ZooBank="F642CEB2-2DCB-4B25-BA8E-A58657767044" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460590613857" checkinUser="plazi" docAuthor="Yin, Zi-Wei, Nomura, Shûhei &amp; Li, Li-Zhen" docDate="2013" docId="03E387E42017FFEAFF67DA1D3054F803" docLanguage="en" docName="zt03637p057.pdf" docOrigin="Zootaxa 3637 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrus Aube 1833" docType="treatment" docVersion="6" lastPageNumber="54" masterDocId="FFDAFF9C2016FFEDFFF0DB703322FF8A" masterDocTitle="Redescription of Tyrus Aubé and T. sinensi s Raffray (Coleoptera: Staphylinidae: Pselaphinae), and two new species from Sichuan, Southwest China" masterLastPageNumber="57" masterPageNumber="47" pageNumber="48" updateTime="1698325835414" updateUser="plazi">
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<mods:title id="4D4B9038DB5C4DF18110B39CAA6CB3E4">Redescription of Tyrus Aubé and T. sinensi s Raffray (Coleoptera: Staphylinidae: Pselaphinae), and two new species from Sichuan, Southwest China</mods:title>
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<mods:namePart id="6B86401DA1A35A74840B6A3C8D388C9C">Yin, Zi-Wei</mods:namePart>
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<mods:namePart id="0229F4E571783C0D5388174D6DDC2C09">Nomura, Shûhei</mods:namePart>
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<mods:namePart id="53AB40A2508662824BBF95E1D2ABA8FD">Li, Li-Zhen</mods:namePart>
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<treatment id="03E387E42017FFEAFF67DA1D3054F803" ID-DOI="http://doi.org/10.5281/zenodo.6145968" ID-GBIF-Taxon="127663756" ID-Zenodo-Dep="6145968" LSID="urn:lsid:plazi:treatment:03E387E42017FFEAFF67DA1D3054F803" httpUri="http://treatment.plazi.org/id/03E387E42017FFEAFF67DA1D3054F803" lastPageId="7" lastPageNumber="54" pageId="1" pageNumber="48">
<subSubSection id="C35065792017FFECFF67DA1D318EFD86" pageId="1" pageNumber="48" type="nomenclature">
<paragraph id="8BF536F22017FFECFF67DA1D32E4FE0D" blockId="1.[151,454,365,423]" box="[151,454,365,391]" pageId="1" pageNumber="48">
<heading id="D0BD819E2017FFECFF67DA1D32E4FE0D" bold="true" box="[151,454,365,391]" fontSize="11" level="1" pageId="1" pageNumber="48" reason="1">
<emphasis id="B93EEAE02017FFECFF67DA1D32E4FE0D" bold="true" box="[151,454,365,391]" pageId="1" pageNumber="48">
Genus
<taxonomicName id="4C4A4D712017FFECFF1EDA1D32E4FE0D" ID-CoL="8VS6F" authority="Aube, 1833" authorityName="Aube" authorityYear="1833" box="[238,454,365,391]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02017FFECFF1EDA1D3210FE0D" bold="true" box="[238,306,365,391]" italics="true" pageId="1" pageNumber="48">Tyrus</emphasis>
Aubé, 1833
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF536F22017FFECFF67DAFF3229FE2D" blockId="1.[151,454,365,423]" box="[151,267,399,423]" pageId="1" pageNumber="48">
(
<figureCitation id="13712A772017FFECFF6FDAFF3220FE2D" box="[159,258,399,423]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="1.[151,250,1960,1983]" captionStartId-1="2.[151,250,1741,1764]" captionStartId-2="3.[151,250,1511,1534]" captionStartId-3="5.[151,250,1321,1344]" captionStartId-4="6.[151,250,1849,1872]" captionStartId-5="8.[151,250,1637,1660]" captionTargetBox-0="[180,1402,786,1918]" captionTargetBox-1="[151,1425,195,1685]" captionTargetBox-2="[165,1425,200,1463]" captionTargetBox-3="[156,1426,199,1295]" captionTargetBox-4="[159,1430,457,1819]" captionTargetBox-5="[166,1419,205,1606]" captionTargetId-0="figure@1.[171,1408,769,1924]" captionTargetId-1="figure@2.[151,1436,195,1720]" captionTargetId-2="figure@3.[151,1436,193,1490]" captionTargetId-3="figure@5.[151,1436,193,1300]" captionTargetId-4="figure@6.[151,1436,447,1827]" captionTargetId-5="figure@8.[151,1436,195,1616]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="8" captionText-0="FIGURE 1. Dorsal habitus of Tyrus sichuanicus (A) and T. sinensis (B). Scales: 0.5 mm." captionText-1="FIGURE 2. Diagnostic features of Tyrus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Pronotum. E. Proventrite. F. Left elytron. Scales: 0.2 mm. Abbreviations: as—antebasal sulcus, at—antennal tubercle, bef—basal elytral foveae, ds—discal stria, ff—frontal fovea, gf—gular foveae, laf—lateral antebasal foveae, lpcf—lateral procoxal foveae, lpp—lateral postantennal pits, maf—median antebasal foveae, oc—ocular canthi, omc—ocular-mandibular carinae, sef—subbasal elytral foveae, ss—sutural stria, vf—vertexal foveae." captionText-2="FIGURE 3. Diagnostic features of Tyrus. A. Meso- and metaventrite. B. Abdomen, in lateral view. C. Same, in dorsal view. D. Same, in ventral view. Scales: 0.2 mm. Abbreviations: blf—basolateral foveae, bs—basal sulcus, lmcf—lateral mesocoxal foveae, lmvf—lateral mesoventral foveae, mbf—mediobasal foveae, mdc—median discal carina, mmsf—median mesoventral foveae, mmtf—median metaventral fovea, msff—mesofurcal foveae." captionText-3="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." captionText-4="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." captionText-5="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri-0="https://zenodo.org/record/283595/files/figure.png" httpUri-1="https://zenodo.org/record/283596/files/figure.png" httpUri-2="https://zenodo.org/record/283597/files/figure.png" httpUri-3="https://zenodo.org/record/283598/files/figure.png" httpUri-4="https://zenodo.org/record/283599/files/figure.png" httpUri-5="https://zenodo.org/record/283600/files/figure.png" pageId="1" pageNumber="48">Figs 16</figureCitation>
)
</paragraph>
<paragraph id="8BF536F22017FFECFF67DAA9318EFD86" blockId="1.[151,1056,472,524]" pageId="1" pageNumber="48">
<taxonomicName id="4C4A4D712017FFECFF67DAA93256FE64" authority="Aube, 1833: 15" authorityName="Aube" authorityPageNumber="15" authorityYear="1833" box="[151,372,472,494]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02017FFECFF67DAA933EFFE64" box="[151,205,473,494]" italics="true" pageId="1" pageNumber="48">Tyrus</emphasis>
Aubé, 1833: 15
</taxonomicName>
.
<typeStatus id="54F188502017FFECFE8EDAA93292FE64" box="[382,432,473,494]" pageId="1" pageNumber="48">Type</typeStatus>
species:
<taxonomicName id="4C4A4D712017FFECFDFDDAA8305EFE67" authority="Panzer, 1803" authorityName="Panzer" authorityYear="1803" box="[525,892,472,494]" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B93EEAE02017FFECFDFDDAA831D0FE64" box="[525,754,472,494]" italics="true" pageId="1" pageNumber="48">Pselaphus mucronatus</emphasis>
Panzer, 1803
</taxonomicName>
(by monotypy). (list of synonyms see Hlaváč &amp; Chandler 2005: 116)
</paragraph>
</subSubSection>
<subSubSection id="C35065792017FFE9FF67D9483122FE7E" lastPageId="4" lastPageNumber="51" pageId="1" pageNumber="48" type="diagnosis">
<paragraph id="8BF536F22017FFECFF67D94832A1FD13" blockId="1.[151,1436,568,737]" pageId="1" pageNumber="48">
<emphasis id="B93EEAE02017FFECFF67D9483233FDDB" bold="true" box="[151,273,568,593]" pageId="1" pageNumber="48">Diagnosis.</emphasis>
Head with frontal and vertexal foveae; maxillary palpomeres IIIV basally pedunculate; pronotum with antebasal sulcus connecting median and lateral antebasal foveae; abdomen with tergite IV as long as to slightly longer than tergite V.
</paragraph>
<paragraph id="8BF536F22017FFECFF39D9D53238FD6B" blockId="1.[151,1436,568,737]" pageId="1" pageNumber="48">
<emphasis id="B93EEAE02017FFECFF39D9D53254FD34" bold="true" box="[201,374,677,702]" pageId="1" pageNumber="48">Redescription.</emphasis>
Length
<quantity id="4CB29B172017FFECFE25D9D5317CFD37" box="[469,606,677,701]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.7" metricValueMin="1.5" pageId="1" pageNumber="48" unit="mm" value="2.1" valueMax="2.7" valueMin="1.5">1.52.7 mm</quantity>
(minimal size taken from Blattný 1925: 217; Bryant 1915: 299). Habitus as in
<figureCitation id="13712A772017FFECFF22D9B93236FD6B" box="[210,276,713,737]" captionStart="FIGURE 1" captionStartId="1.[151,250,1960,1983]" captionTargetBox="[180,1402,786,1918]" captionTargetId="figure@1.[171,1408,769,1924]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Tyrus sichuanicus (A) and T. sinensis (B). Scales: 0.5 mm." httpUri="https://zenodo.org/record/283595/files/figure.png" pageId="1" pageNumber="48">Fig. 1</figureCitation>
.
</paragraph>
<caption id="DF35667A2017FFECFF67DCD8373CF834" box="[151,1054,1960,1983]" httpUri="https://zenodo.org/record/283595/files/figure.png" pageId="1" pageNumber="48" targetBox="[180,1402,786,1918]" targetPageId="1">
<paragraph id="8BF536F22017FFECFF67DCD8373CF834" blockId="1.[151,1054,1960,1983]" box="[151,1054,1960,1983]" pageId="1" pageNumber="48">
<emphasis id="B93EEAE02017FFECFF67DCD83231F834" bold="true" box="[151,275,1960,1983]" pageId="1" pageNumber="48">FIGURE 1.</emphasis>
Dorsal habitus of
<taxonomicName id="4C4A4D712017FFECFE3DDCDA31A1F834" box="[461,643,1960,1983]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="sichuanicus">
<emphasis id="B93EEAE02017FFECFE3DDCDA31A1F834" box="[461,643,1960,1983]" italics="true" pageId="1" pageNumber="48">Tyrus sichuanicus</emphasis>
</taxonomicName>
(A) and
<taxonomicName id="4C4A4D712017FFECFD2EDCDA3067F835" box="[734,837,1962,1983]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B93EEAE02017FFECFD2EDCDA3067F835" box="[734,837,1962,1983]" italics="true" pageId="1" pageNumber="48">T. sinensis</emphasis>
</taxonomicName>
(B). Scales: 0.5 mm.
</paragraph>
</caption>
<caption id="DF35667A2014FFEFFF67DDBD30B8F8D4" httpUri="https://zenodo.org/record/283596/files/figure.png" pageId="2" pageNumber="49" targetBox="[151,1425,195,1685]" targetPageId="2">
<paragraph id="8BF536F22014FFEFFF67DDBD30B8F8D4" blockId="2.[151,1436,1741,1886]" pageId="2" pageNumber="49">
<emphasis id="B93EEAE02014FFEFFF67DDBD3234F969" bold="true" box="[151,278,1741,1764]" pageId="2" pageNumber="49">FIGURE 2.</emphasis>
Diagnostic features of
<taxonomicName id="4C4A4D712014FFEFFDFEDDBF3166F96E" box="[526,580,1743,1764]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02014FFEFFDFEDDBF3166F96E" box="[526,580,1743,1764]" italics="true" pageId="2" pageNumber="49">Tyrus</emphasis>
</taxonomicName>
.
