837 lines
108 KiB
XML
837 lines
108 KiB
XML
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<mods:title id="6EC9D5103F5B1B96DF59BCB4193D29A0">Revision of the South American genera Andinocopris new genus and Homocopris Burmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae: Homocoprini new tribe)</mods:title>
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<mods:namePart id="B6C2AC50A17B4E97D080807C75B1F1D2">Génier, François</mods:namePart>
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<mods:affiliation id="F055F2945B58CF7A032CAC421CE05F33">Beaty Center for Species Discovery, Canadian Museum of Nature, 1740 Pink Road, Gatineau, Québec, J 9 J 3 N 7, Canada.</mods:affiliation>
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<mods:namePart id="AD95AF65044007CBA09B36AF4E7A51BE">Darling, James D. G.</mods:namePart>
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<mods:affiliation id="251448BD5C8EAF4716DA05F0E199C19E">Beaty Center for Species Discovery, Canadian Museum of Nature, 1740 Pink Road, Gatineau, Québec, J 9 J 3 N 7, Canada.</mods:affiliation>
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<mods:nameIdentifier id="C794463D3AA72C914B8A5ADBCDA4B50A" type="email">james21darling@gmail.com</mods:nameIdentifier>
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<mods:title id="42E2EA1F822858E9FD774BCF5371467A">European Journal of Taxonomy</mods:title>
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<mods:date id="BD280CD6D13D61AABB6CAAC4E6811256">2024</mods:date>
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<treatment id="03DC87B7EF2DFFB7FE63FCCACDF3E3C0" ID-DOI="http://doi.org/10.5281/zenodo.10845101" ID-GBIF-Taxon="224814833" ID-Zenodo-Dep="10845101" LSID="urn:lsid:plazi:treatment:03DC87B7EF2DFFB7FE63FCCACDF3E3C0" httpUri="http://treatment.plazi.org/id/03DC87B7EF2DFFB7FE63FCCACDF3E3C0" lastPageId="9" lastPageNumber="232" pageId="8" pageNumber="228">
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<subSubSection id="C36F652AEF2DFFB6FE63FCCACDB3E107" box="[428,1160,853,880]" pageId="8" pageNumber="228" type="nomenclature">
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<paragraph id="8BCA36A1EF2DFFB6FE63FCCACDB3E107" blockId="8.[428,1160,853,915]" box="[428,1160,853,880]" pageId="8" pageNumber="228">
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<heading id="D08281CDEF2DFFB6FE63FCCACDB3E107" box="[428,1160,853,880]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="228" reason="2">
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<taxonomicName id="4C754D22EF2DFFB6FE63FCCACA91E118" authority="(Waterhouse, 1891)" authorityName="Génier & Darling" authorityYear="2024" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[428,938,853,880]" class="Insecta" family="Scarabaeidae" genus="Andinocopris" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi" status="gen. et comb. nov.">
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<emphasis id="B901EAB3EF2DFFB6FE63FCCACB87E107" bold="true" box="[428,700,853,880]" italics="true" pageId="8" pageNumber="228">Andinocopris buckleyi</emphasis>
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(
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<bibRefCitation id="EFE44B50EF2DFFB6FD05FCCACA9BE118" author="Waterhouse C. O." box="[714,928,853,880]" pageId="8" pageNumber="228" pagination="348 - 363" refId="ref28366" refString="Waterhouse C. O. 1891. New Scarabaeidae in the British Museum: a fourth contribution. The Annals and Magazine of Natural History, including Zoology, Botany, and Geology. Sixth Series 7 (40): 348 - 363. https: // doi. org / 10.1080 / 00222939109460622" type="journal article" year="1891">Waterhouse, 1891</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A23257C8EF2DFFB6FC7EFCC9CDB3E107" box="[945,1160,853,880]" pageId="8" pageNumber="228" rank="species">gen. et comb. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F652AEF2DFFB6FDF8FCE7CAC7E1E4" box="[567,1020,888,915]" pageId="8" pageNumber="228" type="description">
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<paragraph id="8BCA36A1EF2DFFB6FDF8FCE7CAC7E1E4" blockId="8.[428,1160,853,915]" box="[567,1020,888,915]" pageId="8" pageNumber="228">
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<figureCitation id="134E2A24EF2DFFB6FDF8FCE7CBA4E1E5" box="[567,671,888,914]" captionStart="Figs 1–4" captionStartId="6.[189,242,1892,1918]" captionTargetBox="[234,1360,273,1801]" captionTargetId="figure-12@6.[218,1370,265,1852]" captionTargetPageId="6" captionText="Figs 1–4. Dorsal habitus. 1. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 2. Andinocopris achamas, alloreferent paralectotype, ♀ (MNHN). 3. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 4. Andinocopris buckleyi, specimen ♀ (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.10824970" httpUri="https://zenodo.org/record/10824970/files/figure.png" pageId="8" pageNumber="228">Figs 3–4</figureCitation>
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,
|
||
<figureCitation id="134E2A24EF2DFFB6FD64FCE7CBC3E1E4" box="[683,760,888,915]" captionStart="Figs13–16" captionStartId="14.[189,242,1891,1917]" captionTargetBox="[221,1346,305,1841]" captionTargetId="figure-12@14.[221,1366,263,1841]" captionTargetPageId="14" captionText="Figs13–16.Headandpronotum,lateralview.13. Andinocoprisachamas(Harold,1867)gen.etcomb.nov., lectotype, ♂ (MNHN). 14. Andinocopris achamas, paralectotype, ♀ (MNHN). 15. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 16. Andinocopris buckleyi, specimen ♀ (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.10824978" httpUri="https://zenodo.org/record/10824978/files/figure.png" pageId="8" pageNumber="228">15–16</figureCitation>
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,
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||
<figureCitation id="134E2A24EF2DFFB6FCCCFCE7CA18E1E5" box="[771,803,888,914]" captionStart="Figs25–30" captionStartId="19.[189,242,1857,1883]" captionTargetBox="[239,1354,353,1767]" captionTargetId="figure-17@19.[234,1354,265,1809]" captionTargetPageId="19" captionText="Figs25–30. Paramere,lateralview.25. Andinocoprisachamas(Harold,1867)gen.etcomb.nov.,specimen ♂ (CMNC). 26. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 27. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 28. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 29. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 30. Homocopris williami Darling & Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824984" httpUri="https://zenodo.org/record/10824984/files/figure.png" pageId="8" pageNumber="228">26</figureCitation>
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,
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||
<figureCitation id="134E2A24EF2DFFB6FCE0FCE7CA74E1E5" box="[815,847,888,914]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling & Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="8" pageNumber="228">32</figureCitation>
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,
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<figureCitation id="134E2A24EF2DFFB6FC94FCE7CA40E1E5" box="[859,891,888,914]" captionStart="Figs 37–41. 37–38" captionStartId="23.[189,242,1892,1918]" captionTargetBox="[257,1314,294,1816]" captionTargetId="figure-17@23.[221,1366,265,1843]" captionTargetPageId="23" captionText="Figs 37–41. 37–38. Genital segment, dorsal view. 37. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 38. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 39–41. Hindwing, dorsal view. 39. Andinocopris achamas (Harold, 1867) gen. et comb. nov. 40. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov. 41. Homocopris torulosus." figureDoi="http://doi.org/10.5281/zenodo.10824988" httpUri="https://zenodo.org/record/10824988/files/figure.png" pageId="8" pageNumber="228">40</figureCitation>
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,
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<figureCitation id="134E2A24EF2DFFB6FC48FCE7CA9CE1E5" box="[903,935,888,914]" captionStart="Figs 42–45" captionStartId="25.[189,242,1822,1848]" captionTargetBox="[232,1356,285,1784]" captionTargetId="figure-18@25.[232,1356,265,1787]" captionTargetPageId="25" captionText="Figs 42–45. Metasternum, ventral view. In specimens with a metasternal depression, the lateral depression is bordered with white dotted lines. For Homocopris grossiorum Darling & Génier sp. nov., a red dotted line indicates where the metasternal depression is interrupted. 42. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♀ (MNHN) 43. Homocopris grossiorum, holotype ♂ (MZSP). 44. Homocopris punctatissimus (Curtis, 1845), specimen ♀ (NMPC). 45. Homocopris torulosus (Eschscholtz, 1822), specimen ♂ (CMNC)." figureDoi="http://doi.org/10.5281/zenodo.10824990" httpUri="https://zenodo.org/record/10824990/files/figure.png" pageId="8" pageNumber="228">42</figureCitation>
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,
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<figureCitation id="134E2A24EF2DFFB6FC7CFCE7CAE8E1E4" box="[947,979,888,915]" captionStart="Figs 46–57. 46–49" captionStartId="29.[189,242,1787,1813]" captionTargetBox="[278,1306,298,1712]" captionTargetId="figure-17@29.[258,1330,265,1743]" captionTargetPageId="29" captionText="Figs 46–57. 46–49. Elytral striae, posterolateral view. 46. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 47. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). Anterior pronotal surface. 48. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 49. Homocopris torulosus, neotype, ♂ (CMNC). 50–57. Type specimen labels. 50. Pinotus achamas Harold, 1867. 51. Pinotus buckleyi (Waterhouse, 1891). 52. Homocopris grossiorum. 53. Copris punctatissimus (Curtis, 1845). 54. Copris torulosus Eschscholtz, 1822. 55. Homocopris williami Darling & Génier sp. nov. 56. Pinotus simulator Luederwaldt, 1936. 57. Pinotus dahli Landin, 1955." figureDoi="http://doi.org/10.5281/zenodo.10824992" httpUri="https://zenodo.org/record/10824992/files/figure.png" pageId="8" pageNumber="228">51</figureCitation>
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,
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<figureCitation id="134E2A24EF2DFFB6FC10FCE7CAC7E1E4" box="[991,1020,888,915]" captionStart="Fig" captionStartId="36.[189,232,1922,1948]" captionTargetBox="[189,1398,975,1873]" captionTargetId="figure-182@36.[189,1398,975,1876]" captionTargetPageId="36" captionText="Fig. 65. Andinocopris achamas (Harold, 1867) gen. et comb. nov. and Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., distribution map. Legend and scale bar included in the top and bottom right corners, respectively." figureDoi="http://doi.org/10.5281/zenodo.10824996" httpUri="https://zenodo.org/record/10824996/files/figure.png" pageId="8" pageNumber="228">65</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F652AEF2DFFB6FF72FC24CAB1E758" pageId="8" pageNumber="228" type="reference_group">
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<paragraph id="8BCA36A1EF2DFFB6FF72FC24CA9AE1A3" blockId="8.[189,970,954,1016]" box="[189,929,954,981]" pageId="8" pageNumber="228">
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<treatmentCitationGroup id="AB65118FEF2DFFB6FF72FC24CA9AE1A3" box="[189,929,954,981]" pageId="8" pageNumber="228">
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<treatmentCitation id="0AD410B0EF2DFFB6FF72FC24CBA2E1A2" author="Waterhouse C. O." box="[189,665,954,981]" page="359" pageId="8" pageNumber="228" year="1891">
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<taxonomicName id="4C754D22EF2DFFB6FF72FC24CBA2E1A2" ID-CoL="4HZTH" authority="Waterhouse, 1891: 359" authorityName="Waterhouse" authorityPageNumber="359" authorityYear="1891" box="[189,665,954,981]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi">
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<emphasis id="B901EAB3EF2DFFB6FF72FC24C8B8E1A3" box="[189,387,954,980]" italics="true" pageId="8" pageNumber="228">Pinotus Buckleyi</emphasis>
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<bibRefCitation id="EFE44B50EF2DFFB6FE45FC25CBA2E1A2" author="Waterhouse C. O." box="[394,665,954,981]" pageId="8" pageNumber="228" pagination="348 - 363" refId="ref28366" refString="Waterhouse C. O. 1891. New Scarabaeidae in the British Museum: a fourth contribution. The Annals and Magazine of Natural History, including Zoology, Botany, and Geology. Sixth Series 7 (40): 348 - 363. https: // doi. org / 10.1080 / 00222939109460622" type="journal article" year="1891">Waterhouse, 1891: 359</bibRefCitation>
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</taxonomicName>
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(original description).
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</paragraph>
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<paragraph id="8BCA36A1EF2DFFB6FF72FC41CAF1E180" blockId="8.[189,970,954,1016]" box="[189,970,989,1016]" pageId="8" pageNumber="228">
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<treatmentCitationGroup id="AB65118FEF2DFFB6FF72FC41CAF1E180" box="[189,970,989,1016]" pageId="8" pageNumber="228">
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[
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<taxonomicName id="4C754D22EF2DFFB6FF08FC41C81AE180" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[199,289,990,1015]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="genus">
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<emphasis id="B901EAB3EF2DFFB6FF08FC41C81AE180" box="[199,289,990,1015]" italics="true" pageId="8" pageNumber="228">Pinotus</emphasis>
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]
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<treatmentCitation id="0AD410B0EF2DFFB6FEFDFC42CBF8E180" author="Luederwaldt H." box="[306,707,989,1016]" page="209" pageId="8" pageNumber="228" year="1936">
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<taxonomicName id="4C754D22EF2DFFB6FEFDFC42CBF8E180" authority="Luederwaldt, 1936: 209" authorityName=", Luederwaldt" authorityPageNumber="209" authorityYear="1936" box="[306,707,989,1016]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="simulator">
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<emphasis id="B901EAB3EF2DFFB6FEFDFC42C899E180" box="[306,418,989,1015]" italics="true" pageId="8" pageNumber="228">simulator</emphasis>
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<bibRefCitation id="EFE44B50EF2DFFB6FE66FC42CBF8E180" author="Luederwaldt H." box="[425,707,989,1016]" pageId="8" pageNumber="228" pagination="207 - 216" refId="ref26756" refString="Luederwaldt H. 1936. Notas complementares " As especies brasileiras do genero Pinotus ". Revista do Museu Paulista 20: 207 - 216." type="journal article" year="1936">Luederwaldt, 1936: 209</bibRefCitation>
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</taxonomicName>
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(original description).