<emphasis id="B93EEAE02014FFEFFDA5DDBE314CF969" bold="true" box="[597,622,1742,1763]" pageId="2" pageNumber="49">A.</emphasis>
Head, in dorsal view.
<emphasis id="B93EEAE02014FFEFFC90DDBE3055F969" bold="true" box="[864,887,1742,1763]" pageId="2" pageNumber="49">B.</emphasis>
Same, in lateral view.
<emphasis id="B93EEAE02014FFEFFB9DDDBD37A4F96E" bold="true" box="[1133,1158,1741,1764]" pageId="2" pageNumber="49">C.</emphasis>
Same, in ventral view.
<emphasis id="B93EEAE02014FFEFFA73DDBE36BEF969" bold="true" box="[1411,1436,1742,1763]" pageId="2" pageNumber="49">D.</emphasis>
Pronotum.
<emphasis id="B93EEAE02014FFEFFEFEDD9D3207F888" bold="true" box="[270,293,1773,1794]" pageId="2" pageNumber="49">E.</emphasis>
Proventrite.
<emphasis id="B93EEAE02014FFEFFE46DD9D32EAF888" bold="true" box="[438,456,1773,1794]" pageId="2" pageNumber="49">F.</emphasis>
Left elytron. Scales: 0.2 mm. Abbreviations: as—antebasal sulcus, at—antennal tubercle, bef—basal elytral foveae, ds—discal stria, ff—frontal fovea, gf—gular foveae, laf—lateral antebasal foveae, lpcf—lateral procoxal foveae, lpp—lateral postantennal pits, maf—median antebasal foveae, oc—ocular canthi, omc—ocular-mandibular carinae, sef—subbasal elytral foveae, ss—sutural stria, vf—vertexal foveae.
</paragraph>
</caption>
<caption id="DF35667A2015FFEEFF67DE973076F9D3" httpUri="https://zenodo.org/record/283597/files/figure.png" pageId="3" pageNumber="50" targetBox="[165,1425,200,1463]" targetPageId="3">
<paragraph id="8BF536F22015FFEEFF67DE973076F9D3" blockId="3.[151,1436,1511,1625]" pageId="3" pageNumber="50">
<emphasis id="B93EEAE02015FFEEFF67DE973230FA77" bold="true" box="[151,274,1511,1534]" pageId="3" pageNumber="50">FIGURE 3.</emphasis>
Diagnostic features of
<taxonomicName id="4C4A4D712015FFEEFE0EDE993116FA74" box="[510,564,1513,1534]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02015FFEEFE0EDE993116FA74" box="[510,564,1513,1534]" italics="true" pageId="3" pageNumber="50">Tyrus</emphasis>
</taxonomicName>
.
<emphasis id="B93EEAE02015FFEEFDB2DE983179FA77" bold="true" box="[578,603,1512,1533]" pageId="3" pageNumber="50">A.</emphasis>
Meso- and metaventrite.
<emphasis id="B93EEAE02015FFEEFCADDE983056FA77" bold="true" box="[861,884,1512,1533]" pageId="3" pageNumber="50">B.</emphasis>
Abdomen, in lateral view.
<emphasis id="B93EEAE02015FFEEFB76DE9737BDFA74" bold="true" box="[1158,1183,1511,1534]" pageId="3" pageNumber="50">C.</emphasis>
Same, in dorsal view.
<emphasis id="B93EEAE02015FFEEFA73DE9836BEFA77" bold="true" box="[1411,1436,1512,1533]" pageId="3" pageNumber="50">D.</emphasis>
Same, in ventral view. Scales: 0.2 mm. Abbreviations: blf—basolateral foveae, bs—basal sulcus, lmcf—lateral mesocoxal foveae, lmvf—lateral mesoventral foveae, mbf—mediobasal foveae, mdc—median discal carina, mmsf—median mesoventral foveae, mmtf—median metaventral fovea, msff—mesofurcal foveae.
</paragraph>
</caption>
<paragraph id="8BF536F22015FFEEFF39DDF133DBF8FB" blockId="3.[151,1436,1665,2013]" pageId="3" pageNumber="50">
Head (
<figureCitation id="13712A772015FFEEFEE6DDF1327EF913" box="[278,348,1665,1689]" captionStart="FIGURE 2" captionStartId="2.[151,250,1741,1764]" captionTargetBox="[151,1425,195,1685]" captionTargetId="figure@2.[151,1436,195,1720]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tyrus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Pronotum. E. Proventrite. F. Left elytron. Scales: 0.2 mm. Abbreviations: as—antebasal sulcus, at—antennal tubercle, bef—basal elytral foveae, ds—discal stria, ff—frontal fovea, gf—gular foveae, laf—lateral antebasal foveae, lpcf—lateral procoxal foveae, lpp—lateral postantennal pits, maf—median antebasal foveae, oc—ocular canthi, omc—ocular-mandibular carinae, sef—subbasal elytral foveae, ss—sutural stria, vf—vertexal foveae." httpUri="https://zenodo.org/record/283596/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
A) with short frontal rostrum, antennal tubercles moderately prominent and close; with setose frontal and vertexal foveae; with eleven antennomeres, clubs formed by apical three antennomeres; lacking postantennal notches at lateral margins; with indistinct lateral postantennal pits (
<figureCitation id="13712A772015FFEEFBA4DDB937BDF96B" box="[1108,1183,1737,1761]" captionStart="FIGURE 2" captionStartId="2.[151,250,1741,1764]" captionTargetBox="[151,1425,195,1685]" captionTargetId="figure@2.[151,1436,195,1720]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tyrus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Pronotum. E. Proventrite. F. Left elytron. Scales: 0.2 mm. Abbreviations: as—antebasal sulcus, at—antennal tubercle, bef—basal elytral foveae, ds—discal stria, ff—frontal fovea, gf—gular foveae, laf—lateral antebasal foveae, lpcf—lateral procoxal foveae, lpp—lateral postantennal pits, maf—median antebasal foveae, oc—ocular canthi, omc—ocular-mandibular carinae, sef—subbasal elytral foveae, ss—sutural stria, vf—vertexal foveae." httpUri="https://zenodo.org/record/283596/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
B); ocular-mandibular carinae present; ocular canthi variably present; maxillary palpi (
<figureCitation id="13712A772015FFEEFC69DD9D30CAF88F" box="[921,1000,1773,1797]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="3" pageNumber="50">Figs 4</figureCitation>
D, 5D) with palpomeres II lengthily pedunculate, III and IV briefly pedunculate at bases, III triangular, IV elongately oval to fusiform, apices broadly rounded, with distinct palpal cone; gula (
<figureCitation id="13712A772015FFEEFD97DC453189F8C7" box="[615,683,1845,1869]" captionStart="FIGURE 2" captionStartId="2.[151,250,1741,1764]" captionTargetBox="[151,1425,195,1685]" captionTargetId="figure@2.[151,1436,195,1720]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tyrus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Pronotum. E. Proventrite. F. Left elytron. Scales: 0.2 mm. Abbreviations: as—antebasal sulcus, at—antennal tubercle, bef—basal elytral foveae, ds—discal stria, ff—frontal fovea, gf—gular foveae, laf—lateral antebasal foveae, lpcf—lateral procoxal foveae, lpp—lateral postantennal pits, maf—median antebasal foveae, oc—ocular canthi, omc—ocular-mandibular carinae, sef—subbasal elytral foveae, ss—sutural stria, vf—vertexal foveae." httpUri="https://zenodo.org/record/283596/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
C) flat, lacking median carina or sulcus, foveae close in transverse opening.
</paragraph>
<paragraph id="8BF536F22015FFEEFF39DC0E3134F833" blockId="3.[151,1436,1665,2013]" pageId="3" pageNumber="50">
Pronotum (
<figureCitation id="13712A772015FFEEFEB9DC0D32AFF81F" box="[329,397,1917,1941]" captionStart="FIGURE 2" captionStartId="2.[151,250,1741,1764]" captionTargetBox="[151,1425,195,1685]" captionTargetId="figure@2.[151,1436,195,1720]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tyrus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Pronotum. E. Proventrite. F. Left elytron. Scales: 0.2 mm. Abbreviations: as—antebasal sulcus, at—antennal tubercle, bef—basal elytral foveae, ds—discal stria, ff—frontal fovea, gf—gular foveae, laf—lateral antebasal foveae, lpcf—lateral procoxal foveae, lpp—lateral postantennal pits, maf—median antebasal foveae, oc—ocular canthi, omc—ocular-mandibular carinae, sef—subbasal elytral foveae, ss—sutural stria, vf—vertexal foveae." httpUri="https://zenodo.org/record/283596/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
D) with transverse antebasal sulcus connecting median and lateral antebasal foveae; lateral procoxal foveae present (
<figureCitation id="13712A772015FFEEFE42DCD132D7F833" box="[434,501,1953,1977]" captionStart="FIGURE 2" captionStartId="2.[151,250,1741,1764]" captionTargetBox="[151,1425,195,1685]" captionTargetId="figure@2.[151,1436,195,1720]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tyrus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Pronotum. E. Proventrite. F. Left elytron. Scales: 0.2 mm. Abbreviations: as—antebasal sulcus, at—antennal tubercle, bef—basal elytral foveae, ds—discal stria, ff—frontal fovea, gf—gular foveae, laf—lateral antebasal foveae, lpcf—lateral procoxal foveae, lpp—lateral postantennal pits, maf—median antebasal foveae, oc—ocular canthi, omc—ocular-mandibular carinae, sef—subbasal elytral foveae, ss—sutural stria, vf—vertexal foveae." httpUri="https://zenodo.org/record/283596/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
E).
</paragraph>
<paragraph id="8BF536F22015FFE9FF39DCB5309CFF7D" blockId="3.[151,1436,1665,2013]" lastBlockId="4.[151,1436,151,1364]" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50">
Each elytron (
<figureCitation id="13712A772015FFEEFE99DCB5328FF857" box="[361,429,1989,2013]" captionStart="FIGURE 2" captionStartId="2.[151,250,1741,1764]" captionTargetBox="[151,1425,195,1685]" captionTargetId="figure@2.[151,1436,195,1720]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tyrus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Pronotum. E. Proventrite. F. Left elytron. Scales: 0.2 mm. Abbreviations: as—antebasal sulcus, at—antennal tubercle, bef—basal elytral foveae, ds—discal stria, ff—frontal fovea, gf—gular foveae, laf—lateral antebasal foveae, lpcf—lateral procoxal foveae, lpp—lateral postantennal pits, maf—median antebasal foveae, oc—ocular canthi, omc—ocular-mandibular carinae, sef—subbasal elytral foveae, ss—sutural stria, vf—vertexal foveae." httpUri="https://zenodo.org/record/283596/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
F) with two basal foveae; with subbasal fovea; sutural stria entire; discal stria exceeding elytral midpoint; apicolateral clefts present. Thorax (
<figureCitation id="13712A772012FFE9FD1BDBE7300CFF25" box="[747,814,151,175]" captionStart="FIGURE 3" captionStartId="3.[151,250,1511,1534]" captionTargetBox="[165,1425,200,1463]" captionTargetId="figure@3.[151,1436,193,1490]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic features of Tyrus. A. Meso- and metaventrite. B. Abdomen, in lateral view. C. Same, in dorsal view. D. Same, in ventral view. Scales: 0.2 mm. Abbreviations: blf—basolateral foveae, bs—basal sulcus, lmcf—lateral mesocoxal foveae, lmvf—lateral mesoventral foveae, mbf—mediobasal foveae, mdc—median discal carina, mmsf—median mesoventral foveae, mmtf—median metaventral fovea, msff—mesofurcal foveae." httpUri="https://zenodo.org/record/283597/files/figure.png" pageId="4" pageNumber="51">Fig. 3</figureCitation>
A) with median mesoventral foveae in shared opening, lateral mesoventral foveae forked; lateral mesocoxal foveae and mesofurcal foveae present; with median metaventral fovea, posterior margin of metathorax medially emarginate.
</paragraph>
<paragraph id="8BF536F22012FFE9FF39DA75375BFE96" blockId="4.[151,1436,151,1364]" box="[201,1145,260,285]" pageId="4" pageNumber="51">Legs with third tarsomeres longer than second tarsomeres; claws subequal in length.</paragraph>
<paragraph id="8BF536F22012FFE9FF39DA573735FE26" blockId="4.[151,1436,151,1364]" pageId="4" pageNumber="51">
Abdomen (
<figureCitation id="13712A772012FFE9FEB6DA5732A8FEB5" box="[326,394,295,319]" captionStart="FIGURE 3" captionStartId="3.[151,250,1511,1534]" captionTargetBox="[165,1425,200,1463]" captionTargetId="figure@3.[151,1436,193,1490]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic features of Tyrus. A. Meso- and metaventrite. B. Abdomen, in lateral view. C. Same, in dorsal view. D. Same, in ventral view. Scales: 0.2 mm. Abbreviations: blf—basolateral foveae, bs—basal sulcus, lmcf—lateral mesocoxal foveae, lmvf—lateral mesoventral foveae, mbf—mediobasal foveae, mdc—median discal carina, mmsf—median mesoventral foveae, mmtf—median metaventral fovea, msff—mesofurcal foveae." httpUri="https://zenodo.org/record/283597/files/figure.png" pageId="4" pageNumber="51">Figs 3</figureCitation>
BD) with tergite IV as long as to slightly longer than tergite V; tergite IV with shallow basal sulcus, large mediobasal foveae moving laterally approaching small anterior and posterior basolateral foveae on each side, median discal carina usually present; tergites VVIII each with one pair of basolateral foveae; sternite IV with shallow basal sulcus; sternites IVVII each with pair of basolateral foveae.
</paragraph>
<paragraph id="8BF536F22012FFE9FF39DAC73122FE7E" blockId="4.[151,1436,151,1364]" pageId="4" pageNumber="51">Sexual dimorphism usually not obvious. Males of Asian species with the scape lightly modified and the mesotibiae apically protuberant.</paragraph>
</subSubSection>
<subSubSection id="C35065792012FFE9FF39DA8F3123FDD5" pageId="4" pageNumber="51" type="distribution">
<paragraph id="8BF536F22012FFE9FF39DA8F3123FDD5" blockId="4.[151,1436,151,1364]" pageId="4" pageNumber="51">
<emphasis id="B93EEAE02012FFE9FF39DA8F3240FD92" bold="true" box="[201,354,511,536]" pageId="4" pageNumber="51">Distribution.</emphasis>
Seven species are known from the Oriental region, three species and one subspecies from the Palearctic region, three species from the Nearctic region, and one extinct species described from Baltic amber (Hlaváč &amp; Chandler 2005: 117).
</paragraph>
</subSubSection>
<subSubSection id="C35065792012FFE9FF39D91B301AFADE" pageId="4" pageNumber="51" type="biology_ecology">
<paragraph id="8BF536F22012FFE9FF39D91B326AFBDD" blockId="4.[151,1436,151,1364]" pageId="4" pageNumber="51">
<emphasis id="B93EEAE02012FFE9FF39D91B320BFD0E" bold="true" box="[201,297,619,644]" pageId="4" pageNumber="51">Biology.</emphasis>
Biological information for
<taxonomicName id="4C4A4D712012FFE9FD91D91D31BFFD0E" box="[609,669,621,644]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FD91D91D31BFFD0E" box="[609,669,621,644]" italics="true" pageId="4" pageNumber="51">Tyrus</emphasis>
</taxonomicName>
species, if known, is generally limited to statements on the habitats in which the adults have been collected. For the Nearctic species,
<taxonomicName id="4C4A4D712012FFE9FC7DD9E13758FD2D" authority="Casey" authorityName="Casey" box="[909,1146,655,680]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="corticinus">
<emphasis id="B93EEAE02012FFE9FC7DD9E13738FD22" box="[909,1050,657,680]" italics="true" pageId="4" pageNumber="51">T. corticinus</emphasis>
(Casey)
</taxonomicName>
was found under bark of pines and Douglas fir, under bark of
<taxonomicName id="4C4A4D712012FFE9FDC9D9C43033FD46" box="[569,785,692,716]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="occidentalis">
<emphasis id="B93EEAE02012FFE9FDC9D9C43033FD46" box="[569,785,692,716]" italics="true" pageId="4" pageNumber="51">Betula occidentalis</emphasis>
</taxonomicName>
, and with ant
<taxonomicName id="4C4A4D712012FFE9FC44D9C537B2FD46" box="[948,1168,693,716]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="sanguinea">
<emphasis id="B93EEAE02012FFE9FC44D9C537B2FD46" box="[948,1168,693,716]" italics="true" pageId="4" pageNumber="51">Formica sanguinea</emphasis>
</taxonomicName>
(Casey 1887; Chandler 1997, 2003; Schwarz 1895), and
<taxonomicName id="4C4A4D712012FFE9FDE5D9A931D9FD65" authority="Aube" authorityName="Aube" box="[533,763,727,752]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B93EEAE02012FFE9FDE5D9A93181FD7A" box="[533,675,728,752]" italics="true" pageId="4" pageNumber="51">T. humeralis</emphasis>
(Aubé)
</taxonomicName>
was found under bark of pine, spruce and balsam fir, also from tree holes, in rotten pine log, at UV light, and with ants
<taxonomicName id="4C4A4D712012FFE9FC72D98C37EDFC9E" box="[898,1231,764,788]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis id="B93EEAE02012FFE9FC72D98C37EDFC9E" box="[898,1231,764,788]" italics="true" pageId="4" pageNumber="51">Aphaenogaster tennesseensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4A4D712012FFE9FAFED98D3260FCB2" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="pennsylvanicus">
<emphasis id="B93EEAE02012FFE9FAFED98D3260FCB2" italics="true" pageId="4" pageNumber="51">Camponotus pennsylvanicus</emphasis>
</taxonomicName>
(Chandler 1997; Schwarz 1890; Eldredge pers. comm.). In the Palaearctic region,
<taxonomicName id="4C4A4D712012FFE9FAE0D85133FAFCD6" authority="Sharp" authorityName="Sharp" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B93EEAE02012FFE9FAE0D85136BEFCB2" box="[1296,1436,801,824]" italics="true" pageId="4" pageNumber="51">T. japonicus</emphasis>
Sharp
</taxonomicName>
was collected under bark of large fallen trees, rotten woods and by flight intercept trap in the deciduous forest (Nomura pers. comm.),
<taxonomicName id="4C4A4D712012FFE9FE19D8193054FCF5" authority="Panzer" authorityName="Panzer" box="[489,886,871,896]" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="subSpecies" species="mucronatus" subSpecies="mucronatus">
<emphasis id="B93EEAE02012FFE9FE19D8193032FC0A" box="[489,784,873,896]" italics="true" pageId="4" pageNumber="51">T. mucronatus mucronatus</emphasis>
(Panzer)
</taxonomicName>
was found under bark of dead trees, in moss, and with ant
<taxonomicName id="4C4A4D712012FFE9FF09D8FD32F2FC2E" box="[249,464,908,932]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B93EEAE02012FFE9FF09D8FD32F2FC2E" box="[249,464,908,932]" italics="true" pageId="4" pageNumber="51">Formica fuliginosa</emphasis>
</taxonomicName>
(Besuchet 1974; Casey 1887; Fleischer 1888; Jeannel 1950; Schatzmayr 1904), and
<taxonomicName id="4C4A4D712012FFE9FA77D8FD3263FC4D" authority="Saulcy" authorityName="Saulcy" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="peyroni">
<emphasis id="B93EEAE02012FFE9FA77D8FD33CEFC42" italics="true" pageId="4" pageNumber="51">T. peyroni</emphasis>
Saulcy
</taxonomicName>
was taken under a stone (Saulcy 1874). The Oriental
<taxonomicName id="4C4A4D712012FFE9FC4CD8C13794FC4D" authority="Raffray" authorityName="Raffray" box="[956,1206,943,968]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="corticalis">
<emphasis id="B93EEAE02012FFE9FC4CD8C13760FC42" box="[956,1090,944,968]" italics="true" pageId="4" pageNumber="51">T. corticalis</emphasis>
(Raffray)
</taxonomicName>
was collected from under tree bark (Raffray 1890),
<taxonomicName id="4C4A4D712012FFE9FDF6D8A53001FC67" authority="Bryant" authorityName="Bryant" box="[518,803,980,1005]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="myrmecophilus">
<emphasis id="B93EEAE02012FFE9FDF6D8A531ECFC66" box="[518,718,980,1004]" italics="true" pageId="4" pageNumber="51">T. myrmecophilus</emphasis>
Bryant
</taxonomicName>
was found under bark with ants (Bryant 1915, without statement of species or genus),
<taxonomicName id="4C4A4D712012FFE9FE07D88931B7FB9A" box="[503,661,1017,1040]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B93EEAE02012FFE9FE07D88931B7FB9A" box="[503,661,1017,1040]" italics="true" pageId="4" pageNumber="51">Tyrus sinensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C4A4D712012FFE9FD23D8893050FB9A" box="[723,882,1016,1040]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="sichuanicus">
<emphasis id="B93EEAE02012FFE9FD23D8893050FB9A" box="[723,882,1016,1040]" italics="true" pageId="4" pageNumber="51">T. sichuanicus</emphasis>
</taxonomicName>
described below were taken from ant colonies of
<taxonomicName id="4C4A4D712012FFE9FF67DF6D33DAFBBE" box="[151,248,1053,1076]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FF67DF6D33DAFBBE" box="[151,248,1053,1076]" italics="true" pageId="4" pageNumber="51">Formica</emphasis>
</taxonomicName>
spp. nesting under rock or fallen tree, and
<taxonomicName id="4C4A4D712012FFE9FD02DF6D30BEFBBE" box="[754,924,1053,1076]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="yajiangensis">
<emphasis id="B93EEAE02012FFE9FD02DF6D30BEFBBE" box="[754,924,1053,1076]" italics="true" pageId="4" pageNumber="51">T. yajiangensis</emphasis>
</taxonomicName>
was possibly taken from sifted leaf litter or under tree bark.
</paragraph>
<paragraph id="8BF536F22012FFE9FF39DF13301AFADE" blockId="4.[151,1436,151,1364]" pageId="4" pageNumber="51">
<emphasis id="B93EEAE02012FFE9FF39DF133291FBF6" bold="true" box="[201,435,1123,1148]" pageId="4" pageNumber="51">Comparative notes.</emphasis>
<taxonomicName id="4C4A4D712012FFE9FE4CDF1532DAFBF6" box="[444,504,1125,1148]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FE4CDF1532DAFBF6" box="[444,504,1125,1148]" italics="true" pageId="4" pageNumber="51">Tyrus</emphasis>
</taxonomicName>
is allied to the largely Oriental
<taxonomicName id="4C4A4D712012FFE9FC95DF143738FBF6" authority="Raffray" authorityName="Raffray" box="[869,1050,1124,1148]" class="Insecta" family="Staphylinidae" genus="Tyrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FC95DF14309EFBF6" box="[869,956,1124,1148]" italics="true" pageId="4" pageNumber="51">Tyrodes</emphasis>
Raffray
</taxonomicName>
, and
<collectingCountry id="F35D76622012FFE9FBA7DF1537CCFBF6" box="[1111,1262,1124,1148]" name="New Zealand" pageId="4" pageNumber="51">New Zealand</collectingCountry>
and Australian
<taxonomicName id="4C4A4D712012FFE9FF67DFF832AFFB2A" authority="Schaufuss" authorityName="Schaufuss" box="[151,397,1160,1184]" class="Insecta" family="Staphylinidae" genus="Hamotulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FF67DFF83230FB2A" box="[151,274,1160,1184]" italics="true" pageId="4" pageNumber="51">Hamotulus</emphasis>
Schaufuss
</taxonomicName>
based on the basally pedunculate, elongate maxillary palpomeres IIIV, and the pronotum with an antebasal sulcus that connects the median and lateral antebasal foveae.