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</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72FB80CBDFE64F" blockId="8.[189,1398,1054,1327]" box="[189,740,1054,1081]" pageId="8" pageNumber="228">
|
||
<treatmentCitationGroup id="AB65118FEF2DFFB6FF72FB80CBDFE64F" box="[189,740,1054,1081]" pageId="8" pageNumber="228">
|
||
<taxonomicName id="4C754D22EF2DFFB6FF72FB80C8B8E64F" ID-CoL="4HZTH" authorityName="Waterhouse" authorityYear="1891" box="[189,387,1054,1080]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72FB80C8B8E64F" box="[189,387,1054,1080]" italics="true" pageId="8" pageNumber="228">Pinotus Buckleyi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AD410B0EF2DFFB6FE6FFB81CB6AE64E" author="Gillet J. J. E." box="[416,593,1054,1081]" page="59" pageId="8" pageNumber="228" year="1911">
|
||
<bibRefCitation id="EFE44B50EF2DFFB6FE6FFB81CB6AE64E" author="Gillet J. J. E." box="[416,593,1054,1081]" pageId="8" pageNumber="228" pagination="38" refId="ref25749" refString="Gillet J. J. E. 1911. Scarabaeidae: Coprinae I. In: Junk W. & Schenkling S. (eds) Coleopterorum Catalogus. Pars 38. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accessed 6 Feb. 2024]." type="book chapter" year="1911">Gillet 1911: 59</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogue).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72FBDDCAB0E62B" blockId="8.[189,1398,1054,1327]" box="[189,907,1089,1116]" pageId="8" pageNumber="228">
|
||
<treatmentCitationGroup id="AB65118FEF2DFFB6FF72FBDDCAB0E62B" box="[189,907,1089,1116]" pageId="8" pageNumber="228">
|
||
<taxonomicName id="4C754D22EF2DFFB6FF72FBDDC888E62B" ID-CoL="4HZTH" authorityName="Waterhouse" authorityYear="1891" box="[189,435,1090,1116]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi" subGenus="P.">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72FBDDC82CE62C" box="[189,279,1090,1115]" italics="true" pageId="8" pageNumber="228">Pinotus</emphasis>
|
||
(
|
||
<emphasis id="B901EAB3EF2DFFB6FEE7FBDDC805E62C" box="[296,318,1090,1115]" italics="true" pageId="8" pageNumber="228">P.</emphasis>
|
||
)
|
||
<emphasis id="B901EAB3EF2DFFB6FE81FBDDC888E62B" box="[334,435,1090,1116]" italics="true" pageId="8" pageNumber="228">Buckleyi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AD410B0EF2DFFB6FE1EFBDDCBD8E62B" author="Luederwaldt H." box="[465,739,1089,1116]" page="652" pageId="8" pageNumber="228" year="1929">
|
||
<bibRefCitation id="EFE44B50EF2DFFB6FE1EFBDDCBD8E62B" author="Luederwaldt H." box="[465,739,1089,1116]" pageId="8" pageNumber="228" pagination="603 - 775" refId="ref26718" refString="Luederwaldt H. 1929. As especies b rasileiras do genero Pinotus (Coleoptera - Lamellicornidae - Coprini), com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929: 652</bibRefCitation>
|
||
</treatmentCitation>
|
||
(monograph).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72FBF9CD3CE608" blockId="8.[189,1398,1054,1327]" box="[189,1031,1125,1151]" pageId="8" pageNumber="228">
|
||
<treatmentCitationGroup id="AB65118FEF2DFFB6FF72FBF9CD3CE608" box="[189,1031,1125,1151]" pageId="8" pageNumber="228">
|
||
<taxonomicName id="4C754D22EF2DFFB6FF72FBF9C88BE608" ID-CoL="4HZTH" authorityName="Waterhouse" authorityYear="1891" box="[189,432,1125,1151]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi" subGenus="P.">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72FBF9C82CE608" box="[189,279,1126,1151]" italics="true" pageId="8" pageNumber="228">Pinotus</emphasis>
|
||
(
|
||
<emphasis id="B901EAB3EF2DFFB6FEE7FBF9C805E608" box="[296,318,1126,1151]" italics="true" pageId="8" pageNumber="228">P.</emphasis>
|
||
)
|
||
<emphasis id="B901EAB3EF2DFFB6FE81FBFAC88BE608" box="[334,432,1125,1151]" italics="true" pageId="8" pageNumber="228">buckleyi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AD410B0EF2DFFB6FE01FBFACBE4E608" author="Luederwaldt H." box="[462,735,1125,1151]" page="208" pageId="8" pageNumber="228" year="1936">
|
||
<bibRefCitation id="EFE44B50EF2DFFB6FE01FBFACBE4E608" author="Luederwaldt H." box="[462,735,1125,1151]" pageId="8" pageNumber="228" pagination="207 - 216" refId="ref26756" refString="Luederwaldt H. 1936. Notas complementares " As especies brasileiras do genero Pinotus ". Revista do Museu Paulista 20: 207 - 216." type="journal article" year="1936">Luederwaldt 1936: 208</bibRefCitation>
|
||
</treatmentCitation>
|
||
(diagnosis, distribution).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72FB16CA01E6D5" blockId="8.[189,1398,1054,1327]" box="[189,826,1160,1187]" pageId="8" pageNumber="228">
|
||
<treatmentCitationGroup id="AB65118FEF2DFFB6FF72FB16CA01E6D5" box="[189,826,1160,1187]" pageId="8" pageNumber="228">
|
||
<taxonomicName id="4C754D22EF2DFFB6FF72FB16C8BBE6D5" ID-CoL="4HZTH" authorityName="Waterhouse" authorityYear="1891" box="[189,384,1160,1186]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72FB16C8BBE6D5" box="[189,384,1160,1186]" italics="true" pageId="8" pageNumber="228">Pinotus buckleyi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AD410B0EF2DFFB6FE52FB17CB94E6D5" author="Blackwelder R. E." box="[413,687,1160,1186]" page="206" pageId="8" pageNumber="228" year="1944">
|
||
<bibRefCitation id="EFE44B50EF2DFFB6FE52FB17CB94E6D5" author="Blackwelder R. E." box="[413,687,1160,1186]" pageId="8" pageNumber="228" pagination="189 - 341" refId="ref24818" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
|
||
</treatmentCitation>
|
||
(checklist).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72FB34CDFFE6B2" blockId="8.[189,1398,1054,1327]" box="[189,1220,1195,1222]" pageId="8" pageNumber="228">
|
||
<treatmentCitationGroup id="AB65118FEF2DFFB6FF72FB34CDFFE6B2" box="[189,1220,1195,1222]" pageId="8" pageNumber="228">
|
||
<taxonomicName id="4C754D22EF2DFFB6FF72FB34C8F1E6B2" box="[189,458,1195,1221]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="simulator">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72FB34C8F1E6B2" box="[189,458,1195,1221]" italics="true" pageId="8" pageNumber="228">Dichotomius simulator</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AD410B0EF2DFFB6FE27FB34CA04E6B1" author="Vulcano M. A. & Pereira F. S." box="[488,831,1195,1222]" page="584" pageId="8" pageNumber="228" year="1967">
|
||
<bibRefCitation id="EFE44B50EF2DFFB6FE27FB34CA04E6B1" author="Vulcano M. A. & Pereira F. S." box="[488,831,1195,1222]" pageId="8" pageNumber="228" pagination="533 - 603" refId="ref28319" refString="Vulcano M. A. & Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica (Zoologia) 5: 533 - 603." type="journal article" year="1967">Vulcano & Pereira 1967: 584</bibRefCitation>
|
||
</treatmentCitation>
|
||
(distribution, identification key).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72FB50CA2EE77B" blockId="8.[189,1398,1054,1327]" pageId="8" pageNumber="228">
|
||
<treatmentCitationGroup id="AB65118FEF2DFFB6FF72FB50CA2EE77B" pageId="8" pageNumber="228">
|
||
<taxonomicName id="4C754D22EF2DFFB6FF72FB50C883E69F" ID-CoL="3MFLJ" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[189,440,1230,1256]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72FB50C883E69F" box="[189,440,1230,1256]" italics="true" pageId="8" pageNumber="228">Homocopris buckleyi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AD410B0EF2DFFB6FE1BFB51CA0EE69E" author="Vaz-de-Mello F. Z. & Genier F. & Smith A. B. T." box="[468,821,1230,1257]" page="192" pageId="8" pageNumber="228" year="2010">
|
||
<bibRefCitation id="EFE44B50EF2DFFB6FE1BFB51CA0EE69E" author="Vaz-de-Mello F. Z. & Genier F. & Smith A. B. T." box="[468,821,1230,1257]" pageId="8" pageNumber="228" pagination="192" refId="ref28185" refString="Vaz-de-Mello F. Z., Genier F. & Smith A. B. T. 2010. Reclassification of Homocopris Burmeister as a valid genus to accomodate three species formerly in Dichotomius Hope (Coleoptera: Scarabaeinae: Coprini). The Coleopterists Bulletin 64 (3): 192. https: // doi. org / 10.1649 / 0010 - 065 X- 64.3.192.3" type="journal article" year="2010">
|
||
Vaz-de-Mello
|
||
<emphasis id="B901EAB3EF2DFFB6FDB3FB50CB8BE69F" box="[636,688,1230,1256]" italics="true" pageId="8" pageNumber="228">et al</emphasis>
|
||
. 2010: 192
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(new combination). —
|
||
<treatmentCitation id="0AD410B0EF2DFFB6FB80FB51CC4DE69F" author="Chamorro W. & Marin-Armijos D. & Granda V. & Vaz-de-Mello F. Z." box="[1103,1398,1230,1257]" page="96" pageId="8" pageNumber="228" year="2018">
|
||
<bibRefCitation id="EFE44B50EF2DFFB6FB80FB51CC4DE69F" author="Chamorro W. & Marin-Armijos D. & Granda V. & Vaz-de-Mello F. Z." box="[1103,1398,1230,1257]" pageId="8" pageNumber="228" pagination="72 - 100" refId="ref25178" refString="Chamorro W., Marin-Armijos D., Granda V. & Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / SOCOLEN. V 44 I 1.6545" type="journal article" year="2018">
|
||
Chamorro
|
||
<emphasis id="B901EAB3EF2DFFB6FB02FB50CC33E69F" box="[1229,1288,1230,1256]" italics="true" pageId="8" pageNumber="228">et al.</emphasis>
|
||
2018: 96
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(distribution); 2019: 159 (catalogue, faunistic).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72FA8ACAB1E758" blockId="8.[189,1398,1054,1327]" box="[189,906,1301,1327]" pageId="8" pageNumber="228">
|
||
<treatmentCitationGroup id="AB65118FEF2DFFB6FF72FA8ACAB1E758" box="[189,906,1301,1327]" pageId="8" pageNumber="228">
|
||
<taxonomicName id="4C754D22EF2DFFB6FF72FA8AC887E758" authorityName="Pereira" authorityYear="1953" box="[189,444,1301,1327]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72FA8AC887E758" box="[189,444,1301,1327]" italics="true" pageId="8" pageNumber="228">Dichotomius buckleyi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AD410B0EF2DFFB6FE16FA8ACA39E758" author="Saavedra A. D. & Vaz-de-Mello F. & Ugaz C. A. & Pacherre T. C." box="[473,770,1301,1327]" page="111" pageId="8" pageNumber="228" year="2017">
|
||
<bibRefCitation id="EFE44B50EF2DFFB6FE16FA8ACA39E758" author="Saavedra A. D. & Vaz-de-Mello F. & Ugaz C. A. & Pacherre T. C." box="[473,770,1301,1327]" pageId="8" pageNumber="228" pagination="108 - 116" refId="ref27601" refString="Saavedra A. D., Vaz-de-Mello F., Ugaz C. A. & Pacherre T. C. 2017. Coleopteros (Coleoptera: Scarabaeidae) de los bosques de niebla, Ramos y Chin Chin, Ayabaca-Huancabamba, Piura-Peru. INDES Revista de Investigacion para el Desarrollo Sustentable 3 (1): 108 - 116. https: // doi. org / 10.25127 / indes. 20153.138" type="journal article" year="2017">
|
||
Saavedra
|
||
<emphasis id="B901EAB3EF2DFFB6FD83FA89CBBCE758" box="[588,647,1301,1327]" italics="true" pageId="8" pageNumber="228">et al.</emphasis>
|
||
2017: 111
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(faunistic).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F652AEF2DFFB7FF72FAC9CA93E330" lastPageId="9" lastPageNumber="229" pageId="8" pageNumber="228" type="diagnosis">
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72FAC9C8F2E707" blockId="8.[189,457,1366,1392]" box="[189,457,1366,1392]" pageId="8" pageNumber="228">
|
||
<heading id="D08281CDEF2DFFB6FF72FAC9C8F2E707" bold="true" box="[189,457,1366,1392]" fontSize="11" level="3" pageId="8" pageNumber="228" reason="3">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72FAC9C8F2E707" bold="true" box="[189,457,1366,1392]" pageId="8" pageNumber="228">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72FA1AC8ECE438" blockId="8.[188,1401,1413,1615]" pageId="8" pageNumber="228">
|
||
Body length
|
||
<quantity id="4C8D9B44EF2DFFB6FE96FA1AC8E6E7E8" box="[345,477,1413,1440]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="8" pageNumber="228" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
|
||
. Dorsal ocular width greater than one quarter of interocular distance. Lateral pronotal carina and pronotal edge distinctly forking anteriorly then merging posteriorly, forming a closed carinated ellipse. Interstriae convex and impunctate. Meso-metasternal suture posteriorly arcuate. Visible abdominal ventrites laterally setose.
|
||
<specimenCount id="9D73FD28EF2DFFB6FD15FA70CBCBE47E" box="[730,752,1519,1545]" count="1" pageId="8" pageNumber="228" type="male">♂</specimenCount>
|
||
cephalic horn emerging centrally from clypeofrontal surface, parallel-sided with horizontal apex, curved throughout, flattened apically and length up to twice the interocular distance.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72F9E9C8D7E4E7" blockId="8.[189,492,1654,1680]" box="[189,492,1654,1680]" pageId="8" pageNumber="228">
|
||
<heading id="D08281CDEF2DFFB6FF72F9E9C8D7E4E7" bold="true" box="[189,492,1654,1680]" fontSize="11" level="3" pageId="8" pageNumber="228" reason="3">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72F9E9C8D7E4E7" bold="true" box="[189,492,1654,1680]" pageId="8" pageNumber="228">Name-bearing type data</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB6FF72F939CDEBE495" blockId="8.[189,1398,1701,1762]" pageId="8" pageNumber="228">
|
||
<taxonomicName id="4C754D22EF2DFFB6FF72F939CB27E4C8" authority="Waterhouse" authorityName="Waterhouse" authorityYear="1891" box="[189,540,1701,1727]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72F939C8B8E4C8" box="[189,387,1701,1727]" italics="true" pageId="8" pageNumber="228">Pinotus buckleyi</emphasis>
|
||
Waterhouse
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54CE8803EF2DFFB6FDE5F93ACBAAE4C8" box="[554,657,1701,1727]" pageId="8" pageNumber="228" type="holotype">holotype</typeStatus>
|
||
<specimenCount id="9D73FD28EF2DFFB6FD54F93ACB8AE4C8" box="[667,689,1701,1727]" count="1" pageId="8" pageNumber="228" type="male">♂</specimenCount>
|
||
(BMNH) (
|
||
<figureCitation id="134E2A24EF2DFFB6FCF5F93ACAAEE4B7" box="[826,917,1701,1728]" captionStart="Figs 46–57. 46–49" captionStartId="29.[189,242,1787,1813]" captionTargetBox="[278,1306,298,1712]" captionTargetId="figure-17@29.[258,1330,265,1743]" captionTargetPageId="29" captionText="Figs 46–57. 46–49. Elytral striae, posterolateral view. 46. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 47. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). Anterior pronotal surface. 48. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 49. Homocopris torulosus, neotype, ♂ (CMNC). 50–57. Type specimen labels. 50. Pinotus achamas Harold, 1867. 51. Pinotus buckleyi (Waterhouse, 1891). 52. Homocopris grossiorum. 53. Copris punctatissimus (Curtis, 1845). 54. Copris torulosus Eschscholtz, 1822. 55. Homocopris williami Darling & Génier sp. nov. 56. Pinotus simulator Luederwaldt, 1936. 57. Pinotus dahli Landin, 1955." figureDoi="http://doi.org/10.5281/zenodo.10824992" httpUri="https://zenodo.org/record/10824992/files/figure.png" pageId="8" pageNumber="228">Fig. 51</figureCitation>
|
||
): “Chiguin | -da | 80.14” [handwritten]; “Type” [white disc with red border]; “
|
||
<taxonomicName id="4C754D22EF2DFFB6FDB2F956CBECE495" authorityName="Erichson" authorityYear="1847" box="[637,727,1737,1762]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAB3EF2DFFB6FDB2F956CBECE495" box="[637,727,1737,1762]" italics="true" pageId="8" pageNumber="228">Pinotus</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C754D22EF2DFFB6FD24F957CA6BE495" authorityName="Waterhouse" authorityYear="1891" box="[747,848,1736,1762]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF2DFFB6FD24F957CA6BE495" box="[747,848,1736,1762]" italics="true" pageId="8" pageNumber="228">Buckleyi</emphasis>
|
||
</taxonomicName>
|
||
| (Type) Waterh.” [handwritten].
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2DFFB7FF72F895CA93E330" blockId="8.[189,1399,1801,1969]" lastBlockId="9.[189,1399,265,327]" lastPageId="9" lastPageNumber="229" pageId="8" pageNumber="228">
|
||
<taxonomicName id="4C754D22EF2DFFB6FF72F895CB09E554" authority="Luederwaldt" authorityName=", Luederwaldt" authorityYear="1936" box="[189,562,1801,1827]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="simulator">
|
||
<emphasis id="B901EAB3EF2DFFB6FF72F895C8B4E554" box="[189,399,1801,1827]" italics="true" pageId="8" pageNumber="228">Pinotus simulator</emphasis>
|
||
Luederwaldt
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54CE8803EF2DFFB6FDF1F896CB93E554" box="[574,680,1801,1827]" pageId="8" pageNumber="228" type="holotype">holotype</typeStatus>
|
||
, by monotypy,
|
||
<specimenCount id="9D73FD28EF2DFFB6FC90F895CA4EE553" box="[863,885,1802,1828]" count="1" pageId="8" pageNumber="228" type="male">♂</specimenCount>
|
||
(BMNH) (
|
||
<figureCitation id="134E2A24EF2DFFB6FC34F896CD6EE553" box="[1019,1109,1801,1828]" captionStart="Figs 46–57. 46–49" captionStartId="29.[189,242,1787,1813]" captionTargetBox="[278,1306,298,1712]" captionTargetId="figure-17@29.[258,1330,265,1743]" captionTargetPageId="29" captionText="Figs 46–57. 46–49. Elytral striae, posterolateral view. 46. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 47. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). Anterior pronotal surface. 48. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 49. Homocopris torulosus, neotype, ♂ (CMNC). 50–57. Type specimen labels. 50. Pinotus achamas Harold, 1867. 51. Pinotus buckleyi (Waterhouse, 1891). 52. Homocopris grossiorum. 53. Copris punctatissimus (Curtis, 1845). 54. Copris torulosus Eschscholtz, 1822. 55. Homocopris williami Darling & Génier sp. nov. 56. Pinotus simulator Luederwaldt, 1936. 57. Pinotus dahli Landin, 1955." figureDoi="http://doi.org/10.5281/zenodo.10824992" httpUri="https://zenodo.org/record/10824992/files/figure.png" pageId="8" pageNumber="228">Fig. 56</figureCitation>
|
||
): “Amazon”; “Degand”; “17140” [handwritten]; “Fry Coll. | 1905-100”; “
|
||
<emphasis id="B901EAB3EF2DFFB6FD3AF8B2CA0AE530" box="[757,817,1837,1863]" italics="true" pageId="8" pageNumber="228">Copr</emphasis>
|
||
. |
|
||
<taxonomicName id="4C754D22EF2DFFB6FC85F8B2CA81E530" authorityName=", Luederwaldt" authorityYear="1936" box="[842,954,1837,1863]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="simulator">
|
||
<emphasis id="B901EAB3EF2DFFB6FC85F8B2CA81E530" box="[842,954,1837,1863]" italics="true" pageId="8" pageNumber="228">simulator</emphasis>
|
||
</taxonomicName>
|
||
” [handwritten]; “
|
||
<taxonomicName id="4C754D22EF2DFFB6FB45F8B2CDDFE531" authorityName="Erichson" authorityYear="1847" box="[1162,1252,1837,1862]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAB3EF2DFFB6FB45F8B2CDDFE531" box="[1162,1252,1837,1862]" italics="true" pageId="8" pageNumber="228">Pinotus</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="9D73FD28EF2DFFB6FB26F8B2CC3BE530" box="[1257,1280,1837,1863]" count="1" pageId="8" pageNumber="228" type="female">♀</specimenCount>
|
||
|
|
||
<taxonomicName id="4C754D22EF2DFFB6FADFF8B2C9CDE51D" authority="Wat." authorityName="Waterhouse" authorityYear="1891" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF2DFFB6FADFF8B2CC4EE530" box="[1296,1397,1837,1863]" italics="true" pageId="8" pageNumber="228">Buckleyi</emphasis>
|
||
Wat.
|
||
</taxonomicName>
|
||
? | illegible | Lüderw. det. 1934” [Luederwaldt’s handwriting]; “
|
||
<typeStatus id="54CE8803EF2DFFB6FC17F8CFCD43E51D" box="[984,1144,1872,1898]" pageId="8" pageNumber="228" type="holotype">HOLOTYPE</typeStatus>
|
||
” [black border on red card]; “WORLD | SCARAB. | DATABASE | WSD00001803” [barcode label]; “
|
||
<taxonomicName id="4C754D22EF2DFFB6FBA1F8EBCDC4E5FA" authorityName="Burmeister" authorityYear="1846" box="[1134,1279,1908,1933]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAB3EF2DFFB6FBA1F8EBCDC4E5FA" box="[1134,1279,1908,1933]" italics="true" pageId="8" pageNumber="228">Homocopris</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C754D22EF2DFFB6FADAF8ECCC4CE5FA" authorityName="Waterhouse" authorityYear="1891" box="[1301,1399,1907,1933]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF2DFFB6FADAF8ECCC4CE5FA" box="[1301,1399,1907,1933]" italics="true" pageId="8" pageNumber="228">buckleyi</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="9D73FD28EF2DFFB6FF72F808C9E8E5C6" box="[189,211,1943,1969]" count="1" pageId="8" pageNumber="228" type="male">♂</specimenCount>
|
||
| dét. F. Génier, 2008” [partly handwritten]; “
|
||
<taxonomicName id="4C754D22EF2DFFB6FD3CF808CAA3E5C7" authority="Holot" authorityName="Holot" box="[755,920,1942,1968]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAB3EF2DFFB6FD3CF808CA76E5C7" box="[755,845,1943,1968]" italics="true" pageId="8" pageNumber="228">Pinotus</emphasis>
|
||
Holot
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C754D22EF2DFFB6FC7FF809CD57E5C7" authority="Lued." authorityName=", Luederwaldt" authorityYear="1936" box="[944,1132,1942,1968]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="228" phylum="Arthropoda" rank="species" species="simulator">
|
||
<emphasis id="B901EAB3EF2DFFB6FC7FF809CD1BE5C7" box="[944,1056,1942,1968]" italics="true" pageId="8" pageNumber="228">simulator</emphasis>
|
||
Lued.
|
||
</taxonomicName>
|
||
| des. F.