<taxonomicName id="4C4A4D712012FFE9FBD8DFDC375FFB4E" box="[1064,1149,1196,1220]" class="Insecta" family="Staphylinidae" genus="Tyrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FBD8DFDC375FFB4E" box="[1064,1149,1196,1220]" italics="true" pageId="4" pageNumber="51">Tyrodes</emphasis>
</taxonomicName>
is generally smaller, has small, nude, easily overlooked frontal fovea (Hlaváč &amp; Chandler 2005: 93) while
<taxonomicName id="4C4A4D712012FFE9FB92DFA137BCFB62" box="[1122,1182,1233,1256]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FB92DFA137BCFB62" box="[1122,1182,1233,1256]" italics="true" pageId="4" pageNumber="51">Tyrus</emphasis>
</taxonomicName>
has greater size, and distinct and setose frontal fovea.
<taxonomicName id="4C4A4D712012FFE9FDFDDF8431AAFA86" box="[525,648,1268,1292]" class="Insecta" family="Staphylinidae" genus="Hamotulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FDFDDF8431AAFA86" box="[525,648,1268,1292]" italics="true" pageId="4" pageNumber="51">Hamotulus</emphasis>
</taxonomicName>
has a pair of tubercles between the vertexal and frontal foveae on the head (absent in one species), and a deep sulcus at the base of tergite IV (Chandler 2001: 433), while
<taxonomicName id="4C4A4D712012FFE9FAECDE693675FABA" box="[1308,1367,1305,1328]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FAECDE693675FABA" box="[1308,1367,1305,1328]" italics="true" pageId="4" pageNumber="51">Tyrus</emphasis>
</taxonomicName>
lacks these tubercles, and the basal sulcus of tergite IV is shallow.
</paragraph>
</subSubSection>
<subSubSection id="C35065792012FFE9FF67DED83239FA6E" pageId="4" pageNumber="51" type="description">
<paragraph id="8BF536F22012FFE9FF67DED83239FA6E" blockId="4.[151,722,1447,1508]" pageId="4" pageNumber="51">
<emphasis id="B93EEAE02012FFE9FF67DED831F0FA4A" bold="true" box="[151,722,1447,1472]" pageId="4" pageNumber="51">
<taxonomicName id="4C4A4D712012FFE9FF67DED83119FA4A" authority="Yin and Nomura" authorityName="Yin and Nomura" box="[151,571,1447,1472]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="sichuanicus" status="sp. nov.">
<emphasis id="B93EEAE02012FFE9FF67DED83249FA4A" bold="true" box="[151,363,1447,1472]" italics="true" pageId="4" pageNumber="51">Tyrus sichuanicus</emphasis>
Yin and Nomura
</taxonomicName>
,
<taxonomicNameLabel id="A20D579B2012FFE9FDB8DED831F0FA4A" box="[584,722,1447,1472]" pageId="4" pageNumber="51" rank="species">new species</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13712A772012FFE9FF6FDEBC33FCFA6E" box="[159,222,1484,1508]" captionStart="FIGURE 1" captionStartId="1.[151,250,1960,1983]" captionTargetBox="[180,1402,786,1918]" captionTargetId="figure@1.[171,1408,769,1924]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Tyrus sichuanicus (A) and T. sinensis (B). Scales: 0.5 mm." httpUri="https://zenodo.org/record/283595/files/figure.png" pageId="4" pageNumber="51">Fig. 1</figureCitation>
A, 4)
</paragraph>
</subSubSection>
<subSubSection id="C35065792012FFE9FF67DD6432F6F81E" pageId="4" pageNumber="51" type="materials_examined">
<paragraph id="8BF536F22012FFE9FF67DD6432F6F81E" blockId="4.[151,1436,1555,2012]" pageId="4" pageNumber="51">
<emphasis id="B93EEAE02012FFE9FF67DD643260F9A6" bold="true" box="[151,322,1555,1580]" pageId="4" pageNumber="51">
<typeStatus id="54F188502012FFE9FF67DD6433F0F9A6" box="[151,210,1556,1580]" pageId="4" pageNumber="51">Type</typeStatus>
material
</emphasis>
(
<date id="FFF410322012FFE9FEA7DD6432E8F9A6" box="[343,458,1556,1580]" pageId="4" pageNumber="51" value="1933-06-23">6 33, 23</date>
♀♀).
<emphasis id="B93EEAE02012FFE9FDECDD643027F9A6" bold="true" box="[540,773,1555,1580]" pageId="4" pageNumber="51">
<typeStatus id="54F188502012FFE9FDECDD6431AFF9A6" box="[540,653,1556,1580]" pageId="4" pageNumber="51" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F35D76622012FFE9FD6EDD6331DDF9A6" box="[670,767,1555,1580]" name="China" pageId="4" pageNumber="51">CHINA</collectingCountry>
:
</emphasis>
3, labeled
<collectingCountry id="F35D76622012FFE9FC69DD6430D8F9A6" box="[921,1018,1556,1580]" name="China" pageId="4" pageNumber="51">CHINA</collectingCountry>
: Sichuan, Yaan County / Shimian, Liziping N. R., Chaheba / roadside, 28°57ʹ21″N / 102°15ʹ0 1″E, alt.
<quantity id="4CB29B172012FFE9FC65DD4730CFF9C5" box="[917,1005,1591,1615]" metricMagnitude="3" metricUnit="m" metricValue="2.6149999999999998" pageId="4" pageNumber="51" unit="m" value="2615.0">2615 m</quantity>
, colony / of
<taxonomicName id="4C4A4D712012FFE9FB8CDD4937FEF9DA" box="[1148,1244,1593,1616]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FB8CDD4937FEF9DA" box="[1148,1244,1593,1616]" italics="true" pageId="4" pageNumber="51">Formica</emphasis>
</taxonomicName>
sp., under rock /
<date id="FFF410322012FFE9FF67DD2C323BF9FE" box="[151,281,1628,1652]" pageId="4" pageNumber="51" value="2012-07-16">16.vii.2012</date>
, Dazi, Peng &amp; Yin leg. [1 ex.
<taxonomicName id="4C4A4D712012FFE9FD98DD2D31EBF9FE" box="[616,713,1629,1652]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FD98DD2D31EBF9FE" box="[616,713,1629,1652]" italics="true" pageId="4" pageNumber="51">Formica</emphasis>
</taxonomicName>
sp. pinned under
<typeStatus id="54F188502012FFE9FC7FDD2C30DAF9FE" box="[911,1016,1628,1652]" pageId="4" pageNumber="51" type="holotype">holotype</typeStatus>
] (SNUC).
<emphasis id="B93EEAE02012FFE9FB86DD2C3646F9FE" bold="true" box="[1142,1380,1627,1652]" pageId="4" pageNumber="51">
<typeStatus id="54F188502012FFE9FB86DD2C37D0F9FE" box="[1142,1266,1628,1652]" pageId="4" pageNumber="51" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F35D76622012FFE9FB0EDD2B367DF9FE" box="[1278,1375,1627,1652]" name="China" pageId="4" pageNumber="51">CHINA</collectingCountry>
:
</emphasis>
<date id="FFF410322012FFE9FA9BDD2C3391F91D" pageId="4" pageNumber="51" value="2010-01-03">1 3, 10</date>
♀♀, same label data as
<typeStatus id="54F188502012FFE9FE10DD0F3164F91D" box="[480,582,1663,1687]" pageId="4" pageNumber="51" type="holotype">holotype</typeStatus>
;
<date id="FFF410322012FFE9FDA6DD0F31E5F91D" box="[598,711,1663,1688]" pageId="4" pageNumber="51" value="1933-02-11">2 33, 11</date>
♀♀, same label data, except
<date id="FFF410322012FFE9FBC2DD0F37E3F91D" box="[1074,1217,1663,1687]" pageId="4" pageNumber="51" value="2012-08-02">02.viii.2012</date>
, Zhong Peng leg. (SNUC); 1 3, labeled Mt., Jiudingshan (
<quantity id="4CB29B172012FFE9FD82DDD431F0F936" box="[626,722,1700,1724]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="51" unit="m" value="3200.0">3,200 m</quantity>
) / Chapingshan Mts. / Mao Xian, Sichuan Prov. // (SE-China) [should be SW-China!] /
<date id="FFF410322012FFE9FE41DDB73108F955" box="[433,554,1735,1759]" pageId="4" pageNumber="51" value="1996-09-22">22.ix.1996</date>
, S. Nomura leg. [1 ex.
<taxonomicName id="4C4A4D712012FFE9FCDFDDB930ADF96A" box="[815,911,1737,1760]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FCDFDDB930ADF96A" box="[815,911,1737,1760]" italics="true" pageId="4" pageNumber="51">Formica</emphasis>
</taxonomicName>
sp. pinned under the
<typeStatus id="54F188502012FFE9FB71DDB737CBF955" box="[1153,1257,1735,1759]" pageId="4" pageNumber="51" type="paratype">paratype</typeStatus>
] (NSMT); 1 ♀, labeled
<collectingCountry id="F35D76622012FFE9FF03DD9C321DF88E" box="[243,319,1772,1796]" name="China" pageId="4" pageNumber="51">China</collectingCountry>
: Sichuan Prov. / Yakou (Meigu to Yuexi) / 28°39ʹN, 102°59ʹ E /
<quantity id="4CB29B172012FFE9FBE8DD9C3752F88E" box="[1048,1136,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="2.55" pageId="4" pageNumber="51" unit="m" value="2550.0">2550 m</quantity>
,
<date id="FFF410322012FFE9FB8CDD9C3622F88E" box="[1148,1280,1772,1796]" pageId="4" pageNumber="51" value="2012-06-14">14-VI-2012</date>
/ Ye LIU leg. [2 ex.
<taxonomicName id="4C4A4D712012FFE9FF13DC613261F8A2" box="[227,323,1809,1832]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02012FFE9FF13DC613261F8A2" box="[227,323,1809,1832]" italics="true" pageId="4" pageNumber="51">Formica</emphasis>
</taxonomicName>
sp. pinned under the
<typeStatus id="54F188502012FFE9FDB3DC7F3189F8AD" box="[579,683,1807,1831]" pageId="4" pageNumber="51" type="paratype">paratype</typeStatus>
] (SNUC); 1 3, labeled
<collectingCountry id="F35D76622012FFE9FC3BDC7F370FF8AD" box="[971,1069,1807,1831]" name="China" pageId="4" pageNumber="51">CHINA</collectingCountry>
: W-Sichuan (15) / Daxue Shan, Hailuogou Glacier / Park, Camp 2, 2550
<quantity id="4CB29B172012FFE9FD7BDC4431C7F8C6" box="[651,741,1844,1868]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="51" unit="m" value="2700.0">2700 m</quantity>
/ 29°35ʹ16″N, 102°01ʹ53″E / 30./
<date id="FFF410322012FFE9FB88DC4437DEF8C6" box="[1144,1276,1844,1868]" pageId="4" pageNumber="51" value="1997-05-31">31.05.1997</date>
, M. Schülke (pcMS); 1 ♀, labeled
<collectingCountry id="F35D76622012FFE9FE65DC2732FDF8E5" box="[405,479,1879,1903]" name="China" pageId="4" pageNumber="51">China</collectingCountry>
, Sichuan Prov., / Liziping env., near / Shimian,
<quantity id="4CB29B172012FFE9FBF1DC27377AF8E5" box="[1025,1112,1879,1903]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="51" unit="km" value="200.0">200 km</quantity>
SW / of Yaan,
<date id="FFF410322012FFE9FAE3DC2733F3F81E" pageId="4" pageNumber="51" value="1991-06-27" valueMax="1991-07-03" valueMin="1991-06-27">27.vi3.vii / 1991</date>
, Z. Kejval lgt (pcPH).
</paragraph>
</subSubSection>
<subSubSection id="C35065792012FFE9FF39DCEF370CF856" pageId="4" pageNumber="51" type="diagnosis">
<paragraph id="8BF536F22012FFE9FF39DCEF370CF856" blockId="4.[151,1436,1555,2012]" pageId="4" pageNumber="51">
<emphasis id="B93EEAE02012FFE9FF39DCEF3261F832" bold="true" box="[201,323,1951,1976]" pageId="4" pageNumber="51">Diagnosis.</emphasis>
Antennomeres IXX each subcylindrical; ocular canthi present; maxillary palpomeres IV nearly fusiform; mesotrochanters with two long spines; mesotibiae with long apical spur.
</paragraph>
</subSubSection>
<caption id="DF35667A2013FFE8FF67DE59365BFA10" httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52" targetBox="[156,1426,199,1295]" targetPageId="5">
<paragraph id="8BF536F22013FFE8FF67DE59365BFA10" blockId="5.[151,1437,1321,1435]" pageId="5" pageNumber="52">
<emphasis id="B93EEAE02013FFE8FF67DE593230FAB4" bold="true" box="[151,274,1321,1344]" pageId="5" pageNumber="52">FIGURE 4.</emphasis>
Diagnostic features of
<taxonomicName id="4C4A4D712013FFE8FE0CDE5A3193FAB5" box="[508,689,1321,1343]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="sichuanicus">
<emphasis id="B93EEAE02013FFE8FE0CDE5A3193FAB5" box="[508,689,1321,1343]" italics="true" pageId="5" pageNumber="52">Tyrus sichuanicus</emphasis>
</taxonomicName>
.
<emphasis id="B93EEAE02013FFE8FD4DDE5931F7FAB4" bold="true" box="[701,725,1321,1342]" pageId="5" pageNumber="52">A.</emphasis>
Antenna.
<emphasis id="B93EEAE02013FFE8FCCDDE5A3076FAB5" bold="true" box="[829,852,1322,1343]" pageId="5" pageNumber="52">B.</emphasis>
antennal club, enlarged.
<emphasis id="B93EEAE02013FFE8FBA0DE59374AFACA" bold="true" box="[1104,1128,1321,1344]" pageId="5" pageNumber="52">C.</emphasis>
Scape, enlarged.
<emphasis id="B93EEAE02013FFE8FAEADE5A3610FAB5" bold="true" box="[1306,1330,1322,1343]" pageId="5" pageNumber="52">D.</emphasis>
Maxillary palpus.
<emphasis id="B93EEAE02013FFE8FF15DE3833DFFAD7" bold="true" box="[229,253,1352,1373]" pageId="5" pageNumber="52">E.</emphasis>
Protrochanter and protibia.
<emphasis id="B93EEAE02013FFE8FDE7DE38310BFAD7" bold="true" box="[535,553,1352,1373]" pageId="5" pageNumber="52">F.</emphasis>
Mesotrochanter and mesotibia.
<emphasis id="B93EEAE02013FFE8FC9EDE3730A1FAD4" bold="true" box="[878,899,1351,1374]" pageId="5" pageNumber="52">G.</emphasis>
Mesotibia.
<emphasis id="B93EEAE02013FFE8FC0DDE383735FAD7" bold="true" box="[1021,1047,1352,1373]" pageId="5" pageNumber="52">H.</emphasis>
Apical portion of mesotibia, enlarged.
<emphasis id="B93EEAE02013FFE8FF67DE173385FAF6" bold="true" box="[151,167,1383,1404]" pageId="5" pageNumber="52">I.</emphasis>
Male sternite IX.
<emphasis id="B93EEAE02013FFE8FE91DE173256FAF6" bold="true" box="[353,372,1383,1404]" pageId="5" pageNumber="52">J.</emphasis>
Aedeagus, in dorsal view.
<emphasis id="B93EEAE02013FFE8FD76DE173182FAF6" bold="true" box="[646,672,1383,1404]" pageId="5" pageNumber="52">K.</emphasis>
Same, in lateral view.
<emphasis id="B93EEAE02013FFE8FC78DE1730BDFAF6" bold="true" box="[904,927,1383,1404]" pageId="5" pageNumber="52">L.</emphasis>
Same, in ventral view.
<emphasis id="B93EEAE02013FFE8FB7EDE17378FFAF6" bold="true" box="[1166,1197,1383,1404]" pageId="5" pageNumber="52">M.</emphasis>
Female tergite VIII.
<emphasis id="B93EEAE02013FFE8FA73DE1736BEFAF6" bold="true" box="[1411,1436,1383,1404]" pageId="5" pageNumber="52">N.</emphasis>
Female sternite VIII.
<emphasis id="B93EEAE02013FFE8FE81DEF432A9FA11" bold="true" box="[369,395,1412,1435]" pageId="5" pageNumber="52">O.</emphasis>
Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1.
</paragraph>
</caption>
<subSubSection id="C35065792013FFE8FF39DEB231FBF800" pageId="5" pageNumber="52" type="description">
<paragraph id="8BF536F22013FFE8FF39DEB23053F970" blockId="5.[151,1436,1474,2003]" pageId="5" pageNumber="52">
<emphasis id="B93EEAE02013FFE8FF39DEB23278FA51" bold="true" box="[201,346,1474,1499]" pageId="5" pageNumber="52">Description.</emphasis>
Male (
<figureCitation id="13712A772013FFE8FE40DEB232D0FA50" box="[432,498,1474,1498]" captionStart="FIGURE 1" captionStartId="1.[151,250,1960,1983]" captionTargetBox="[180,1402,786,1918]" captionTargetId="figure@1.[171,1408,769,1924]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Tyrus sichuanicus (A) and T. sinensis (B). Scales: 0.5 mm." httpUri="https://zenodo.org/record/283595/files/figure.png" pageId="5" pageNumber="52">Fig. 1</figureCitation>
A). Length
<quantity id="4CB29B172013FFE8FD8DDEB23004FA50" box="[637,806,1474,1498]" metricMagnitude="-3" metricUnit="m" metricValue="2.495" metricValueMax="2.7" metricValueMin="2.29" pageId="5" pageNumber="52" unit="mm" value="2.495" valueMax="2.7" valueMin="2.29">2.292.70 mm</quantity>
. Head about as long as wide, HL
<quantity id="4CB29B172013FFE8FB47DEB23642FA50" box="[1207,1376,1474,1498]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="5.5" metricValueMin="4.699999999999999" pageId="5" pageNumber="52" unit="mm" value="0.51" valueMax="0.55" valueMin="0.47">0.470.55 mm</quantity>
, HW
<quantity id="4CB29B172013FFE8FF67DE96321EFA74" box="[151,316,1510,1534]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="4.9" metricValueMin="4.5" pageId="5" pageNumber="52" unit="mm" value="0.47" valueMax="0.49" valueMin="0.45">0.450.49 mm</quantity>
; eyes each composed of about 4050 facets. Antennae (
<figureCitation id="13712A772013FFE8FC5BDE9630CFFA74" box="[939,1005,1510,1534]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
A) elongate, scapes laterally expanded (
<figureCitation id="13712A772013FFE8FF6FDD7A33C0F9A8" box="[159,226,1546,1570]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
C), clubs as in
<figureCitation id="13712A772013FFE8FE76DD7A32EAF9A8" box="[390,456,1546,1570]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
B. Maxillary palpi (
<figureCitation id="13712A772013FFE8FD52DD7A31C6F9A8" box="[674,740,1546,1570]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
D) with palpomeres I short, II lengthily pedunculate, III nearly triangular, IV fusiform. Pronotum slightly longer than wide, PL
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, PW
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, with lateral margins nearly straight. Elytra wider than long, EL
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, EW
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. Protrochanters with triangular ventral spine, profemora with two blunt spines on ventral margin (
<figureCitation id="13712A772013FFE8FBF7DD06376EF904" box="[1031,1100,1654,1678]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
E); mesotrochanters (
<figureCitation id="13712A772013FFE8FACFDD0636A1F904" box="[1343,1411,1654,1678]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
F) with two large ventral spines; mesotibiae (
<figureCitation id="13712A772013FFE8FD77DDEA31EFF938" box="[647,717,1690,1714]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
G) slightly arched, with long apical spur (
<figureCitation id="13712A772013FFE8FB49DDEA37DDF938" box="[1209,1279,1690,1714]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
H). Abdomen broad at base and narrowed apically, AL
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, AW
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. Sternite IX as in
<figureCitation id="13712A772013FFE8FB21DDCE3634F95C" box="[1233,1302,1726,1750]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">
Fig.
<date id="FFF410322013FFE8FAF6DDCE3634F95C" box="[1286,1302,1726,1750]" pageId="5" pageNumber="52">4</date>
</figureCitation>
I. Aedeagus length
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, with asymmetric median lobe (
<figureCitation id="13712A772013FFE8FD1FDD923015F970" box="[751,823,1762,1786]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Figs 4</figureCitation>
JL).
</paragraph>
<paragraph id="8BF536F22013FFE8FF39DC7631FBF800" blockId="5.[151,1436,1474,2003]" pageId="5" pageNumber="52">
Female. Similar to male in general; BL
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, HL
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, HW
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, PL
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, PW
<quantity id="4CB29B172013FFE8FEF1DC5A3285F8C8" box="[257,423,1834,1858]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.6" metricValueMin="4.8" pageId="5" pageNumber="52" unit="mm" value="0.52" valueMax="0.56" valueMin="0.48">0.480.56 mm</quantity>
, EL
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, EW
<quantity id="4CB29B172013FFE8FD4CDC5A3043F8C8" box="[700,865,1834,1858]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.07" metricValueMin="0.93" pageId="5" pageNumber="52" unit="mm" value="1.0" valueMax="1.07" valueMin="0.93">0.931.07 mm</quantity>
, AL
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, AW
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. Eyes each composed of about 40 facets. Scape simple; mesotibiae lacking apical spur; tergite VIII (
<figureCitation id="13712A772013FFE8FB88DC3E3798F8EC" box="[1144,1210,1870,1894]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
M) and sternite VIII (
<figureCitation id="13712A772013FFE8FF6FDC0233C0F800" box="[159,226,1906,1930]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
N) transverse; genital complex as in
<figureCitation id="13712A772013FFE8FD8CDC02319DF800" box="[636,703,1906,1930]" captionStart="FIGURE 4" captionStartId="5.[151,250,1321,1344]" captionTargetBox="[156,1426,199,1295]" captionTargetId="figure@5.[151,1436,193,1300]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tyrus sichuanicus. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. M. Female tergite VIII. N. Female sternite VIII. O. Female genital complex. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, H, I, M, N, O = 0.1." httpUri="https://zenodo.org/record/283598/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
O.
</paragraph>
</subSubSection>
<subSubSection id="C35065792013FFE8FF39DCE632E6F858" pageId="5" pageNumber="52" type="distribution">
<paragraph id="8BF536F22013FFE8FF39DCE631BAF824" blockId="5.[151,1436,1474,2003]" box="[201,664,1942,1967]" pageId="5" pageNumber="52">
<emphasis id="B93EEAE02013FFE8FF39DCE63240F825" bold="true" box="[201,354,1942,1967]" pageId="5" pageNumber="52">Distribution.</emphasis>
Southwest
<collectingCountry id="F35D76622013FFE8FE15DCE6310EF824" box="[485,556,1942,1966]" name="China" pageId="5" pageNumber="52">China</collectingCountry>
: Sichuan.
</paragraph>
<paragraph id="8BF536F22013FFE8FF39DCCB32E6F858" blockId="5.[151,1436,1474,2003]" box="[201,452,1979,2003]" pageId="5" pageNumber="52">
<emphasis id="B93EEAE02013FFE8FF39DCCB3214F859" bold="true" box="[201,310,1979,2003]" pageId="5" pageNumber="52">Host ant.</emphasis>
<taxonomicName id="4C4A4D712013FFE8FECDDCCB32BFF858" box="[317,413,1979,2002]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02013FFE8FECDDCCB32BFF858" box="[317,413,1979,2002]" italics="true" pageId="5" pageNumber="52">Formica</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
<subSubSection id="C35065792010FFEBFF39DBE73773FEEE" pageId="6" pageNumber="53" type="biology_ecology">
<paragraph id="8BF536F22010FFEBFF39DBE73773FEEE" blockId="6.[151,1436,151,392]" pageId="6" pageNumber="53">
<emphasis id="B93EEAE02010FFEBFF39DBE7320BFF3A" bold="true" box="[201,297,151,176]" pageId="6" pageNumber="53">Biology.</emphasis>
All adults were collected from ant nests during an investigation of myrmecophilous pselaphine in Liziping Natrual Reserve (
<figureCitation id="13712A772010FFEBFE3FDBCC3138FF5E" box="[463,538,188,212]" captionStart="FIGURE 7" captionStartId="9.[151,250,1910,1933]" captionTargetBox="[151,1435,283,1888]" captionTargetId="figure@9.[151,1436,283,1889]" captionTargetPageId="9" captionText="FIGURE 7. Habitat of Tyrus sichuanicus and T. sinensis. A. Collection site of T. sichuanicus — Chaheba Mt., alt. 2615 m. B. Peng &amp; Yin investigated a nest of Formica sp. at Chaheba Mt. C. A closer view of the nest. D. Collection site of T. sinensis — Hongla Shan Mt., alt. 3700 m (photo by Wen-Xuan Bi)." httpUri="https://zenodo.org/record/283601/files/figure.png" pageId="6" pageNumber="53">Figs 7</figureCitation>
AC). More than one-hundred ant nests were explored, and 203 pselaphines placed in 11 species were collected. The environment was dominated by two ant species,
<taxonomicName id="4C4A4D712010FFEBFB8ADB9137F8FF72" box="[1146,1242,225,248]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02010FFEBFB8ADB9137F8FF72" box="[1146,1242,225,248]" italics="true" pageId="6" pageNumber="53">Formica</emphasis>
</taxonomicName>
sp. and
<taxonomicName id="4C4A4D712010FFEBFAC7DB9136BEFF72" box="[1335,1436,225,248]" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02010FFEBFAC7DB9136BEFF72" box="[1335,1436,225,248]" italics="true" pageId="6" pageNumber="53">Myrmica</emphasis>
</taxonomicName>
sp. Both species build their nests under different sizes of rocks. Unfortunately the two ant species were confused at the collection site. The association of the new species with
<taxonomicName id="4C4A4D712010FFEBFCD9DA5930ABFECA" box="[809,905,297,320]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02010FFEBFCD9DA5930ABFECA" box="[809,905,297,320]" italics="true" pageId="6" pageNumber="53">Formica</emphasis>
</taxonomicName>
sp. was confirmed based on the
<typeStatus id="54F188502010FFEBFB03DA58367CFECA" box="[1267,1374,296,320]" pageId="6" pageNumber="53" type="paratype">paratypes</typeStatus>
from Jiudingshan and from the Pass between Meigu and Yuexi (see collection data above).
</paragraph>
</subSubSection>
<subSubSection id="C35065792010FFEAFF39DA00323BFF5E" lastPageId="7" lastPageNumber="54" pageId="6" pageNumber="53" type="etymology">
<paragraph id="8BF536F22010FFEBFF39DA0036BEFE0D" blockId="6.[151,1436,151,392]" box="[201,1436,367,392]" pageId="6" pageNumber="53">
<emphasis id="B93EEAE02010FFEBFF39DA00326DFE02" bold="true" box="[201,335,368,392]" pageId="6" pageNumber="53">Etymology.</emphasis>
The new species is named after the province where all individuals of the new species were found.
</paragraph>
<caption id="DF35667A2010FFEBFF67DC493164F821" httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="6" pageNumber="53" targetBox="[159,1430,457,1819]" targetPageId="6">
<paragraph id="8BF536F22010FFEBFF67DC493164F821" blockId="6.[151,1437,1849,1963]" pageId="6" pageNumber="53">
<emphasis id="B93EEAE02010FFEBFF67DC493236F8C5" bold="true" box="[151,276,1849,1872]" pageId="6" pageNumber="53">FIGURE 5.</emphasis>
Diagnostic features of
<taxonomicName id="4C4A4D712010FFEBFDF7DC4B31B5F8DA" box="[519,663,1851,1872]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B93EEAE02010FFEBFDF7DC4B31B5F8DA" box="[519,663,1851,1872]" italics="true" pageId="6" pageNumber="53">Tyrus sinensis</emphasis>
</taxonomicName>
.
<emphasis id="B93EEAE02010FFEBFD56DC4A319CF8C5" bold="true" box="[678,702,1850,1871]" pageId="6" pageNumber="53">A.</emphasis>
Antenna.
<emphasis id="B93EEAE02010FFEBFCDBDC4A3060F8C5" bold="true" box="[811,834,1850,1871]" pageId="6" pageNumber="53">B.</emphasis>
antennal club, enlarged.
<emphasis id="B93EEAE02010FFEBFBB7DC49377DF8DA" bold="true" box="[1095,1119,1849,1872]" pageId="6" pageNumber="53">C.</emphasis>
Scape, enlarged.
<emphasis id="B93EEAE02010FFEBFAE7DC4A3612F8C5" bold="true" box="[1303,1328,1850,1871]" pageId="6" pageNumber="53">D.</emphasis>
Maxillary palpus.
<emphasis id="B93EEAE02010FFEBFF15DC2933DFF8E4" bold="true" box="[229,253,1881,1902]" pageId="6" pageNumber="53">E.</emphasis>
Protrochanter and protibia.
<emphasis id="B93EEAE02010FFEBFDE7DC29310BF8E4" bold="true" box="[535,553,1881,1902]" pageId="6" pageNumber="53">F.</emphasis>
Mesotrochanter and mesotibia.
<emphasis id="B93EEAE02010FFEBFC9EDC2830A1F8E5" bold="true" box="[878,899,1880,1903]" pageId="6" pageNumber="53">G.</emphasis>
Mesotibia.
<emphasis id="B93EEAE02010FFEBFC0DDC293735F8E4" bold="true" box="[1021,1047,1881,1902]" pageId="6" pageNumber="53">H.</emphasis>
Apical portion of mesotibia, enlarged.
<emphasis id="B93EEAE02010FFEBFF67DC073385F806" bold="true" box="[151,167,1911,1932]" pageId="6" pageNumber="53">I.</emphasis>
Male sternite IX.
<emphasis id="B93EEAE02010FFEBFEAEDC073253F806" bold="true" box="[350,369,1911,1932]" pageId="6" pageNumber="53">J.</emphasis>
Aedeagus, in dorsal view.
<emphasis id="B93EEAE02010FFEBFD71DC0731B9F806" bold="true" box="[641,667,1911,1932]" pageId="6" pageNumber="53">K.</emphasis>
Same, in lateral view.
<emphasis id="B93EEAE02010FFEBFC70DC0730BAF806" bold="true" box="[896,920,1911,1932]" pageId="6" pageNumber="53">L.</emphasis>
Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05.
</paragraph>
</caption>
<paragraph id="8BF536F22011FFEAFF67DBE8323BFF5E" blockId="7.[151,416,151,212]" pageId="7" pageNumber="54">
<taxonomicName id="4C4A4D712011FFEAFF67DBE83282FF3A" authority="Raffray" authorityName="Raffray" box="[151,416,151,176]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B93EEAE02011FFEAFF67DBE83282FF3A" bold="true" box="[151,416,151,176]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF67DBE8321EFF3A" bold="true" box="[151,316,151,176]" italics="true" pageId="7" pageNumber="54">Tyrus sinensis</emphasis>
Raffray
</emphasis>
</taxonomicName>
(
<figureCitation id="13712A772011FFEAFF6FDBCC33FDFF5E" box="[159,223,188,212]" captionStart="FIGURE 1" captionStartId="1.[151,250,1960,1983]" captionTargetBox="[180,1402,786,1918]" captionTargetId="figure@1.[171,1408,769,1924]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Tyrus sichuanicus (A) and T. sinensis (B). Scales: 0.5 mm." httpUri="https://zenodo.org/record/283595/files/figure.png" pageId="7" pageNumber="54">Fig. 1</figureCitation>
B, 5)
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF67DA7530B4FE93" box="[151,918,259,282]" pageId="7" pageNumber="54" type="description">
<paragraph id="8BF536F22011FFEAFF67DA7530B4FE93" blockId="7.[151,918,259,282]" box="[151,918,259,282]" pageId="7" pageNumber="54">
<taxonomicName id="4C4A4D712011FFEAFF67DA7532CEFE93" authority="Raffray, 1912: 442" authorityName="Raffray" authorityPageNumber="442" authorityYear="1912" box="[151,492,259,282]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B93EEAE02011FFEAFF67DA753207FE90" box="[151,293,261,282]" italics="true" pageId="7" pageNumber="54">Tyrus sinensis</emphasis>
Raffray, 1912: 442
</taxonomicName>
.
<typeStatus id="54F188502011FFEAFE06DA75310AFE90" box="[502,552,261,282]" pageId="7" pageNumber="54">Type</typeStatus>
locality: Yunnan, Southwest
<collectingCountry id="F35D76622011FFEAFCA2DA7330B0FE93" box="[850,914,259,281]" name="China" pageId="7" pageNumber="54">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF67DA3631D2FE2C" pageId="7" pageNumber="54" type="materials_examined">
<paragraph id="8BF536F22011FFEAFF67DA3631D2FE2C" blockId="7.[151,1437,326,1143]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF67DA3632A3FED5" bold="true" box="[151,385,326,351]" pageId="7" pageNumber="54">Material examined</emphasis>
(1 3).
<emphasis id="B93EEAE02011FFEAFE1BDA363170FED5" bold="true" box="[491,594,326,351]" pageId="7" pageNumber="54">
<collectingCountry id="F35D76622011FFEAFE1BDA36316FFED5" box="[491,589,326,351]" name="China" pageId="7" pageNumber="54">CHINA</collectingCountry>
:
</emphasis>
3, labeled
<collectingCountry id="F35D76622011FFEAFD01DA363071FED4" box="[753,851,326,350]" name="China" pageId="7" pageNumber="54">CHINA</collectingCountry>
: Xizang, Mangkang Count. / Hongla Shan Mt., 29°12ʹ12ʹʹN / 98°39ʹ40ʹʹE, ca.
<quantity id="4CB29B172011FFEAFE18DA1A311CFE08" box="[488,574,362,386]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="54" unit="m" value="3700.0">3700 m</quantity>
, colony / of
<taxonomicName id="4C4A4D712011FFEAFD38DA1C300BFE09" box="[712,809,364,387]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02011FFEAFD38DA1C300BFE09" box="[712,809,364,387]" italics="true" pageId="7" pageNumber="54">Formica</emphasis>
</taxonomicName>
sp. under dead wood /
<date id="FFF410322011FFEAFBD9DA1A3788FE08" box="[1065,1194,362,386]" pageId="7" pageNumber="54" value="2011-07-25">25.vii.2011</date>
, Wen-Xuan Bi leg. [1 ex.