|
||
<bibRefCitation id="EFE44B50EF2DFFB6FB03F809CC4DE5C7" author="Vaz-de-Mello F. Z. & Edmonds W. D. & Ocampo F. C. & Schoolmeesters P." box="[1228,1398,1942,1968]" pageId="8" pageNumber="228" pagination="1 - 73" refId="ref28252" refString="Vaz-de-Mello F. Z., Edmonds W. D., Ocampo F. C. & Schoolmeesters P. 2011. A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa 2854 (1): 1 - 73. https: // doi. org / 10.11646 / zootaxa. 2854.1.1" type="journal article" year="2011">Vaz-de-Mello,</bibRefCitation>
|
||
2011” [partly handwritten]. The number ‘17140 corresponds in Fry’s notebooks in BMNH to ‘
|
||
<taxonomicName id="4C754D22EF2CFFB7FAD3FE96CC49E354" authorityName="Geoffroy" authorityYear="1762" box="[1308,1394,265,291]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="229" phylum="Arthropoda" rank="genus">Copris</taxonomicName>
|
||
,
|
||
<collectingCountry id="F3627631EF2CFFB7FF71FEB3C832E331" box="[190,265,300,326]" name="Brazil" pageId="9" pageNumber="229">Brazil</collectingCountry>
|
||
, Upper Amazon, Degand’ (M.V.L. Barclay pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F652AEF2CFFB7FF72FEF1CDF3E3C0" pageId="9" pageNumber="229" type="materials_examined">
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72FEF1C866E3FF" blockId="9.[189,349,366,392]" box="[189,349,366,392]" pageId="9" pageNumber="229">
|
||
<heading id="D08281CDEF2CFFB7FF72FEF1C866E3FF" bold="true" box="[189,349,366,392]" fontSize="11" level="3" pageId="9" pageNumber="229" reason="3">
|
||
<emphasis id="B901EAB3EF2CFFB7FF72FEF1C866E3FF" bold="true" box="[189,349,366,392]" pageId="9" pageNumber="229">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72FE02CDF3E3C0" blockId="9.[189,1224,412,439]" box="[189,1224,412,439]" pageId="9" pageNumber="229">
|
||
<taxonomicName id="4C754D22EF2CFFB7FF72FE02C8BBE3C1" authorityName="Waterhouse" authorityYear="1891" box="[189,384,412,438]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="229" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF2CFFB7FF72FE02C8BBE3C1" box="[189,384,412,438]" italics="true" pageId="9" pageNumber="229">Pinotus buckleyi</emphasis>
|
||
</taxonomicName>
|
||
: Chiguinda,
|
||
<collectingRegion id="49B1F843EF2CFFB7FDD7FE03CBD3E3C1" box="[536,744,412,438]" country="Ecuador" name="Morona-Santiago" pageId="9" pageNumber="229">Morona-Santiago</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3627631EF2CFFB7FD3BFE03CA61E3C1" box="[756,858,412,438]" name="Ecuador" pageId="9" pageNumber="229">Ecuador</collectingCountry>
|
||
;
|
||
<taxonomicName id="4C754D22EF2CFFB7FCAAFE02CD0DE3C1" authorityName=", Luederwaldt" authorityYear="1936" box="[869,1078,412,438]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="229" phylum="Arthropoda" rank="species" species="simulator">
|
||
<emphasis id="B901EAB3EF2CFFB7FCAAFE02CD0DE3C1" box="[869,1078,412,438]" italics="true" pageId="9" pageNumber="229">Pinotus simulator</emphasis>
|
||
</taxonomicName>
|
||
: Amazonas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F652AEF2CFFB7FF72FD4ACD52E694" pageId="9" pageNumber="229" type="materials_examined">
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72FD4ACB59E087" blockId="9.[189,610,725,752]" box="[189,610,725,752]" pageId="9" pageNumber="229">
|
||
<emphasis id="B901EAB3EF2CFFB7FF72FD4AC893E098" bold="true" box="[189,424,725,751]" pageId="9" pageNumber="229">Material examined</emphasis>
|
||
(
|
||
<specimenCount id="9D73FD28EF2CFFB7FE78FD49CB31E087" box="[439,522,726,752]" count="16" pageId="9" pageNumber="229" type="male">16 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D73FD28EF2CFFB7FDD9FD49CB6DE087" box="[534,598,726,752]" count="9" pageId="9" pageNumber="229" type="female">9 ♀♀</specimenCount>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72FC9BCC4CE1DB" blockId="9.[189,1399,772,940]" pageId="9" pageNumber="229">
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FF72FC9BCD22E168" box="[189,1049,772,799]" collectionCode="BMNH" collectorName="Buckley" country="Ecuador" location="ECUADOR" pageId="9" pageNumber="229" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<collectingCountry id="F3627631EF2CFFB7FF72FC9BC870E169" box="[189,331,772,798]" name="Ecuador" pageId="9" pageNumber="229">ECUADOR</collectingCountry>
|
||
•
|
||
<specimenCount id="9D73FD28EF2CFFB7FEADFC9BC8B4E168" box="[354,399,772,799]" count="1" pageId="9" pageNumber="229" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D73FD28EF2CFFB7FE54FC9BC8FCE168" box="[411,455,772,799]" count="1" pageId="9" pageNumber="229" type="female">1 ♀</specimenCount>
|
||
; [unspecified locality]; [no date];
|
||
<collectorName id="26805377EF2CFFB7FC9EFC9BCA8CE169" box="[849,951,772,798]" pageId="9" pageNumber="229">Buckley</collectorName>
|
||
;
|
||
<collectionCode id="ED64AE64EF2CFFB7FC0EFC9ACD22E168" box="[961,1049,773,799]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="9" pageNumber="229">BMNH</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FBEFFC9AC8F3E135" ID-GBIF-Occurrence="4602214301" collectionCode="MNHN" pageId="9" pageNumber="229" specimenCount="1" specimenCount-male="1">
|
||
•
|
||
<specimenCount id="9D73FD28EF2CFFB7FBFFFC9BCD65E168" box="[1072,1118,772,799]" count="1" pageId="9" pageNumber="229" type="male">1 ♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED64AE64EF2CFFB7FEA8FCB7C8FEE135" box="[359,453,808,834]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="229">MNHN</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FE01FCB7CBF6E112" ID-GBIF-Occurrence="4602214303" collectingDate="2005-12-10" collectionCode="CEMT" elevation="2920" latitude="-3.6666667" location="Bosque Natural Huashapamba" longLatPrecision="1307" longitude="-79.26667" pageId="9" pageNumber="229" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loja">
|
||
–
|
||
<emphasis id="B901EAB3EF2CFFB7FE2EFCB8CB27E136" bold="true" box="[481,540,807,833]" pageId="9" pageNumber="229">
|
||
<collectingRegion id="49B1F843EF2CFFB7FE2EFCB8CB27E136" box="[481,540,807,833]" country="Ecuador" name="Loja" pageId="9" pageNumber="229">Loja</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D73FD28EF2CFFB7FDFCFCB7CB5BE135" box="[563,608,808,834]" count="1" pageId="9" pageNumber="229" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D73FD28EF2CFFB7FDA4FCB7CBACE135" box="[619,663,808,834]" count="1" pageId="9" pageNumber="229" type="female">1 ♀</specimenCount>
|
||
;
|
||
<location id="8EAA607AEF2CFFB7FD6BFCB7CD32E136" LSID="urn:lsid:plazi:treatment:03DC87B7EF2DFFB7FE63FCCACDF3E3C0:8EAA607AEF2CFFB7FD6BFCB7CD32E136" box="[676,1033,807,834]" latitude="-3.6666667" longLatPrecision="1307" longitude="-79.26667" name="Bosque Natural Huashapamba" pageId="9" pageNumber="229" stateProvince="Loja">Bosque Natural Huashapamba</location>
|
||
,
|
||
<location id="8EAA607AEF2CFFB7FBDCFCB7CDBAE135" LSID="urn:lsid:plazi:treatment:03DC87B7EF2DFFB7FE63FCCACDF3E3C0:8EAA607AEF2CFFB7FBDCFCB7CDBAE135" box="[1043,1153,808,834]" latitude="-3.6666667" longLatPrecision="1307" longitude="-79.26667" name="Saraguro" pageId="9" pageNumber="229" stateProvince="Loja">Saraguro</location>
|
||
; [
|
||
<geoCoordinate id="EE415066EF2CFFB7FB5DFCB9CDD0E135" box="[1170,1259,806,834]" degrees="3" direction="south" minutes="40" orientation="latitude" pageId="9" pageNumber="229" precision="925" value="-3.6666667">3°40′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE415066EF2CFFB7FB38FCB9CC51E135" box="[1271,1386,806,834]" degrees="79" direction="west" minutes="16" orientation="longitude" pageId="9" pageNumber="229" precision="925" value="-79.26667">79°16′ W</geoCoordinate>
|
||
];
|
||
<quantity id="4C8D9B44EF2CFFB7FF72FCD4C821E112" box="[189,282,843,869]" metricMagnitude="3" metricUnit="m" metricValue="2.92" pageId="9" pageNumber="229" unit="m" value="2920.0">
|
||
<elevation id="0058D192EF2CFFB7FF72FCD4C821E112" box="[189,282,843,869]" metricMagnitude="3" metricUnit="m" metricValue="2.92" pageId="9" pageNumber="229" unit="m" value="2920.0">2920 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFCB1061EF2CFFB7FEE7FCD4C8F8E112" box="[296,451,842,869]" pageId="9" pageNumber="229" value="2005-12-10">
|
||
<collectingDate id="EF8FE989EF2CFFB7FEE7FCD4C8F8E112" box="[296,451,842,869]" pageId="9" pageNumber="229" value="2005-12-10">10 Dec.2005</collectingDate>
|
||
</date>
|
||
; [anonymous];
|
||
<collectionCode id="ED64AE64EF2CFFB7FDB3FCD4CBF6E112" box="[636,717,843,869]" collectionName="CEMT" pageId="9" pageNumber="229">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FD19FCD4C84CE1FF" ID-GBIF-Occurrence="4602214304" collectionCode="CMNC" collectorName="Abbe Gaujon" latitude="-3.9916668" location="Loja" longLatPrecision="21" longitude="-79.19334" pageId="9" pageNumber="229" specimenCount="1" specimenCount-male="1">
|
||
•
|
||
<specimenCount id="9D73FD28EF2CFFB7FD26FCD4CA21E112" box="[745,794,843,869]" count="1" pageId="9" pageNumber="229" type="male">1 ♂</specimenCount>
|
||
;
|
||
<location id="8EAA607AEF2CFFB7FCE7FCD4CA59E112" LSID="urn:lsid:plazi:treatment:03DC87B7EF2DFFB7FE63FCCACDF3E3C0:8EAA607AEF2CFFB7FCE7FCD4CA59E112" box="[808,866,843,869]" latitude="-3.9916668" longLatPrecision="21" longitude="-79.19334" name="Loja" pageId="9" pageNumber="229">Loja</location>
|
||
; [
|
||
<geoCoordinate id="EE415066EF2CFFB7FCBAFCD6CAC0E112" box="[885,1019,841,869]" degrees="3" direction="south" minutes="59" orientation="latitude" pageId="9" pageNumber="229" precision="15" seconds="30" value="-3.9916668">3°59′30″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE415066EF2CFFB7FBC6FCD6CD93E112" box="[1033,1192,841,869]" degrees="79" direction="west" minutes="11" orientation="longitude" pageId="9" pageNumber="229" precision="15" seconds="36" value="-79.19334">79°11′36″ W</geoCoordinate>
|
||
]; [no date]:
|
||
<collectorName id="26805377EF2CFFB7FAF8FCD4C82CE1FF" pageId="9" pageNumber="229">Abbé Gaujon</collectorName>
|
||
;
|
||
<collectionCode id="ED64AE64EF2CFFB7FEEEFCF1C84CE1FF" box="[289,375,878,904]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="9" pageNumber="229">CMNC</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FEB1FCF1CB1BE1DB" ID-GBIF-Occurrence="4602214305" collectionCode="MNHN" pageId="9" pageNumber="229" specimenCount="8" specimenCount-female="2" specimenCount-male="6">
|
||
•
|
||
<specimenCount id="9D73FD28EF2CFFB7FE41FCF1C8EBE1FF" box="[398,464,878,904]" count="6" pageId="9" pageNumber="229" type="male">6 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D73FD28EF2CFFB7FE13FCF1CB27E1FF" box="[476,540,878,904]" count="2" pageId="9" pageNumber="229" type="female">2 ♀♀</specimenCount>
|
||
(one of these was used for the descriptions of both sexes); same collection data as for preceding;
|
||
<collectionCode id="ED64AE64EF2CFFB7FE70FC0DCB26E1DB" box="[447,541,914,940]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="229">MNHN</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FDE9FC0DCC4CE1DB" ID-GBIF-Occurrence="4602214302" box="[550,1399,912,940]" collectionCode="MNHN" collectorName="Buckley" latitude="-1.7333333" location="Sarayacu" longLatPrecision="1308" longitude="-77.48333" pageId="9" pageNumber="229" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza">
|
||
–
|
||
<emphasis id="B901EAB3EF2CFFB7FDF4FC0ECBA0E1DC" bold="true" box="[571,667,913,939]" pageId="9" pageNumber="229">
|
||
<collectingRegion id="49B1F843EF2CFFB7FDF4FC0ECBA0E1DC" box="[571,667,913,939]" country="Ecuador" name="Pastaza" pageId="9" pageNumber="229">Pastaza</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D73FD28EF2CFFB7FD7CFC0ECBD9E1DB" box="[691,738,913,940]" count="1" pageId="9" pageNumber="229" type="male">1 ♂</specimenCount>
|
||
;
|
||
<location id="8EAA607AEF2CFFB7FD22FC0ECA67E1DC" LSID="urn:lsid:plazi:treatment:03DC87B7EF2DFFB7FE63FCCACDF3E3C0:8EAA607AEF2CFFB7FD22FC0ECA67E1DC" box="[749,860,913,939]" latitude="-1.7333333" longLatPrecision="1308" longitude="-77.48333" name="Sarayacu" pageId="9" pageNumber="229" stateProvince="Pastaza">Sarayacu</location>
|
||
; [
|
||
<geoCoordinate id="EE415066EF2CFFB7FCA0FC0FCAF3E1DC" box="[879,968,912,940]" degrees="1" direction="south" minutes="44" orientation="latitude" pageId="9" pageNumber="229" precision="925" value="-1.7333333">1°44′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE415066EF2CFFB7FC1BFC0FCD73E1DB" box="[980,1096,912,940]" degrees="77" direction="west" minutes="29" orientation="longitude" pageId="9" pageNumber="229" precision="925" value="-77.48333">77°29′ W</geoCoordinate>
|
||
]; 1879;
|
||
<collectorName id="26805377EF2CFFB7FB6AFC0ECC30E1DC" box="[1189,1291,913,939]" pageId="9" pageNumber="229">Buckley</collectorName>
|
||
;
|
||
<collectionCode id="ED64AE64EF2CFFB7FADAFC0DCC48E1DB" box="[1301,1395,914,940]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="229">MNHN</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72FC4CCD52E694" blockId="9.[189,1399,977,1251]" pageId="9" pageNumber="229">
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FF72FC4CCA60E667" ID-GBIF-Occurrence="4601643476" collectedFrom="cloud forest" collectingDate="1979-10-16" collectionCode="BDGC" collectorName="J. L. Barkley" country="Peru" county="Cumpang" elevation="2625" latitude="-8.333333" location="above [Rio] Uctubamba on trail to Ongon" longLatPrecision="1301" longitude="-76.98333" municipality="Uctubamba" pageId="9" pageNumber="229" specimenCount="1" specimenCount-female="1" stateProvince="La Libertad">
|
||
<collectingCountry id="F3627631EF2CFFB7FF72FC4CC833E19A" box="[189,264,979,1005]" name="Peru" pageId="9" pageNumber="229">PERU</collectingCountry>
|
||
–
|
||
<emphasis id="B901EAB3EF2CFFB7FEEBFC4CC886E19B" bold="true" box="[292,445,978,1005]" pageId="9" pageNumber="229">
|
||
<collectingRegion id="49B1F843EF2CFFB7FEEBFC4CC886E19B" box="[292,445,978,1005]" country="Peru" name="La Libertad" pageId="9" pageNumber="229">La Libertad</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D73FD28EF2CFFB7FE1AFC4CCB39E19A" box="[469,514,979,1005]" count="1" pageId="9" pageNumber="229" type="female">1 ♀</specimenCount>
|
||
;
|
||
<collectingCounty id="62AB4E2DEF2CFFB7FDDFFC4CCBBEE19A" box="[528,645,979,1005]" pageId="9" pageNumber="229">Cumpang</collectingCounty>
|
||
,
|
||
<location id="8EAA607AEF2CFFB7FD5FFC4DCD46E19A" LSID="urn:lsid:plazi:treatment:03DC87B7EF2DFFB7FE63FCCACDF3E3C0:8EAA607AEF2CFFB7FD5FFC4DCD46E19A" box="[656,1149,978,1005]" country="Peru" county="Cumpang" latitude="-8.333333" longLatPrecision="1301" longitude="-76.98333" municipality="Uctubamba" name="above [Rio] Uctubamba on trail to Ongon" pageId="9" pageNumber="229" stateProvince="La Libertad">above [Río] Uctubamba on trail to Ongón</location>
|
||
; [
|
||
<geoCoordinate id="EE415066EF2CFFB7FB40FC4ECDD2E19A" box="[1167,1257,977,1005]" degrees="8" direction="south" minutes="20" orientation="latitude" pageId="9" pageNumber="229" precision="925" value="-8.333333">8°20′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE415066EF2CFFB7FB39FC4ECC51E19A" box="[1270,1386,977,1005]" degrees="76" direction="west" minutes="59" orientation="longitude" pageId="9" pageNumber="229" precision="925" value="-76.98333">76°59′ W</geoCoordinate>
|
||
];
|
||
<quantity id="4C8D9B44EF2CFFB7FF72FC69C822E667" box="[189,281,1014,1041]" metricMagnitude="3" metricUnit="m" metricValue="2.625" pageId="9" pageNumber="229" unit="m" value="2625.0">
|
||
<elevation id="0058D192EF2CFFB7FF72FC69C822E667" box="[189,281,1014,1041]" metricMagnitude="3" metricUnit="m" metricValue="2.625" pageId="9" pageNumber="229" unit="m" value="2625.0">2625 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFCB1061EF2CFFB7FEEAFC69C8FBE667" box="[293,448,1014,1040]" pageId="9" pageNumber="229" value="1979-10-16">
|
||
<collectingDate id="EF8FE989EF2CFFB7FEEAFC69C8FBE667" box="[293,448,1014,1040]" pageId="9" pageNumber="229" value="1979-10-16">16 Oct. 1979</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26805377EF2CFFB7FE03FC69CB5AE667" box="[460,609,1014,1040]" pageId="9" pageNumber="229">J.L. Barkley</collectorName>
|
||
;
|
||
<collectedFrom id="3869755CEF2CFFB7FDA3FC69CBC0E667" box="[620,763,1014,1040]" pageId="9" pageNumber="229">cloud forest</collectedFrom>
|
||
;
|
||
<collectionCode id="ED64AE64EF2CFFB7FCC9FC69CA60E667" box="[774,859,1014,1040]" pageId="9" pageNumber="229">BDGC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FCA9FC69CBE2E644" ID-GBIF-Occurrence="4602214306" collectingDate="2016-01" collectionCode="CMD" collectorName="Roja & Gonzales" country="Peru" elevation="3000" latitude="-4.633333" location="Ayabaca" longLatPrecision="1306" longitude="-79.71667" pageId="9" pageNumber="229" specimenCount="1" specimenCount-male="1" stateProvince="Piura">
|
||
–
|
||
<emphasis id="B901EAB3EF2CFFB7FCB3FC6ACAF9E678" bold="true" box="[892,962,1013,1039]" pageId="9" pageNumber="229">
|
||
<collectingRegion id="49B1F843EF2CFFB7FCB3FC6ACAF9E678" box="[892,962,1013,1039]" country="Peru" name="Piura" pageId="9" pageNumber="229">Piura</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D73FD28EF2CFFB7FC14FC69CD31E667" box="[987,1034,1014,1040]" count="1" pageId="9" pageNumber="229" type="male">1 ♂</specimenCount>
|
||
;
|
||
<location id="8EAA607AEF2CFFB7FBDBFC69CD47E667" LSID="urn:lsid:plazi:treatment:03DC87B7EF2DFFB7FE63FCCACDF3E3C0:8EAA607AEF2CFFB7FBDBFC69CD47E667" box="[1044,1148,1014,1040]" country="Peru" latitude="-4.633333" longLatPrecision="1306" longitude="-79.71667" name="Ayabaca" pageId="9" pageNumber="229" stateProvince="Piura">Ayabaca</location>
|
||
; [
|
||
<geoCoordinate id="EE415066EF2CFFB7FB40FC6BCDD2E667" box="[1167,1257,1012,1040]" degrees="4" direction="south" minutes="38" orientation="latitude" pageId="9" pageNumber="229" precision="925" value="-4.633333">4°38′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE415066EF2CFFB7FB3AFC6BCC51E667" box="[1269,1386,1012,1040]" degrees="79" direction="west" minutes="43" orientation="longitude" pageId="9" pageNumber="229" precision="925" value="-79.71667">79°43′ W</geoCoordinate>
|
||
];
|
||
<quantity id="4C8D9B44EF2CFFB7FF72FB86C821E644" box="[189,282,1049,1075]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="229" unit="m" value="3000.0">
|
||
<elevation id="0058D192EF2CFFB7FF72FB86C821E644" box="[189,282,1049,1075]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="229" unit="m" value="3000.0">3000 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFCB1061EF2CFFB7FEE9FB86C8A0E644" box="[294,411,1049,1075]" pageId="9" pageNumber="229" value="2016-01">
|
||
<collectingDate id="EF8FE989EF2CFFB7FEE9FB86C8A0E644" box="[294,411,1049,1075]" pageId="9" pageNumber="229" value="2016-01">Jan. 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26805377EF2CFFB7FE67FB86C8E4E644" box="[424,479,1049,1075]" pageId="9" pageNumber="229">Roja</collectorName>
|
||
and
|
||
<collectorName id="26805377EF2CFFB7FDD6FB86CBB1E644" box="[537,650,1049,1075]" pageId="9" pageNumber="229">Gonzales</collectorName>
|
||
;
|
||
<collectionCode id="ED64AE64EF2CFFB7FD59FB86CBE2E644" box="[662,729,1049,1075]" collectionName="CMD" pageId="9" pageNumber="229">CMD</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FD3CFB86CB33E60E" ID-GBIF-Occurrence="4601643438" collectedFrom="cloud forest, bovine dung" collectingDate="2009-04-16" collectionCode="CEMT" collectorName="D. Saavedra" country="Peru" county="Provincia de Ayacaba" elevation="3055" latitude="-4.9997225" location="San Juan" longLatPrecision="21" longitude="-79.65806" municipality="Bosque de Chin Chin" pageId="9" pageNumber="229" specimenCount="1" specimenCount-male="1" stateProvince="Piura">
|
||
<specimenCount id="9D73FD28EF2CFFB7FD3CFB86CA18E644" box="[755,803,1049,1075]" count="1" pageId="9" pageNumber="229" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingCounty id="62AB4E2DEF2CFFB7FCFFFB86CD0AE644" box="[816,1073,1049,1075]" pageId="9" pageNumber="229">Provincia de Ayacaba</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BAEACDBEF2CFFB7FBF1FB86CC05E644" box="[1086,1342,1049,1075]" pageId="9" pageNumber="229">Bosque de Chin Chin</collectingMunicipality>
|
||
,
|
||
<location id="8EAA607AEF2CFFB7FA85FB86C9CDE620" LSID="urn:lsid:plazi:treatment:03DC87B7EF2DFFB7FE63FCCACDF3E3C0:8EAA607AEF2CFFB7FA85FB86C9CDE620" country="Peru" county="Provincia de Ayacaba" latitude="-4.9997225" longLatPrecision="21" longitude="-79.65806" municipality="Bosque de Chin Chin" name="San Juan" pageId="9" pageNumber="229" stateProvince="Piura">San Juan</location>
|
||
,
|
||
<location id="8EAA607AEF2CFFB7FECCFBA3CB26E621" LSID="urn:lsid:plazi:treatment:03DC87B7EF2DFFB7FE63FCCACDF3E3C0:8EAA607AEF2CFFB7FECCFBA3CB26E621" box="[259,541,1084,1110]" country="Peru" county="Provincia de Ayacaba" latitude="-4.9997225" longLatPrecision="21" longitude="-79.65806" municipality="Bosque de Chin Chin" name="Distrito de Pacaipampa" pageId="9" pageNumber="229" stateProvince="Piura">Distrito de Pacaipampa</location>
|
||
; [
|
||
<geoCoordinate id="EE415066EF2CFFB7FDFFFBA4CB8DE621" box="[560,694,1083,1111]" degrees="4" direction="south" minutes="59" orientation="latitude" pageId="9" pageNumber="229" precision="15" seconds="59" value="-4.9997225">4°59′59″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE415066EF2CFFB7FD0AFBA4CA5EE620" box="[709,869,1083,1111]" degrees="79" direction="west" minutes="39" orientation="longitude" pageId="9" pageNumber="229" precision="15" seconds="29" value="-79.65806">79°39′29″ W</geoCoordinate>
|
||
];
|
||
<quantity id="4C8D9B44EF2CFFB7FCB5FBA3CAE3E620" box="[890,984,1084,1111]" metricMagnitude="3" metricUnit="m" metricValue="3.055" pageId="9" pageNumber="229" unit="m" value="3055.0">
|
||
<elevation id="0058D192EF2CFFB7FCB5FBA3CAE3E620" box="[890,984,1084,1111]" metricMagnitude="3" metricUnit="m" metricValue="3.055" pageId="9" pageNumber="229" unit="m" value="3055.0">3055 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFCB1061EF2CFFB7FC29FBA3CDBEE621" box="[998,1157,1084,1110]" pageId="9" pageNumber="229" value="2009-04-16">
|
||
<collectingDate id="EF8FE989EF2CFFB7FC29FBA3CDBEE621" box="[998,1157,1084,1110]" pageId="9" pageNumber="229" value="2009-04-16">16 Apr. 2009</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26805377EF2CFFB7FB5CFBA2CC13E621" box="[1171,1320,1084,1111]" pageId="9" pageNumber="229">D. Saavedra</collectorName>
|
||
;
|
||
<collectedFrom id="3869755CEF2CFFB7FAFAFBA3C838E60E" pageId="9" pageNumber="229">cloud forest</collectedFrom>
|
||
,
|
||
<collectedFrom id="3869755CEF2CFFB7FEDEFBC0C892E60E" box="[273,425,1119,1145]" pageId="9" pageNumber="229">bovine dung</collectedFrom>
|
||
;
|
||
<collectionCode id="ED64AE64EF2CFFB7FE78FBC0CB33E60E" box="[439,520,1119,1145]" collectionName="CEMT" pageId="9" pageNumber="229">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FDE9FBC0CDF7E60E" ID-GBIF-Occurrence="4601643536" box="[550,1228,1119,1146]" collectingDate="2009-04-16" collectionCode="PMOC" collectorName="D. Saavedra" country="Peru" county="Provincia de Ayacaba" elevation="3055" latitude="-4.9997225" location="San Juan" longLatPrecision="21" longitude="-79.65806" municipality="Bosque de Chin Chin" pageId="9" pageNumber="229" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Piura">
|
||
<specimenCount id="9D73FD28EF2CFFB7FDE9FBC0CB6CE60D" box="[550,599,1119,1146]" count="1" pageId="9" pageNumber="229" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D73FD28EF2CFFB7FDA9FBC0CBADE60D" box="[614,662,1119,1146]" count="1" pageId="9" pageNumber="229" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED64AE64EF2CFFB7FBB6FBC0CDF7E60E" box="[1145,1228,1119,1145]" collectionName="PMOC" pageId="9" pageNumber="229">PMOC</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FB25FBC0CD1DE6B7" ID-GBIF-Occurrence="4601643347" collectedFrom="cloud forest" collectingDate="2009-02-16" collectingMethod="pitfall trap" collectionCode="CEMT" collectorName="D. Saavedra" country="Peru" elevation="2834" latitude="-4.701111" location="Comunidad Campesina de Samanga del Sector Espindola" longLatPrecision="21" longitude="-79.461945" municipality="Bosque de Ramos" pageId="9" pageNumber="229" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Piura">
|
||
<specimenCount id="9D73FD28EF2CFFB7FB25FBC0CC20E60D" box="[1258,1307,1119,1146]" count="1" pageId="9" pageNumber="229" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D73FD28EF2CFFB7FAE5FBC0CC54E60D" box="[1322,1391,1119,1146]" count="2" pageId="9" pageNumber="229" type="female">2 ♀♀</specimenCount>
|
||
;
|
||
<collectingMunicipality id="6BAEACDBEF2CFFB7FF72FB1CC8AAE6EA" box="[189,401,1154,1181]" pageId="9" pageNumber="229">Bosque de Ramos</collectingMunicipality>
|
||
,
|
||
<location id="8EAA607AEF2CFFB7FE53FB1DCD00E6EB" LSID="urn:lsid:plazi:treatment:03DC87B7EF2DFFB7FE63FCCACDF3E3C0:8EAA607AEF2CFFB7FE53FB1DCD00E6EB" box="[412,1083,1154,1181]" country="Peru" latitude="-4.701111" longLatPrecision="21" longitude="-79.461945" municipality="Bosque de Ramos" name="Comunidad Campesina de Samanga del Sector Espindola" pageId="9" pageNumber="229" stateProvince="Piura">Comunidad Campesina de Samanga del Sector Espíndola</location>
|
||
; [
|
||
<geoCoordinate id="EE415066EF2CFFB7FB82FB1ECDFAE6EA" box="[1101,1217,1153,1181]" degrees="4" direction="south" minutes="42" orientation="latitude" pageId="9" pageNumber="229" precision="15" seconds="4" value="-4.701111">4°42′4″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE415066EF2CFFB7FB02FB1ECC51E6EA" box="[1229,1386,1153,1181]" degrees="79" direction="west" minutes="27" orientation="longitude" pageId="9" pageNumber="229" precision="15" seconds="43" value="-79.461945">79°27′43″ W</geoCoordinate>