<taxonomicName id="4C4A4D712011FFEAFF4FDAFF323DFE2C" box="[191,287,399,422]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02011FFEAFF4FDAFF323DFE2C" box="[191,287,399,422]" italics="true" pageId="7" pageNumber="54">Formica</emphasis>
</taxonomicName>
sp. pinned under the
<typeStatus id="54F188502011FFEAFDE1DAFE315BFE2C" box="[529,633,398,422]" pageId="7" pageNumber="54" type="paratype">paratype</typeStatus>
] (SNUC).
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF39DAC237A1FCDC" pageId="7" pageNumber="54" type="diagnosis">
<paragraph id="8BF536F22011FFEAFF39DAC23652FE64" blockId="7.[151,1437,326,1143]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39DAC23261FE41" bold="true" box="[201,323,434,459]" pageId="7" pageNumber="54">Diagnosis.</emphasis>
Antennomeres IX obconical, X nearly trapezoidal, slightly transverse; ocular canthi absent; maxillary palpomeres IV nearly oval; mesotrochanters with two long spines; mesotibiae with short apical spur.
</paragraph>
<paragraph id="8BF536F22011FFEAFF39DA8A3118FCB8" blockId="7.[151,1437,326,1143]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39DA8A3254FD99" bold="true" box="[201,374,506,531]" pageId="7" pageNumber="54">Redescription.</emphasis>
Male (
<figureCitation id="13712A772011FFEAFE3ADA8A3129FD98" box="[458,523,506,530]" captionStart="FIGURE 1" captionStartId="1.[151,250,1960,1983]" captionTargetBox="[180,1402,786,1918]" captionTargetId="figure@1.[171,1408,769,1924]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Tyrus sichuanicus (A) and T. sinensis (B). Scales: 0.5 mm." httpUri="https://zenodo.org/record/283595/files/figure.png" pageId="7" pageNumber="54">Fig. 1</figureCitation>
B). Length
<quantity id="4CB29B172011FFEAFD62DA8A31D9FD98" box="[658,763,506,530]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="54" unit="mm" value="2.6">2.60 mm</quantity>
. Head about as long as wide, HL
<quantity id="4CB29B172011FFEAFB72DA8A37C8FD98" box="[1154,1258,506,530]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="54" unit="mm" value="0.4">0.40 mm</quantity>
, HW
<quantity id="4CB29B172011FFEAFADFDA8A36BAFD98" box="[1327,1432,506,530]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="7" pageNumber="54" unit="mm" value="0.41">0.41 mm</quantity>
; eyes each composed of about 45 facets. Antennae (
<figureCitation id="13712A772011FFEAFD27D96E303BFDBC" box="[727,793,542,566]" captionStart="FIGURE 5" captionStartId="6.[151,250,1849,1872]" captionTargetBox="[159,1430,457,1819]" captionTargetId="figure@6.[151,1436,447,1827]" captionTargetPageId="6" captionText="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
A) elongate, scapes slightly expanded laterally (
<figureCitation id="13712A772011FFEAFAC6D96E365BFDBC" box="[1334,1401,542,566]" captionStart="FIGURE 5" captionStartId="6.[151,250,1849,1872]" captionTargetBox="[159,1430,457,1819]" captionTargetId="figure@6.[151,1436,447,1827]" captionTargetPageId="6" captionText="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
C), clubs as in
<figureCitation id="13712A772011FFEAFED6D932324DFDD0" box="[294,367,578,602]" captionStart="FIGURE 5" captionStartId="6.[151,250,1849,1872]" captionTargetBox="[159,1430,457,1819]" captionTargetId="figure@6.[151,1436,447,1827]" captionTargetPageId="6" captionText="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
B. Maxillary palpi (
<figureCitation id="13712A772011FFEAFDAED9323187FDD0" box="[606,677,578,602]" captionStart="FIGURE 5" captionStartId="6.[151,250,1849,1872]" captionTargetBox="[159,1430,457,1819]" captionTargetId="figure@6.[151,1436,447,1827]" captionTargetPageId="6" captionText="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
D) with palpomeres I short, II lengthily pedunculate, III nearly triangular, IV oval. Pronotum about as long as wide, PL
<quantity id="4CB29B172011FFEAFCCAD9163086FDF4" box="[826,932,614,638]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="7" pageNumber="54" unit="mm" value="0.52">0.52 mm</quantity>
, PW
<quantity id="4CB29B172011FFEAFC19D9163771FDF4" box="[1001,1107,614,638]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="54" unit="mm" value="0.51">0.51 mm</quantity>
, with lateral margins nearly straight. Elytra wider than long, EL
<quantity id="4CB29B172011FFEAFDCCD9FA3184FD28" box="[572,678,650,674]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="7" pageNumber="54" unit="mm" value="0.68">0.68 mm</quantity>
, EW
<quantity id="4CB29B172011FFEAFD19D9FA3071FD28" box="[745,851,650,674]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" pageId="7" pageNumber="54" unit="mm" value="1.02">1.02 mm</quantity>
. Protrochanters with short triangular ventral spine, profemora with two blunt spines on ventral margin (
<figureCitation id="13712A772011FFEAFD08D9DE301CFD4C" box="[760,830,686,710]" captionStart="FIGURE 5" captionStartId="6.[151,250,1849,1872]" captionTargetBox="[159,1430,457,1819]" captionTargetId="figure@6.[151,1436,447,1827]" captionTargetPageId="6" captionText="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
E); mesotrochanters (
<figureCitation id="13712A772011FFEAFBC4D9DE375BFD4C" box="[1076,1145,686,710]" captionStart="FIGURE 5" captionStartId="6.[151,250,1849,1872]" captionTargetBox="[159,1430,457,1819]" captionTargetId="figure@6.[151,1436,447,1827]" captionTargetPageId="6" captionText="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
F) with two large ventral spines; mesotibiae (
<figureCitation id="13712A772011FFEAFE70D9A232E5FD60" box="[384,455,722,746]" captionStart="FIGURE 5" captionStartId="6.[151,250,1849,1872]" captionTargetBox="[159,1430,457,1819]" captionTargetId="figure@6.[151,1436,447,1827]" captionTargetPageId="6" captionText="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
G) strongly arched, with short apical spur (
<figureCitation id="13712A772011FFEAFC22D9A23739FD60" box="[978,1051,722,746]" captionStart="FIGURE 5" captionStartId="6.[151,250,1849,1872]" captionTargetBox="[159,1430,457,1819]" captionTargetId="figure@6.[151,1436,447,1827]" captionTargetPageId="6" captionText="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
H). Abdomen broad at base and narrowed apically, AL 1.00 mm, AW
<quantity id="4CB29B172011FFEAFD93D98631F3FC84" box="[611,721,758,782]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="7" pageNumber="54" unit="mm" value="1.08">1.08 mm</quantity>
. Sternite IX as in
<figureCitation id="13712A772011FFEAFC48D9863723FC84" box="[952,1025,758,782]" captionStart="FIGURE 5" captionStartId="6.[151,250,1849,1872]" captionTargetBox="[159,1430,457,1819]" captionTargetId="figure@6.[151,1436,447,1827]" captionTargetPageId="6" captionText="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="7" pageNumber="54">
Fig.
<date id="FFF410322011FFEAFC02D9863723FC84" box="[1010,1025,758,782]" pageId="7" pageNumber="54">5</date>
</figureCitation>
I. Aedeagus length
<quantity id="4CB29B172011FFEAFB1CD986367AFC84" box="[1260,1368,758,782]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="54" unit="mm" value="0.51">0.51 mm</quantity>
, with asymmetric median lobe (
<figureCitation id="13712A772011FFEAFE49D86A32DDFCB8" box="[441,511,794,818]" captionStart="FIGURE 5" captionStartId="6.[151,250,1849,1872]" captionTargetBox="[159,1430,457,1819]" captionTargetId="figure@6.[151,1436,447,1827]" captionTargetPageId="6" captionText="FIGURE 5. Diagnostic features of Tyrus sinensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283599/files/figure.png" pageId="7" pageNumber="54">Figs 5</figureCitation>
JL).
</paragraph>
<paragraph id="8BF536F22011FFEAFF39D84E37A1FCDC" blockId="7.[151,1437,326,1143]" box="[201,1155,830,854]" pageId="7" pageNumber="54">
Female. Not examined. Similar to male in general; BL
<quantity id="4CB29B172011FFEAFCC1D84E30B7FCDC" box="[817,917,830,854]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="54" unit="mm" value="2.4">2.40 mm</quantity>
(Raffray 1912: 443).
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF39D81232E6FC14" pageId="7" pageNumber="54" type="distribution">
<paragraph id="8BF536F22011FFEAFF39D8123752FCF0" blockId="7.[151,1437,326,1143]" box="[201,1136,866,891]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39D8123240FCF1" bold="true" box="[201,354,866,891]" pageId="7" pageNumber="54">Distribution.</emphasis>
Southwest
<collectingCountry id="F35D76622011FFEAFE15D812310EFCF0" box="[485,556,866,890]" name="China" pageId="7" pageNumber="54">China</collectingCountry>
: Yunnan, Xizang (= Tibet,
<emphasis id="B93EEAE02011FFEAFCAAD8133742FCF1" bold="true" box="[858,1120,866,891]" pageId="7" pageNumber="54">new provincial record</emphasis>
).
</paragraph>
<paragraph id="8BF536F22011FFEAFF39D8F732E6FC14" blockId="7.[151,1437,326,1143]" box="[201,452,903,927]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39D8F73214FC15" bold="true" box="[201,310,903,927]" pageId="7" pageNumber="54">Host ant.</emphasis>
<taxonomicName id="4C4A4D712011FFEAFECCD8F832BEFC15" box="[316,412,904,927]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02011FFEAFECCD8F832BEFC15" box="[316,412,904,927]" italics="true" pageId="7" pageNumber="54">Formica</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF39D8DA329CFC6C" pageId="7" pageNumber="54" type="biology_ecology">
<paragraph id="8BF536F22011FFEAFF39D8DA329CFC6C" blockId="7.[151,1437,326,1143]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39D8DA320BFC49" bold="true" box="[201,297,938,963]" pageId="7" pageNumber="54">Biology.</emphasis>
The single male was collected from a colony of
<taxonomicName id="4C4A4D712011FFEAFCBFD8DC308DFC49" box="[847,943,940,963]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAE02011FFEAFCBFD8DC308DFC49" box="[847,943,940,963]" italics="true" pageId="7" pageNumber="54">Formica</emphasis>
</taxonomicName>
sp. nesting on the ground under a dead tree (Bi per. comm.) (
<figureCitation id="13712A772011FFEAFEA7D8BE32B9FC6C" box="[343,411,974,998]" captionStart="FIGURE 7" captionStartId="9.[151,250,1910,1933]" captionTargetBox="[151,1435,283,1888]" captionTargetId="figure@9.[151,1436,283,1889]" captionTargetPageId="9" captionText="FIGURE 7. Habitat of Tyrus sichuanicus and T. sinensis. A. Collection site of T. sichuanicus — Chaheba Mt., alt. 2615 m. B. Peng &amp; Yin investigated a nest of Formica sp. at Chaheba Mt. C. A closer view of the nest. D. Collection site of T. sinensis — Hongla Shan Mt., alt. 3700 m (photo by Wen-Xuan Bi)." httpUri="https://zenodo.org/record/283601/files/figure.png" pageId="7" pageNumber="54">Fig. 7</figureCitation>
D).
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF39D88233C9FA8B" pageId="7" pageNumber="54" type="discussion">
<paragraph id="8BF536F22011FFEAFF39D88236B9FBFC" blockId="7.[151,1437,326,1143]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39D8823273FB81" bold="true" box="[201,337,1010,1035]" pageId="7" pageNumber="54">Comments.</emphasis>
The
<typeStatus id="54F188502011FFEAFE7CD883329EFB81" box="[396,444,1011,1035]" pageId="7" pageNumber="54">type</typeStatus>
material of
<taxonomicName id="4C4A4D712011FFEAFDB8D8843199FB81" box="[584,699,1012,1035]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B93EEAE02011FFEAFDB8D8843199FB81" box="[584,699,1012,1035]" italics="true" pageId="7" pageNumber="54">T. sinensis</emphasis>
</taxonomicName>
(a female) was not examined, but the form of the antennomeres, the simple postgenae, and the nearly oval maxillary palpomeres IV of the single male from the Hongla-Shan Mountain highly agree with the descriptive details provided by Raffray (1912: 443), in combination with the fact that the collection site of this male is near the border of Xizang and Yunnan, it is therefore identified as
<taxonomicName id="4C4A4D712011FFEAFAD0DF1036B0FBFD" box="[1312,1426,1120,1143]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B93EEAE02011FFEAFAD0DF1036B0FBFD" box="[1312,1426,1120,1143]" italics="true" pageId="7" pageNumber="54">T. sinensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF536F22011FFEAFF67DFB633C9FA8B" blockId="7.[151,653,1221,1281]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF67DFB631AFFB54" bold="true" box="[151,653,1221,1246]" pageId="7" pageNumber="54">
<taxonomicName id="4C4A4D712011FFEAFF67DFB63283FB54" authority="Yin" authorityName="Yin" box="[151,417,1221,1246]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="yajiangensis">
<emphasis id="B93EEAE02011FFEAFF67DFB6324DFB54" bold="true" box="[151,367,1221,1246]" italics="true" pageId="7" pageNumber="54">Tyrus yajiangensis</emphasis>
Yin
</taxonomicName>
and Li, new species
</emphasis>
(
<figureCitation id="13712A772011FFEAFF6FDF9933C1FA8B" box="[159,227,1257,1281]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF67DE4232A7FA1B" pageId="7" pageNumber="54" type="materials_examined">
<paragraph id="8BF536F22011FFEAFF67DE4232A7FA1B" blockId="7.[151,1436,1329,1930]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF67DE42321FFAC0" bold="true" box="[151,317,1329,1354]" pageId="7" pageNumber="54">
<typeStatus id="54F188502011FFEAFF67DE4233F0FAC0" box="[151,210,1330,1354]" pageId="7" pageNumber="54">Type</typeStatus>
material
</emphasis>
(1 3).
<emphasis id="B93EEAE02011FFEAFE7EDE423153FAC0" bold="true" box="[398,625,1329,1354]" pageId="7" pageNumber="54">
<typeStatus id="54F188502011FFEAFE7EDE4232DDFAC0" box="[398,511,1330,1354]" pageId="7" pageNumber="54" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F35D76622011FFEAFDFADE41314EFAC0" box="[522,620,1329,1354]" name="China" pageId="7" pageNumber="54">CHINA</collectingCountry>
:
</emphasis>
3, labeled
<collectingCountry id="F35D76622011FFEAFD07DE41307BFAC3" box="[759,857,1329,1353]" name="China" pageId="7" pageNumber="54">CHINA</collectingCountry>
: W-Sichuan 1999 / Ganzi, Tibet. Aut. Pref., Yajiang Co. / Shalui Shan, Bachtal
<quantity id="4CB29B172011FFEAFE33DE2532D9FAE7" box="[451,507,1365,1389]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="54" unit="km" value="6.0">6 km</quantity>
WSW / Yajiang,
<quantity id="4CB29B172011FFEAFD4EDE253036FAE7" box="[702,788,1365,1389]" metricMagnitude="3" metricUnit="m" metricValue="3.25" pageId="7" pageNumber="54" unit="m" value="3250.0">3250 m</quantity>
, 30°01ʹN / 100°57ʹE, Laubstreu, Rinde, Pilze /
<date id="FFF410322011FFEAFAD2DE25364AFAE7" box="[1314,1384,1365,1389]" pageId="7" pageNumber="54">4.VII.</date>
, leg. M. Schülke (pcMS).
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF39DEED37DFFA53" pageId="7" pageNumber="54" type="diagnosis">
<paragraph id="8BF536F22011FFEAFF39DEED37DFFA53" blockId="7.[151,1436,1329,1930]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39DEED3261FA3C" bold="true" box="[201,323,1437,1462]" pageId="7" pageNumber="54">Diagnosis.</emphasis>
Antennomeres IXX each elongate, narrowed apically; ocular canthi present; maxillary palpomeres IV nearly oval; mesotrochanters with two long spines; mesotibiae with short apical spur.
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF39DE9532BDF897" pageId="7" pageNumber="54" type="description">
<paragraph id="8BF536F22011FFEAFF39DE9537D4F973" blockId="7.[151,1436,1329,1930]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39DE953278FA74" bold="true" box="[201,346,1509,1534]" pageId="7" pageNumber="54">Description.</emphasis>
Male. Length
<quantity id="4CB29B172011FFEAFDF8DE953150FA77" box="[520,626,1509,1533]" metricMagnitude="-3" metricUnit="m" metricValue="2.48" pageId="7" pageNumber="54" unit="mm" value="2.48">2.48 mm</quantity>
. Head about as long as wide, HL
<quantity id="4CB29B172011FFEAFBF3DE95374EFA77" box="[1027,1132,1509,1533]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="7" pageNumber="54" unit="mm" value="0.52">0.52 mm</quantity>
, HW
<quantity id="4CB29B172011FFEAFB44DE95363CFA77" box="[1204,1310,1509,1533]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="54" unit="mm" value="0.5">0.50 mm</quantity>
; eyes each composed of about 45 facets. Antennae (
<figureCitation id="13712A772011FFEAFDACDD7931BCF9AB" box="[604,670,1545,1569]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
A) elongate, scapes expanded laterally (
<figureCitation id="13712A772011FFEAFBABDD7937BFF9AB" box="[1115,1181,1545,1569]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
C), clubs as in
<figureCitation id="13712A772011FFEAFAB1DD7936A1F9AB" box="[1345,1411,1545,1569]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
B. Maxillary palpi (
<figureCitation id="13712A772011FFEAFEA8DD5D32BEF9CF" box="[344,412,1581,1605]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
D) with palpomeres I short, II lengthily pedunculate, III nearly triangular, IV nearly oval. Pronotum about as long as wide, PL
<quantity id="4CB29B172011FFEAFDC2DD2131BBF9E3" box="[562,665,1617,1641]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="7" pageNumber="54" unit="mm" value="0.52">0.52 mm</quantity>
, PW
<quantity id="4CB29B172011FFEAFD24DD213019F9E3" box="[724,827,1617,1641]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="7" pageNumber="54" unit="mm" value="0.54">0.54 mm</quantity>
, with lateral margins nearly straight. Elytra wider than long, EL
<quantity id="4CB29B172011FFEAFEF0DD053244F907" box="[256,358,1653,1677]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="7" pageNumber="54" unit="mm" value="0.68">0.68 mm</quantity>
, EW
<quantity id="4CB29B172011FFEAFE53DD053128F907" box="[419,522,1653,1677]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" pageId="7" pageNumber="54" unit="mm" value="1.07">1.07 mm</quantity>
. Protrochanters triangularly projecting ventrally, profemora with two blunt spines on ventral margin (
<figureCitation id="13712A772011FFEAFE9EDDE93293F93B" box="[366,433,1689,1713]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
E); mesotrochanters (
<figureCitation id="13712A772011FFEAFD51DDE931C6F93B" box="[673,740,1689,1713]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
F) with two large ventral spines; mesotibiae (
<figureCitation id="13712A772011FFEAFB10DDE93601F93B" box="[1248,1315,1689,1713]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
G) slightly arched, with short apical spur (
<figureCitation id="13712A772011FFEAFE0BDDCD3163F95F" box="[507,577,1725,1749]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
H). Abdomen broad at base and narrowed apically, AL
<quantity id="4CB29B172011FFEAFB30DDCD360AF95F" box="[1216,1320,1725,1749]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="54" unit="mm" value="0.76">0.76 mm</quantity>
, AW 1.00 mm. Sternite IX as in
<figureCitation id="13712A772011FFEAFE7DDD9132F3F973" box="[397,465,1761,1785]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">
Fig.
<date id="FFF410322011FFEAFE31DD9132F3F973" box="[449,465,1761,1785]" pageId="7" pageNumber="54">6</date>
</figureCitation>
I. Aedeagus length
<quantity id="4CB29B172011FFEAFD58DD91302CF973" box="[680,782,1761,1785]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="54" unit="mm" value="0.45">0.45 mm</quantity>
, with asymmetric median lobe (
<figureCitation id="13712A772011FFEAFB84DD913799F973" box="[1140,1211,1761,1785]" captionStart="FIGURE 6" captionStartId="8.[151,250,1637,1660]" captionTargetBox="[166,1419,205,1606]" captionTargetId="figure@8.[151,1436,195,1616]" captionTargetPageId="8" captionText="FIGURE 6. Diagnostic features of Tyrus yajiangensis. A. Antenna. B. antennal club, enlarged. C. Scape, enlarged. D. Maxillary palpus. E. Protrochanter and protibia. F. Mesotrochanter and mesotibia. G. Mesotibia. H. Apical portion of mesotibia, enlarged. I. Male sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): A = 0.5; B, E, F, G, J, K, L = 0.2; C, D, I = 0.1; H = 0.05." httpUri="https://zenodo.org/record/283600/files/figure.png" pageId="7" pageNumber="54">Figs 6</figureCitation>
JL).
</paragraph>
<paragraph id="8BF536F22011FFEAFF39DC7532BDF897" blockId="7.[151,1436,1329,1930]" box="[201,415,1797,1821]" pageId="7" pageNumber="54">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF39DC5931BAF8CB" box="[201,664,1833,1858]" pageId="7" pageNumber="54" type="distribution">
<paragraph id="8BF536F22011FFEAFF39DC5931BAF8CB" blockId="7.[151,1436,1329,1930]" box="[201,664,1833,1858]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39DC593240F8C8" bold="true" box="[201,354,1833,1858]" pageId="7" pageNumber="54">Distribution.</emphasis>
Southwest
<collectingCountry id="F35D76622011FFEAFE15DC59310EF8CB" box="[485,556,1833,1857]" name="China" pageId="7" pageNumber="54">China</collectingCountry>
: Sichuan.
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF39DC3D3719F8EF" box="[201,1083,1869,1894]" pageId="7" pageNumber="54" type="biology_ecology">
<paragraph id="8BF536F22011FFEAFF39DC3D3719F8EF" blockId="7.[151,1436,1329,1930]" box="[201,1083,1869,1894]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39DC3D320BF8EC" bold="true" box="[201,297,1869,1894]" pageId="7" pageNumber="54">Biology.</emphasis>
The adult was possibly taken from sifted leaf litter or under tree bark.
</paragraph>
</subSubSection>
<subSubSection id="C35065792011FFEAFF39DC023054F803" box="[201,886,1905,1930]" pageId="7" pageNumber="54" type="etymology">
<paragraph id="8BF536F22011FFEAFF39DC023054F803" blockId="7.[151,1436,1329,1930]" box="[201,886,1905,1930]" pageId="7" pageNumber="54">
<emphasis id="B93EEAE02011FFEAFF39DC02326DF800" bold="true" box="[201,335,1906,1930]" pageId="7" pageNumber="54">Etymology.</emphasis>
The new species is named after the
<typeStatus id="54F188502011FFEAFD15DC023037F800" box="[741,789,1906,1930]" pageId="7" pageNumber="54">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>