|
||
];
|
||
<quantity id="4C8D9B44EF2CFFB7FF72FB39C821E6B7" box="[189,282,1190,1216]" metricMagnitude="3" metricUnit="m" metricValue="2.8339999999999996" pageId="9" pageNumber="229" unit="m" value="2834.0">
|
||
<elevation id="0058D192EF2CFFB7FF72FB39C821E6B7" box="[189,282,1190,1216]" metricMagnitude="3" metricUnit="m" metricValue="2.8339999999999996" pageId="9" pageNumber="229" unit="m" value="2834.0">2834 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFCB1061EF2CFFB7FEE8FB39C8FDE6B7" box="[295,454,1190,1216]" pageId="9" pageNumber="229" value="2009-02-16">
|
||
<collectingDate id="EF8FE989EF2CFFB7FEE8FB39C8FDE6B7" box="[295,454,1190,1216]" pageId="9" pageNumber="229" value="2009-02-16">16 Feb. 2009</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26805377EF2CFFB7FE1CFB39CB5DE6B7" box="[467,614,1190,1216]" pageId="9" pageNumber="229">D. Saavedra</collectorName>
|
||
;
|
||
<collectedFrom id="3869755CEF2CFFB7FDBCFB39CA39E6B7" box="[627,770,1190,1216]" pageId="9" pageNumber="229">cloud forest</collectedFrom>
|
||
,
|
||
<collectingMethod id="52344EB6EF2CFFB7FCC1FB39CAB4E6B7" box="[782,911,1190,1216]" pageId="9" pageNumber="229">pitfall trap</collectingMethod>
|
||
: fish;
|
||
<collectionCode id="ED64AE64EF2CFFB7FC1AFB39CD1DE6B7" box="[981,1062,1190,1216]" collectionName="CEMT" pageId="9" pageNumber="229">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B1D3CFCEF2CFFB7FB8FFB39CD59E694" ID-GBIF-Occurrence="4601643358" collectedFrom="cloud forest" collectingDate="2009-05-27" collectingMethod="pitfall trap" collectionCode="CEMT" collectorName="D. Saavedra" country="Peru" elevation="2834" latitude="-4.701111" location="Comunidad Campesina de Samanga del Sector Espindola" longLatPrecision="21" longitude="-79.461945" municipality="Bosque de Ramos" pageId="9" pageNumber="229" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Piura">
|
||
<specimenCount id="9D73FD28EF2CFFB7FB8FFB39CD54E6B7" box="[1088,1135,1190,1216]" count="1" pageId="9" pageNumber="229" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D73FD28EF2CFFB7FBB2FB39CD91E6B7" box="[1149,1194,1190,1216]" count="1" pageId="9" pageNumber="229" type="female">1 ♀</specimenCount>
|
||
; same locality as for preceding;
|
||
<date id="FFCB1061EF2CFFB7FEA5FB56CB32E694" box="[362,521,1225,1251]" pageId="9" pageNumber="229" value="2009-05-27">
|
||
<collectingDate id="EF8FE989EF2CFFB7FEA5FB56CB32E694" box="[362,521,1225,1251]" pageId="9" pageNumber="229" value="2009-05-27">27 May 2009</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26805377EF2CFFB7FDD9FB56CB93E694" box="[534,680,1225,1251]" pageId="9" pageNumber="229">D. Saavedra</collectorName>
|
||
;
|
||
<collectedFrom id="3869755CEF2CFFB7FD7BFB56CA78E694" box="[692,835,1225,1251]" pageId="9" pageNumber="229">cloud forest</collectedFrom>
|
||
,
|
||
<collectingMethod id="52344EB6EF2CFFB7FC81FB56CAF5E694" box="[846,974,1225,1251]" pageId="9" pageNumber="229">pitfall trap</collectingMethod>
|
||
: fish;
|
||
<collectionCode id="ED64AE64EF2CFFB7FBDCFB56CD59E694" box="[1043,1122,1225,1251]" collectionName="CEMT" pageId="9" pageNumber="229">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F652AEF2CFFB2FF72FA95C83BE4AA" lastPageId="12" lastPageNumber="232" pageId="9" pageNumber="229" type="description">
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72FA95C875E753" blockId="9.[189,334,1290,1316]" box="[189,334,1290,1316]" pageId="9" pageNumber="229">
|
||
<heading id="D08281CDEF2CFFB7FF72FA95C875E753" bold="true" box="[189,334,1290,1316]" fontSize="11" level="3" pageId="9" pageNumber="229" reason="3">
|
||
<emphasis id="B901EAB3EF2CFFB7FF72FA95C875E753" bold="true" box="[189,334,1290,1316]" pageId="9" pageNumber="229">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72FAA6CDB5E724" blockId="9.[189,1166,1337,1363]" box="[189,1166,1337,1363]" pageId="9" pageNumber="229">
|
||
The
|
||
<typeStatus id="54CE8803EF2CFFB7FF3DFAA6C862E724" box="[242,345,1337,1363]" pageId="9" pageNumber="229" type="holotype">holotype</typeStatus>
|
||
was previously studied, but is not available for a detailed description.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2CFFB7FF25FAE5CBDFE7E3" blockId="9.[189,740,1402,1464]" box="[234,740,1402,1429]" pageId="9" pageNumber="229">
|
||
<emphasis id="B901EAB3EF2CFFB7FF25FAE5C89EE7E3" bold="true" box="[234,421,1402,1428]" pageId="9" pageNumber="229">Male specimen</emphasis>
|
||
(
|
||
<figureCitation id="134E2A24EF2CFFB7FE7BFAE5CB3BE7E3" box="[436,512,1402,1428]" captionStart="Figs 1–4" captionStartId="6.[189,242,1892,1918]" captionTargetBox="[234,1360,273,1801]" captionTargetId="figure-12@6.[218,1370,265,1852]" captionTargetPageId="6" captionText="Figs 1–4. Dorsal habitus. 1. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 2. Andinocopris achamas, alloreferent paralectotype, ♀ (MNHN). 3. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 4. Andinocopris buckleyi, specimen ♀ (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.10824970" httpUri="https://zenodo.org/record/10824970/files/figure.png" pageId="9" pageNumber="229">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="134E2A24EF2CFFB7FDC2FAE5CB17E7E2" box="[525,556,1402,1429]" captionStart="Figs13–16" captionStartId="14.[189,242,1891,1917]" captionTargetBox="[221,1346,305,1841]" captionTargetId="figure-12@14.[221,1366,263,1841]" captionTargetPageId="14" captionText="Figs13–16.Headandpronotum,lateralview.13. Andinocoprisachamas(Harold,1867)gen.etcomb.nov., lectotype, ♂ (MNHN). 14. Andinocopris achamas, paralectotype, ♀ (MNHN). 15. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 16. Andinocopris buckleyi, specimen ♀ (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.10824978" httpUri="https://zenodo.org/record/10824978/files/figure.png" pageId="9" pageNumber="229">15</figureCitation>
|
||
,
|
||
<figureCitation id="134E2A24EF2CFFB7FDF6FAE5CB62E7E3" box="[569,601,1402,1428]" captionStart="Figs25–30" captionStartId="19.[189,242,1857,1883]" captionTargetBox="[239,1354,353,1767]" captionTargetId="figure-17@19.[234,1354,265,1809]" captionTargetPageId="19" captionText="Figs25–30. Paramere,lateralview.25. Andinocoprisachamas(Harold,1867)gen.etcomb.nov.,specimen ♂ (CMNC). 26. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 27. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 28. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 29. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 30. Homocopris williami Darling & Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824984" httpUri="https://zenodo.org/record/10824984/files/figure.png" pageId="9" pageNumber="229">26</figureCitation>
|
||
,
|
||
<figureCitation id="134E2A24EF2CFFB7FDAAFAE5CBBEE7E3" box="[613,645,1402,1428]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling & Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="9" pageNumber="229">32</figureCitation>
|
||
,
|
||
<figureCitation id="134E2A24EF2CFFB7FD5EFAE5CB8AE7E3" box="[657,689,1402,1428]" captionStart="Figs 37–41. 37–38" captionStartId="23.[189,242,1892,1918]" captionTargetBox="[257,1314,294,1816]" captionTargetId="figure-17@23.[221,1366,265,1843]" captionTargetPageId="23" captionText="Figs 37–41. 37–38. Genital segment, dorsal view. 37. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 38. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 39–41. Hindwing, dorsal view. 39. Andinocopris achamas (Harold, 1867) gen. et comb. nov. 40. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov. 41. Homocopris torulosus." figureDoi="http://doi.org/10.5281/zenodo.10824988" httpUri="https://zenodo.org/record/10824988/files/figure.png" pageId="9" pageNumber="229">40</figureCitation>
|
||
,
|
||
<figureCitation id="134E2A24EF2CFFB7FD72FAE5CBE7E7E3" box="[701,732,1402,1428]" captionStart="Figs 42–45" captionStartId="25.[189,242,1822,1848]" captionTargetBox="[232,1356,285,1784]" captionTargetId="figure-18@25.[232,1356,265,1787]" captionTargetPageId="25" captionText="Figs 42–45. Metasternum, ventral view. In specimens with a metasternal depression, the lateral depression is bordered with white dotted lines. For Homocopris grossiorum Darling & Génier sp. nov., a red dotted line indicates where the metasternal depression is interrupted. 42. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♀ (MNHN) 43. Homocopris grossiorum, holotype ♂ (MZSP). 44. Homocopris punctatissimus (Curtis, 1845), specimen ♀ (NMPC). 45. Homocopris torulosus (Eschscholtz, 1822), specimen ♂ (CMNC)." figureDoi="http://doi.org/10.5281/zenodo.10824990" httpUri="https://zenodo.org/record/10824990/files/figure.png" pageId="9" pageNumber="229">42</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72FA01CBBEE7CF" blockId="9.[189,740,1402,1464]" box="[189,645,1437,1464]" pageId="9" pageNumber="229">
|
||
<smallCapsWord id="8D2CA07DEF2CFFB7FF72FA01C855E7C1" baselines="1457,1457" box="[189,366,1438,1464]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="9" pageNumber="229">MEASUREMENTS</smallCapsWord>
|
||
. Body length
|
||
<quantity id="4C8D9B44EF2CFFB7FDDAFA02CBBBE7CF" box="[533,640,1437,1464]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="9" pageNumber="229" unit="mm" value="16.4">16.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72FA7FCBCFE4F1" blockId="9.[188,1400,1503,1671]" pageId="9" pageNumber="229">
|
||
<smallCapsWord id="8D2CA07DEF2CFFB7FF72FA7FC9C7E78F" baselines="1523,1523" box="[189,252,1504,1530]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="9" pageNumber="229">HEAD</smallCapsWord>
|
||
. Clypeal edge broadly arcuate, lacking teeth. Anteroventral clypeal tooth acute. Clypeal and genal surface rugose, clypeogenal suture ill-defined. Posterior genal angle obtuse. Frontal surface reduced, smooth. Dorsal ocular width greater than one-quarter interocular distance. Interocular surface smooth. Cephalic horn emerging centrally from clypeofrontal surface, parallel-sided with quadrate apex, curved throughout, apically flattened,
|
||
<quantity id="4C8D9B44EF2CFFB7FDEBF9F3CB45E4F1" box="[548,638,1644,1671]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="9" pageNumber="229" unit="mm" value="5.7">5.7 mm</quantity>
|
||
in length.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72F931CDC2E500" blockId="9.[188,1399,1709,1912]" pageId="9" pageNumber="229">
|
||
<smallCapsWord id="8D2CA07DEF2CFFB7FF72F931C87BE4B1" baselines="1729,1729" box="[189,320,1710,1736]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Prothorax" pageId="9" pageNumber="229">PROTHORAX</smallCapsWord>
|
||
. Carina along anterior pronotal edge broad medially, narrowing laterally. Anterolateral pronotal lobe smooth, sectioned posteriorly by abrupt shelf. Lateral pronotal carina and pronotal edge distinctly forking anteriorly then merging posteriorly, forming closed carinated ellipse. Pronotal surface smooth anteriorly, transitioning posterolaterally into chagrined texture. Centrally raised projection bifurcated with deeply excavated anteroventral surface defined by uneven texture. Propleuron and prosternum broadly setose; with long (~
|
||
<quantity id="4C8D9B44EF2CFFB7FD59F8C2CBE2E500" box="[662,729,1885,1911]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="229" unit="mm" value="1.0">1 mm</quantity>
|
||
) dark setae. Anterior prosternal apex rounded.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF2CFFB7FF72F800CD3FE5AB" blockId="9.[189,1399,1951,2012]" pageId="9" pageNumber="229">
|
||
<smallCapsWord id="8D2CA07DEF2CFFB7FF72F800C829E5C0" baselines="1970,1970" box="[189,274,1951,1977]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Elytra" pageId="9" pageNumber="229">ELYTRA</smallCapsWord>
|
||
. Glossy overall. Striae 1–6 widest anteriorly; narrowing posteriorly. Elytral stria eighth absent. Elytral striae nine absent on basal sixth. Interstriae convex, impunctate.
|
||
</paragraph>
|
||
<caption id="DF0A6629EF2FFFB4FF72F817CD05E59E" ID-DOI="http://doi.org/10.5281/zenodo.10824972" ID-Zenodo-Dep="10824972" httpUri="https://zenodo.org/record/10824972/files/figure.png" pageId="10" pageNumber="230" startId="10.[189,242,1928,1954]" targetBox="[215,1383,296,1868]" targetPageId="10">
|
||
<paragraph id="8BCA36A1EF2FFFB4FF72F817CD05E59E" blockId="10.[189,1399,1928,2025]" pageId="10" pageNumber="230">
|
||
<emphasis id="B901EAB3EF2FFFB4FF72F817C815E5D5" bold="true" box="[189,302,1928,1954]" pageId="10" pageNumber="230">Figs 5–8.</emphasis>
|
||
Dorsal habitus.
|
||
<emphasis id="B901EAB3EF2FFFB4FE3EF817C8C4E5D5" bold="true" box="[497,511,1928,1954]" pageId="10" pageNumber="230">5</emphasis>
|
||
.
|
||
<taxonomicName id="4C754D22EF2FFFB4FDC1F816CD3EE5D5" authority="Darling & Genier" authorityName="Darling & Genier" authorityYear="2024" box="[526,1029,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="230" phylum="Arthropoda" rank="species" species="grossiorum" status="sp. nov.">
|
||
<emphasis id="B901EAB3EF2FFFB4FDC1F816CA17E5D5" box="[526,812,1929,1954]" italics="true" pageId="10" pageNumber="230">Homocopris grossiorum</emphasis>
|
||
Darling & Génier
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257C8EF2FFFB4FBC2F816CD51E5D4" box="[1037,1130,1929,1955]" pageId="10" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (CEMT).
|
||
<emphasis id="B901EAB3EF2FFFB4FF72F834C9F7E5B2" bold="true" box="[189,204,1963,1989]" pageId="10" pageNumber="230">6</emphasis>
|
||
.
|
||
<taxonomicName id="4C754D22EF2FFFB4FF2CF833CB31E5B2" authorityName="Darling & Genier" authorityYear="2024" box="[227,522,1964,1989]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="230" phylum="Arthropoda" rank="species" species="grossiorum">
|
||
<emphasis id="B901EAB3EF2FFFB4FF2CF833CB31E5B2" box="[227,522,1964,1989]" italics="true" pageId="10" pageNumber="230">Homocopris grossiorum</emphasis>
|
||
</taxonomicName>
|
||
, allotype, ♀ (CEMT).
|
||
<emphasis id="B901EAB3EF2FFFB4FCF4F833CA71E5B1" bold="true" box="[827,842,1964,1990]" pageId="10" pageNumber="230">7</emphasis>
|
||
.
|
||
<taxonomicName id="4C754D22EF2FFFB4FCAEF833CC49E5B1" authority="(Curtis, 1845)" baseAuthorityName="Curtis" baseAuthorityYear="1845" box="[865,1394,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="230" phylum="Arthropoda" rank="species" species="punctatissimus">
|
||
<emphasis id="B901EAB3EF2FFFB4FCAEF833CD8AE5B2" box="[865,1201,1964,1989]" italics="true" pageId="10" pageNumber="230">Homocopris punctatissimus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE44B50EF2FFFB4FB05F834CC52E5B1" author="Curtis J." box="[1226,1385,1963,1990]" pageId="10" pageNumber="230" pagination="441 - 475" refId="ref25304" refString="Curtis J. 1845. Descriptions etc. of the insects collected by Captain P. P. King, R. N. F. R. S. & L. S. in the survey of the Straits of Magellan. The Transactions of the Linnean Society of London 19: 441 - 475. https: // doi. org / 10.1111 / j. 1096 - 3642.1842. tb 00371. x" type="journal article" year="1845">Curtis, 1845</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, specimen ♂ (CMNC).
|
||
<emphasis id="B901EAB3EF2FFFB4FE06F851C8E3E59F" bold="true" box="[457,472,1998,2024]" pageId="10" pageNumber="230">8</emphasis>
|
||
.
|
||
<taxonomicName id="4C754D22EF2FFFB4FE29F850CA16E59F" baseAuthorityName="Curtis" baseAuthorityYear="1845" box="[486,813,1999,2024]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="230" phylum="Arthropoda" rank="species" species="punctatissimus">
|
||
<emphasis id="B901EAB3EF2FFFB4FE29F850CA16E59F" box="[486,813,1999,2024]" italics="true" pageId="10" pageNumber="230">Homocopris punctatissimus</emphasis>
|
||
</taxonomicName>
|
||
, specimen ♀ (NMPC).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0A6629EF2EFFB5FF72F817CD9FE59F" ID-DOI="http://doi.org/10.5281/zenodo.10824974" ID-Zenodo-Dep="10824974" httpUri="https://zenodo.org/record/10824974/files/figure.png" pageId="11" pageNumber="231" startId="11.[189,242,1928,1954]" targetBox="[207,1370,321,1842]" targetPageId="11">
|
||
<paragraph id="8BCA36A1EF2EFFB5FF72F817CD9FE59F" blockId="11.[189,1399,1928,2025]" pageId="11" pageNumber="231">
|
||
<emphasis id="B901EAB3EF2EFFB5FF72F817C87EE5D5" bold="true" box="[189,325,1928,1954]" pageId="11" pageNumber="231">Figs 9–12.</emphasis>
|
||
Dorsal habitus.
|
||
<emphasis id="B901EAB3EF2EFFB5FDD1F817CB16E5D5" bold="true" box="[542,557,1928,1954]" pageId="11" pageNumber="231">9</emphasis>
|
||
.
|
||
<taxonomicName id="4C754D22EF2EFFB5FD8BF816CD6CE5D5" authority="(Eschscholtz, 1822)" baseAuthorityName="Eschscholtz" baseAuthorityYear="1822" box="[580,1111,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="231" phylum="Arthropoda" rank="species" species="torulosus">
|
||
<emphasis id="B901EAB3EF2EFFB5FD8BF816CA69E5D5" box="[580,850,1928,1954]" italics="true" pageId="11" pageNumber="231">Homocopris torulosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE44B50EF2EFFB5FCA5F817CD74E5D5" author="Eschscholtz J. F. G. von" box="[874,1103,1928,1954]" pageId="11" pageNumber="231" refId="ref25539" refString="Eschscholtz J. F. G. von 1822. Beschreibungen neuer Insectens. Entomographien. Erster Lieferung, G. Reimer, Berlin. https: // doi. org / 10.1515 / 9783112513941" type="book" year="1822">Eschscholtz, 1822</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, neotype, ♂ (CMNC).
|
||
<emphasis id="B901EAB3EF2EFFB5FF72F834C9E1E5B2" bold="true" box="[189,218,1963,1989]" pageId="11" pageNumber="231">10</emphasis>
|
||
.
|
||
<taxonomicName id="4C754D22EF2EFFB5FF21F833C8C3E5B2" baseAuthorityName="Eschscholtz" baseAuthorityYear="1822" box="[238,504,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="231" phylum="Arthropoda" rank="species" species="torulosus">
|
||
<emphasis id="B901EAB3EF2EFFB5FF21F833C8C3E5B2" box="[238,504,1963,1989]" italics="true" pageId="11" pageNumber="231">Homocopris torulosus</emphasis>
|
||
</taxonomicName>
|
||
, specimen ♀ (repository unknown).
|
||
<emphasis id="B901EAB3EF2EFFB5FC0EF834CAE6E5B2" bold="true" box="[961,989,1963,1989]" pageId="11" pageNumber="231">11</emphasis>
|
||
.
|
||
<taxonomicName id="4C754D22EF2EFFB5FC3FF833C836E59F" authority="Darling & Genier" authorityName="Darling & Genier" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="231" phylum="Arthropoda" rank="species" species="williami" status="sp. nov.">
|
||
<emphasis id="B901EAB3EF2EFFB5FC3FF833CDD6E5B2" box="[1008,1261,1963,1989]" italics="true" pageId="11" pageNumber="231">Homocopris williami</emphasis>
|
||
Darling & Génier
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257C8EF2EFFB5FEDBF850C84AE59E" box="[276,369,1999,2025]" pageId="11" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (MZSP).
|
||
<emphasis id="B901EAB3EF2EFFB5FDB1F851CBA0E59F" bold="true" box="[638,667,1998,2024]" pageId="11" pageNumber="231">12</emphasis>
|
||
.
|
||
<taxonomicName id="4C754D22EF2EFFB5FD65F850CA98E59F" authorityName="Darling & Genier" authorityYear="2024" box="[682,931,1998,2024]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="231" phylum="Arthropoda" rank="species" species="williami">
|
||
<emphasis id="B901EAB3EF2EFFB5FD65F850CA98E59F" box="[682,931,1998,2024]" italics="true" pageId="11" pageNumber="231">Homocopris williami</emphasis>
|
||
</taxonomicName>
|
||
, allotype, ♀ (MZSP).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FE97CD45E356" blockId="12.[189,1150,263,290]" box="[189,1150,263,290]" pageId="12" pageNumber="232">
|
||
<smallCapsWord id="8D2CA07DEF29FFB2FF72FE97C814E357" baselines="283,283" box="[189,303,264,290]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Hindwing" pageId="12" pageNumber="232">HINDWING</smallCapsWord>
|
||
(
|
||
<figureCitation id="134E2A24EF29FFB2FE8EFE98C8ACE356" box="[321,407,263,290]" captionStart="Figs 37–41. 37–38" captionStartId="23.[189,242,1892,1918]" captionTargetBox="[257,1314,294,1816]" captionTargetId="figure-17@23.[221,1366,265,1843]" captionTargetPageId="23" captionText="Figs 37–41. 37–38. Genital segment, dorsal view. 37. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 38. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 39–41. Hindwing, dorsal view. 39. Andinocopris achamas (Harold, 1867) gen. et comb. nov. 40. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov. 41. Homocopris torulosus." figureDoi="http://doi.org/10.5281/zenodo.10824988" httpUri="https://zenodo.org/record/10824988/files/figure.png" pageId="12" pageNumber="232">Fig. 40</figureCitation>
|
||
). Membrane edge between AP and AA lobed. AP vein reduced.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FED6CC23E3DE" blockId="12.[189,1398,328,425]" pageId="12" pageNumber="232">
|
||
<smallCapsWord id="8D2CA07DEF29FFB2FF72FED6C80FE316" baselines="348,348" box="[189,308,329,355]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Ventrites" pageId="12" pageNumber="232">VENTRITES</smallCapsWord>
|
||
. Meso-metasternal suture posteriorly arcuate. Surface of median metasternal lobe (
|
||
<figureCitation id="134E2A24EF29FFB2FAD8FED7CC54E314" box="[1303,1391,328,355]" captionStart="Figs 42–45" captionStartId="25.[189,242,1822,1848]" captionTargetBox="[232,1356,285,1784]" captionTargetId="figure-18@25.[232,1356,265,1787]" captionTargetPageId="25" captionText="Figs 42–45. Metasternum, ventral view. In specimens with a metasternal depression, the lateral depression is bordered with white dotted lines. For Homocopris grossiorum Darling & Génier sp. nov., a red dotted line indicates where the metasternal depression is interrupted. 42. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♀ (MNHN) 43. Homocopris grossiorum, holotype ♂ (MZSP). 44. Homocopris punctatissimus (Curtis, 1845), specimen ♀ (NMPC). 45. Homocopris torulosus (Eschscholtz, 1822), specimen ♂ (CMNC)." figureDoi="http://doi.org/10.5281/zenodo.10824990" httpUri="https://zenodo.org/record/10824990/files/figure.png" pageId="12" pageNumber="232">Fig. 42</figureCitation>
|
||
) finely punctate and convex, with scarce setigerous punctation along mesocoxae. Lateral metasternal lobe densely setose. Visible abdominal ventrites 1–6 laterally setose. Pygidium finely punctate.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FE4ECB6CE046" blockId="12.[188,1399,464,561]" pageId="12" pageNumber="232">
|
||
<smallCapsWord id="8D2CA07DEF29FFB2FF72FE4EC86EE39E" baselines="484,484" box="[189,341,465,491]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Prothoracic" pageId="12" pageNumber="232">PROTHORACIC</smallCapsWord>
|
||
<smallCapsWord id="8D2CA07DEF29FFB2FE94FE49C8B4E39E" baselines="484" box="[347,399,470,489]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="legs" pageId="12" pageNumber="232">LEGS</smallCapsWord>
|
||
. Anterior and posterior surface of profemur with dense row of long dark setae. Ventral profemoral surface finely punctate, lacking setae. Protibia with four distinct teeth. Protibial forespur straight with apex tapering inward.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FDC6CAB0E0CE" blockId="12.[189,1399,600,697]" pageId="12" pageNumber="232">
|
||
<smallCapsWord id="8D2CA07DEF29FFB2FF72FDC6C853E006" baselines="620,620" box="[189,360,601,627]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mesothoracic" pageId="12" pageNumber="232">MESOTHORACIC</smallCapsWord>
|
||
<smallCapsWord id="8D2CA07DEF29FFB2FEBEFDC1C89EE006" baselines="620" box="[369,421,606,625]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="legs" pageId="12" pageNumber="232">LEGS</smallCapsWord>
|
||
. Posterior surface of mesotrochanter with dense tuft of long rufous setae. Anterior and posterior surface of mesofemur with dense row of long dark setae. Outer surface of mesotibia serrate. Width at mesotibial apex one-third mesotibial length.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FD7ECA78E137" blockId="12.[189,1399,736,833]" pageId="12" pageNumber="232">
|
||
<smallCapsWord id="8D2CA07DEF29FFB2FF72FD7EC85EE08E" baselines="756,756" box="[189,357,737,763]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Metathoracic" pageId="12" pageNumber="232">METATHORACIC</smallCapsWord>
|
||
<smallCapsWord id="8D2CA07DEF29FFB2FEBCFD79C89CE08E" baselines="756" box="[371,423,742,761]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="legs" pageId="12" pageNumber="232">LEGS</smallCapsWord>
|
||
. Posterior surface of metatrochanter with tuft of long dark setae. Anterior and posterior surface of metafemur with dense row of long dark setae. Outer surface of metatibia serrate. Width at metatibial apex one-quarter metatibial length.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FCF7C811E19C" blockId="12.[189,1399,872,1003]" pageId="12" pageNumber="232">
|
||
<smallCapsWord id="8D2CA07DEF29FFB2FF72FCF7C9C4E1F7" baselines="891,891" box="[189,255,872,898]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="12" pageNumber="232">MALE</smallCapsWord>
|
||
<smallCapsWord id="8D2CA07DEF29FFB2FEC6FCF2C84CE1F7" baselines="891" box="[265,375,877,896]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="12" pageNumber="232">GENITALIA</smallCapsWord>
|
||
. Aedeagus length:
|
||
<quantity id="4C8D9B44EF29FFB2FD90FCF7CB84E1F5" box="[607,703,872,898]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="12" pageNumber="232" unit="mm" value="3.6">3.6 mm</quantity>
|
||
. Medially and laterally paired sclerites of genital segment broad, ill-defined. Paramere (
|
||
<figureCitation id="134E2A24EF29FFB2FDDBFC14CB5DE1D2" box="[532,614,907,933]" captionStart="Figs25–30" captionStartId="19.[189,242,1857,1883]" captionTargetBox="[239,1354,353,1767]" captionTargetId="figure-17@19.[234,1354,265,1809]" captionTargetPageId="19" captionText="Figs25–30. Paramere,lateralview.25. Andinocoprisachamas(Harold,1867)gen.etcomb.nov.,specimen ♂ (CMNC). 26. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 27. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 28. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 29. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 30. Homocopris williami Darling & Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824984" httpUri="https://zenodo.org/record/10824984/files/figure.png" pageId="12" pageNumber="232">Fig.26</figureCitation>
|
||
) laterally flattened, gently tapered; apex broad and slightly inclined with ventral tooth. FLP endophallite (
|
||
<figureCitation id="134E2A24EF29FFB2FD5AFC31CBCAE1BF" box="[661,753,942,968]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling & Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="12" pageNumber="232">Fig. 32</figureCitation>
|
||
) sigmoidal; broad distally, narrowing and inflecting centrally.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF24FB8CCB6BE65A" blockId="12.[189,1399,1043,1245]" box="[235,592,1043,1069]" pageId="12" pageNumber="232">
|
||
<emphasis id="B901EAB3EF29FFB2FF24FB8CC8FAE65A" bold="true" box="[235,449,1043,1069]" pageId="12" pageNumber="232">Female specimen</emphasis>
|
||
(
|
||
<figureCitation id="134E2A24EF29FFB2FE1FFB8CCB26E65A" box="[464,541,1043,1069]" captionStart="Figs 1–4" captionStartId="6.[189,242,1892,1918]" captionTargetBox="[234,1360,273,1801]" captionTargetId="figure-12@6.[218,1370,265,1852]" captionTargetPageId="6" captionText="Figs 1–4. Dorsal habitus. 1. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 2. Andinocopris achamas, alloreferent paralectotype, ♀ (MNHN). 3. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 4. Andinocopris buckleyi, specimen ♀ (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.10824970" httpUri="https://zenodo.org/record/10824970/files/figure.png" pageId="12" pageNumber="232">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="134E2A24EF29FFB2FDE6FB8CCB7CE65A" box="[553,583,1043,1069]" captionStart="Figs13–16" captionStartId="14.[189,242,1891,1917]" captionTargetBox="[221,1346,305,1841]" captionTargetId="figure-12@14.[221,1366,263,1841]" captionTargetPageId="14" captionText="Figs13–16.Headandpronotum,lateralview.13. Andinocoprisachamas(Harold,1867)gen.etcomb.nov., lectotype, ♂ (MNHN). 14. Andinocopris achamas, paralectotype, ♀ (MNHN). 15. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 16. Andinocopris buckleyi, specimen ♀ (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.10824978" httpUri="https://zenodo.org/record/10824978/files/figure.png" pageId="12" pageNumber="232">16</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FBA9CAEDE6AA" blockId="12.[189,1399,1043,1245]" pageId="12" pageNumber="232">
|
||
Similar to male with the following exceptions: body length
|
||
<quantity id="4C8D9B44EF29FFB2FC14FBA9CD6AE626" box="[987,1105,1078,1105]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="12" pageNumber="232" unit="mm" value="14.7">14.7 mm</quantity>
|
||
. Clypeal edge slightly sinuous medially. Clypeofrontal carina bi-tuberculate, with width equivalent to dorsal ocular width. Anterolateral pronotal lobe surface smooth, immediately transitioning to fused granulate texture posteriorly. Central pronotal ridge slightly overhanging, with lateral tubercles and shallow anterior excavation. Posterolateral and posterior pronotal surface chagrined.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FA9BC808E769" blockId="12.[189,307,1284,1310]" box="[189,307,1284,1310]" pageId="12" pageNumber="232">
|
||
<heading id="D08281CDEF29FFB2FF72FA9BC808E769" bold="true" box="[189,307,1284,1310]" fontSize="11" level="3" pageId="12" pageNumber="232" reason="3">
|
||
<emphasis id="B901EAB3EF29FFB2FF72FA9BC808E769" bold="true" box="[189,307,1284,1310]" pageId="12" pageNumber="232">Variation</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FAACCBD2E7E4" blockId="12.[187,1398,1331,1427]" pageId="12" pageNumber="232">
|
||
Body length
|
||
<quantity id="4C8D9B44EF29FFB2FE9EFAACC8EAE73A" box="[337,465,1331,1358]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="12" pageNumber="232" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
|
||
. Clypeal teeth absent or round and indistinct. Cephalic horn acute in small males; less than
|
||
<quantity id="4C8D9B44EF29FFB2FEE6FAC9C854E706" box="[297,367,1366,1393]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="232" unit="mm" value="1.0">1 mm</quantity>
|
||
in length. Pronotal armament in small males limited to notched anterior ridge. Pronotal ridge lacking lateral tubercles in small females.
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FA25C8FEE7A1" blockId="12.[189,453,1466,1494]" box="[189,453,1466,1494]" pageId="12" pageNumber="232">
|
||
<emphasis id="B901EAB3EF29FFB2FF72FA25C86DE7A3" bold="true" box="[189,342,1466,1492]" pageId="12" pageNumber="232">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="134E2A24EF29FFB2FEA9FA24C880E7A1" box="[358,443,1467,1494]" captionStart="Fig" captionStartId="36.[189,232,1922,1948]" captionTargetBox="[189,1398,975,1873]" captionTargetId="figure-182@36.[189,1398,975,1876]" captionTargetPageId="36" captionText="Fig. 65. Andinocopris achamas (Harold, 1867) gen. et comb. nov. and Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., distribution map. Legend and scale bar included in the top and bottom right corners, respectively." figureDoi="http://doi.org/10.5281/zenodo.10824996" httpUri="https://zenodo.org/record/10824996/files/figure.png" pageId="12" pageNumber="232">Fig. 65</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72FA76CA54E43D" blockId="12.[189,1399,1513,1610]" pageId="12" pageNumber="232">
|
||
High elevation (
|
||
<quantity id="4C8D9B44EF29FFB2FEB5FA76CB24E474" box="[378,543,1513,1540]" metricMagnitude="3" metricUnit="m" metricValue="2.84" metricValueMax="3.055" metricValueMin="2.625" pageId="12" pageNumber="232" unit="m" value="2840.0" valueMax="3055.0" valueMin="2625.0">2625–3055 m</quantity>
|
||
) in the Ecuadorian and northern Peruvian Andes. The single record from “Amazon” is based on an historical specimen, the
|
||
<typeStatus id="54CE8803EF29FFB2FCE3F993CAA8E451" box="[812,915,1548,1574]" pageId="12" pageNumber="232" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C754D22EF29FFB2FC0DF992CC0CE451" authority="Luederwaldt" authorityName=", Luederwaldt" authorityYear="1936" box="[962,1335,1548,1574]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="232" phylum="Arthropoda" rank="species" species="simulator">
|
||
<emphasis id="B901EAB3EF29FFB2FC0DF992CDACE451" box="[962,1175,1548,1574]" italics="true" pageId="12" pageNumber="232">Pinotus simulator</emphasis>
|
||
Luederwaldt
|
||
</taxonomicName>
|
||
with vague data (see Max Barclay pers. com., comment below).
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72F9EEC845E4FC" blockId="12.[189,382,1649,1675]" box="[189,382,1649,1675]" pageId="12" pageNumber="232">
|
||
<heading id="D08281CDEF29FFB2FF72F9EEC845E4FC" bold="true" box="[189,382,1649,1675]" fontSize="11" level="3" pageId="12" pageNumber="232" reason="3">
|
||
<emphasis id="B901EAB3EF29FFB2FF72F9EEC845E4FC" bold="true" box="[189,382,1649,1675]" pageId="12" pageNumber="232">Natural history</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB2FF72F93FC83BE4AA" blockId="12.[189,1399,1696,1757]" pageId="12" pageNumber="232">Specimens with data were collected in cloud forest using pitfall traps baited with fish and in bovine dung.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F652AEF29FFB3FF72F89BCBC2E31D" lastPageId="13" lastPageNumber="233" pageId="12" pageNumber="232" type="discussion">
|
||
<paragraph id="8BCA36A1EF29FFB2FF72F89BC814E569" blockId="12.[189,303,1796,1822]" box="[189,303,1796,1822]" pageId="12" pageNumber="232">
|
||
<heading id="D08281CDEF29FFB2FF72F89BC814E569" bold="true" box="[189,303,1796,1822]" fontSize="11" level="3" pageId="12" pageNumber="232" reason="3">
|
||
<emphasis id="B901EAB3EF29FFB2FF72F89BC814E569" bold="true" box="[189,303,1796,1822]" pageId="12" pageNumber="232">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCA36A1EF29FFB3FF72F8ACCBC2E31D" blockId="12.[188,1399,1843,2010]" lastBlockId="13.[189,1399,265,362]" lastPageId="13" lastPageNumber="233" pageId="12" pageNumber="232">
|
||
In the original description of
|
||
<taxonomicName id="4C754D22EF29FFB2FDEBF8ABCACCE53A" authority=", Luederwaldt (1936)" authorityName=", Luederwaldt" authorityYear="1936" box="[548,1015,1843,1869]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="232" phylum="Arthropoda" rank="species" species="simulator">
|
||
<emphasis id="B901EAB3EF29FFB2FDEBF8ABCBC3E53A" box="[548,760,1843,1869]" italics="true" pageId="12" pageNumber="232">Pinotus simulator</emphasis>
|
||
,
|
||
<bibRefCitation id="EFE44B50EF29FFB2FCC5F8ACCACCE53A" author="Luederwaldt H." box="[778,1015,1843,1869]" pageId="12" pageNumber="232" pagination="207 - 216" refId="ref26756" refString="Luederwaldt H. 1936. Notas complementares " As especies brasileiras do genero Pinotus ". Revista do Museu Paulista 20: 207 - 216." type="journal article" year="1936">Luederwaldt (1936)</bibRefCitation>
|
||
</taxonomicName>
|
||
describes the
|
||
<typeStatus id="54CE8803EF29FFB2FB67F8ACCDE0E53A" box="[1192,1243,1843,1869]" pageId="12" pageNumber="232">type</typeStatus>
|
||
specimen as “seeming atrophied, perhaps belonging to [
|
||
<taxonomicName id="4C754D22EF29FFB2FD74F8C8CA00E507" authorityName="Waterhouse" authorityYear="1891" box="[699,827,1878,1904]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="232" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF29FFB2FD74F8C8CA00E507" box="[699,827,1878,1904]" italics="true" pageId="12" pageNumber="232">P. buckleyi</emphasis>
|
||
</taxonomicName>
|
||
]”. Luederwaldt goes on to describe the cephalic and pronotal armament of the specimen as “a small quadrate hump” in place of a long cephalic horn, and a “small sinuous carina” instead of a large bifurcating pronotal projection. We suspected Luederwaldt was referring to a poorly developed male representative of
|
||
<taxonomicName id="4C754D22EF29FFB2FC92F85FCAE6E5AD" authorityName="Waterhouse" authorityYear="1891" box="[861,989,1984,2010]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="232" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF29FFB2FC92F85FCAE6E5AD" box="[861,989,1984,2010]" italics="true" pageId="12" pageNumber="232">A. buckleyi</emphasis>
|
||
</taxonomicName>
|
||
. Upon examination of photographs of the specimen in question, we have confirmed this to be the case; the external morphology of
|
||
<taxonomicName id="4C754D22EF28FFB3FF72FEB2C875E331" authorityName=", Luederwaldt" authorityYear="1936" box="[189,334,300,326]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="233" phylum="Arthropoda" rank="species" species="simulator">
|
||
<emphasis id="B901EAB3EF28FFB3FF72FEB2C875E331" box="[189,334,300,326]" italics="true" pageId="13" pageNumber="233">P. simulator</emphasis>
|
||
</taxonomicName>
|
||
matches a typical small male of
|
||
<taxonomicName id="4C754D22EF28FFB3FD21FEB2CA4EE331" authorityName="Waterhouse" authorityYear="1891" box="[750,885,300,327]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="233" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF28FFB3FD21FEB2CA4EE331" box="[750,885,300,327]" italics="true" pageId="13" pageNumber="233">A. buckleyi</emphasis>
|
||
</taxonomicName>
|
||
in all respects, and the name is therefore considered a new junior synonym of
|
||
<taxonomicName id="4C754D22EF28FFB3FDBFFECFCBC8E31E" authorityName="Waterhouse" authorityYear="1891" box="[624,755,335,362]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="233" phylum="Arthropoda" rank="species" species="buckleyi">
|
||
<emphasis id="B901EAB3EF28FFB3FDBFFECFCBC8E31E" box="[624,755,335,362]" italics="true" pageId="13" pageNumber="233">A. buckleyi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |